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276 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Schistocarpha oppositifolia (Kuntze) Rydb. This essentially 
tropical species is widely distributed in tropical America, and 
according to Rydberg in his account of the genus (N. Amer. Fl. 
34: 306. 1927) it extends northward to San Luis Potosi. A recent 
collection shows that it occurs, rather unexpectedly, also in Tamau- 
lipas: Jaumave, stony river valley, September 30, 1931, H. W. 
von Rozynski 76. 

Vernonia angusta (Gleason), comb. nov. Eremosis angusta 
Gleason, N. Amer. FL 33: 98. 1922. 









BOTANICAL SERIES 



FIELD MUSEUM OF NATURAL HISTORY 

FOUNDED BY MARSHALL FIELD, 1893 

VOLUME XI NUMBER 6 



REVISION OF THE GENUS COREOPSIS 



BY 

EARL EDWARD SHERFF 

RESEARCH ASSOCIATE IN SYSTEMATIC BOTANY 



B. E. DAHLGREN 

CURATOR, DEPARTMENT OF BOTANY 
EDITOR 



PUBLICATION 366 




CHICAGO, U.S.A. 
OCTOBER 20, 1936 



BOTANICAL SERIES 
FIELD MUSEUM OF NATURAL HISTORY 

FOUNDED BY MARSHALL FIELD, 1893 

VOLUME XI NUMBER 6 



REVISION OF THE GENUS COREOPSIS 

THE LIBRARY OF THE 

QCT281936 

UNIVERSITY OF ILLINOIS 
BY 

EARL EDWARD SHERFF 

RESEARCH ASSOCIATE IN SYSTEMATIC BOTANY 



B. E. DAHLGREN 

CURATOR, DEPARTMENT OF BOTANY 
EDITOR 



PUBLICATION 366 




CHICAGO, U.S.A. 
OCTOBER 20, 1936 



PRINTED IN THE UNITED STATES OF AMERICA 
BY FIELD MUSEUM PRESS 



REVISION OF THE GENUS COREOPSIS 



EARL EDWARD SHERFF 



PREFACE 

The revision of the genus Coreopsis as set forth in the following 

j pages is the outcome of a suggestion received in 1912. Early in 

that year Dr. Jesse M. Greenman, then Assistant Curator of 

- Taxonomy at Field Museum (now Curator of the Herbarium at 
: the Missouri Botanical Garden), selected from the Compositae the 

closely related genera Bidens, Coreopsis, and Cosmos as offering 
great promise for the student who should study them painstakingly. 

- In turn, each of these genera has been studied by me since then. 
3 The results of my investigations in Cosmos have been summarized 
I in large part in an earlier paper (Revision of the Genus Cosmos, 
I Field Mus. Bot. Ser. 8: 401-447. 1932) and will be concluded in 

a subsequent treatment to appear in the North American Flora. 

- My studies in Bidens and Coreopsis have been detailed in many 
articles which have appeared during more than two decades, particu- 

- larly in the Botanical Gazette and the American Journal of Botany, 
but also in the Journal of the Washington Academy of Sciences and 
the Journal of the Pan-Pacific Research Institution. In the present 
state of our knowledge, it is impossible, of course, to render a complete 

^ account of all the species in these two genera. Perhaps many, 

I especially in Bidens, remain to be discovered. I have, however, 

I attempted a revisional survey of each genus which should be as 

complete for the entire world as the many facilities offered me for 

research would permit. Treatments of the species growing in North 

^ America are to appear soon in the North American Flora. A com- 

v prehensive revision of the entire genus Bidens will appear in separate 

' volumes of the Field Museum Botanical Series. The present paper 

attempts a revisional summary of the entire genus of Coreopsis. 

No attempt has been made to present an extended history 
; of the genus Coreopsis. This history has been somewhat interlocked 
5 with that of Bidens and in my monograph of that genus I have 

- presented a historical re"sum to which readers are here referred. In 
addition, numerous references given accompanying the various sec- 

^ tions (infra) may be consulted with profit. 

Many of the world's principal scientific institutions have assisted 

- by lending materials or by permitting access to their herbaria, 

279 



280 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

libraries, or gardens. My very deep gratitude should be expressed 
here to the staffs of all these institutions and particularly to the 
following individuals: Dr. Ivan Murray Johnston, Research Asso- 
ciate, Arnold Arboretum; Dr. Adolph Engler (deceased), former 
Director, Dr. Ludwig Diels, present Director, and Dr. Johannes 
Mattfeld, Curator, Botanical Garden of Berlin; Dr. Alfred Barton 
Rendle, former Keeper of Botany, and Dr. John Ramsbottom, 
present Keeper of Botany, British Museum of Natural History; 
Dr. E. De Wildeman, former Director, Dr. Walter Robyns, present 
Director, and Dr. P. Staner (until recently Attach^ at the Museum 
of the Belgian Congo at Tervueren but now on the staff), of the 
National Botanical Garden at Brussels; Dr. Enrico Carano, former 
Director, Dr. Giovanni Negri, present Director, and Dr. Renato 
Pampanini, former Adjutant and Conservator at the Institute of 
Botany of the University of Florence (now Professor of Botany and 
Director at the Botanical Institute of the Royal University, Cagliari, 
Italy); Dr. John Isaac Briquet (deceased), former Director of the 
Conservatory and Botanical Garden of Geneva; Dr. Benjamin 
Lincoln Robinson (deceased), former Curator, and Mr. Charles A. 
Weatherby, Assistant Curator, Gray Herbarium of Harvard Univer- 
sity; Dr. Edgar Irmscher, Curator of the Herbarium of the Botanical 
Institute at Hamburg; Sir David Prain, former Director, Sir Arthur 
W. Hill, present Director, Dr. Otto Stapf (deceased), former Keeper 
of Herbarium, Mr. A. D. Cotton, present Keeper of Herbarium, and 
Miss M. L. Green, Botanist, Royal Botanical Gardens of Kew; Dr. 
Benjamin Daydon Jackson (deceased), former Secretary of the 
Linnean Society of London; Dr. Jesse More Greenman, Curator of 
the Herbarium, Missouri Botanical Garden; Dr. Nathaniel Lord 
Britton (deceased), former Director, New York Botanical Garden; Dr. 
H. LeComte, honorary Professor, Dr. Henri Humbert, Professor, Mr. 
Louis Anfray (deceased), former Preparator, and Dr. Paul Danguy, 
Vice-Director, Museum of Natural History of Paris; Dr. Philip Munz, 
Professor of Botany, Pomona College; Dr. Ira L. Wiggins, Stanford 
University; Dr. William R. Maxon, Associate Curator, United States 
National Museum; Dr. Richard Wettstein-Westersheim (deceased), 
former Director, Botanical Garden and Institute of Vienna. 

My work has been carried on from time to time in most of the 
above institutions and in numerous others, but principally in the 
Field Museum of Natural History. To the authorities of Field 
Museum am I indeed indebted for the many courtesies and invaluable 
aid extended to me over a long period of time. Especially do I 



REVISION OF THE GENUS COREOPSIS 281 

thank Mr. Stephen C. Simms, Director, Drs. Charles F. Millspaugh 
(deceased), former Curator of Botany, and B. E. Dahlgren, present 
Curator of Botany, and Mr. Paul C. Standley, Associate Curator of 
the Herbarium, for their assistance in many ways. 

Much effort has been made to eliminate typographic errors in 
the text. Throughout the preparation of my original manuscript 
and final text I have been aided greatly in the proof reading by my 
wife, Fern R. Seeley Sherff. It is a pleasure to acknowledge here 
her cooperation. 

During the progress of my work, I have photographed several 
hundred of the more important specimens studied, such as types 
and cotypes. Complete sets of these photographs are in my private 
collection and in the Herbarium of the Field Museum of Natural 
History. Throughout the text the term "cotype" is used to connote 
a duplicate of the type, as is usually shown, for example, by the use 
of the same collection number. 

Genus COREOPSIS: Descriptio 

L. Gen. PI. ed. 5. Num. 879. 1754; Sp. PL 907. 1753 (maxima 
pro parte); Acispermum Neck. Elem. Bot. 1: 34, No. 64. 1790. 
(For other synonyms see under names of various sections.) 

Plantae (familiae Compositarum) herbaceae vel saepe fruticosae, 
glabrae vel pubescentes. Folia opposita vel rarius alterna, indivisa 
atque integra dentatave, vel tripartita, vel semel bis terve ternatim 
vel pinnatim dissecta. Capitula mediocria vel majuscula, manifeste 
pedunculata vel pedicellata, solitaria vel laxe corymboso-paniculata, 
radiata. Involucri bracteae plerumque in 2 raro in 3-4 seriebus 
dispositae, basi plus minusve connatae, exteriores plerumque 
herbaceae vel submembranaceae adpressae vel saepe patentes, 
interiores plerumque majores brunneae vel flavae membranaceae. 
Receptaculum planum vel convexiusculum. Flores radii ligulati, 
1-seriati, neutri vel rarius styliferi ac fertiles vel steriles, lamina 
patente integri vel paucidentati, plerumque flavi rarius rosacei 
vel discolores. Paleae planae vel concaviusculae membranaceae 
striatae flores tubulosos subtendentes. Flores tubulosi flavi vel 
superne colorati, hermaphroditi ac fertiles vel intimi steriles, regulares, 
corollae limbo parum ampliati apice breviter 4- vel saepissime 5- 
dentati, saepe ad gutturis basim anulo glabro pilosove circumscripti ; 
antheris basi integris vel (auriculis minutis) sagittatis; stylorum 
ramis apice truncatis vel conicis vel breviter caudato-appendiculatis. 
Achaenia obcompressa, orbiculata vel oblonga vel plus minusve 



> 





282 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

oblengo-linearia, saepe 2-alata alls membranaceis vel indurato- 
crassatis integris vel fractis vel pectinato-dentatis planis vel incur- 
vatis, glabra vel (marginibus praesertim) villosa, epapposa vel 
aristis lanceolatis linearibusve glabris erecto-hispidulisve vel 2 
dentibus vel 2 paleis papposa (interdum medio apice poculo minuto 
coronata), numquam aristis retrorsum hamosis munita. 
Type species: Coreopsis lanceolata L. 

SECTIONES GENERIS 

Sect. I. Electro, (DC.) Blake, Proc. Amer. Acad. n. ser. 41: 337. 
1913; pro genere DC. Prodr. 5: 630. 1836. Nos. 1-4. (This and the 
next four sections included by Blake under the subgenus Leptosyne 
[DC.] Blake, loc. cit. 336. 1913; pro genere DC. Prodr. 5: 531. 1836; 
etiam A. Gray, Proc. Amer. Acad. 17: 218. 1882; etiam A. Gray, 
Syn. Fl. N. Amer. 1, pt. 2: 299. 1884; pro sect. Coreopsidis 0. Hoffm 
in Engler & Prantl, Nat. Pflanzenf. 4, pt. 5: 243. 1890 [exclud 
Epilepidem, Coreocarpum, et Acomam].) 

Suffruticosae, foliis oppositis, indivisis vel ternatim partitis, 
coriaceis, lamina segmentisve ovatis vel lanceolatis. Capitula 
solitaria vel paniculato-corymbosa. Involucri bracteae exteriores 
circ. 5, plus minusve oblongae; interiores circ. 8, longiores, plus 
minusve ovato-oblongae. Flores ligulati circ. 5, plerumque 2-3- 
dentati, ligula oblongi vel elliptici, fertiles, tubo pubescentes. 
Flores tubulosi limbo raro 4- plerumque 5-dentati, anulo deficiente, 
tube pubescenti quam gutture cylindraceo-infundibuliformi brevi 
ore; styli ramis apice subulato-appendiculatis. Achaenia valde 
obcompressa, glabra, marginata, exteriora lata interiora multo 
angustiora, omnia epapposa vel interiora raro jugo aristarum 
glabrarum angustarum munita. Plantae mexicanae, centrali 
americanae, et haitienses. (Type, Electra mexicana DC. = Coreopsis 
mutica var. leptomera.) 

Sect. II. Anathysana Blake, op. cit. 339. Nos. 5-7. 

Herbae perennes caulibus pluribus e radice lignea foliis oppositis 
integris vel pinnatiformibus lobis paucis filiformi-linearibus. Invo- 
lucrum ut apud Sect. I sed interius 8-12-bracteatum. Flores ligulati 
fertiles. Flores tubulosi saepius exanulati; styli ramis apice incras- 
satis breviter appendiculatis. Achaenia ut apud Sect. I, epapposa. 
Plantae mexicanae (num. 7 ex insula Socorro). (Type, Leptosyne 
mexicana A. Gray=Coreopsis cyclocarpa.} 

Sect. III. Tuckermannia (Nutt.) Blake, loc. cit. 340; pro genere, 
Nutt. Trans. Amer. Phil. Soc. ser. 2. 7: 363. 1841; pro sect. Lep- 



REVISION OF THE GENUS COREOPSIS 283 

tosynei, A. Gray, Bot. Calif. 1: 356. 1876; etiam Syn. Fl. N. Amer. 
l,pt. 2:300. 1884. Nos. 8 and 9. 

Perennes, robustae, foliis alternis carnosis 2-3-pinnatim dissectis, 
capitulis magnis. Involucri bracteae subaequales, exteriores 5-8, 
lanceolato-oblongae; interiores circ. 12, oblongae. Flores ligulati 
magni, fertiles. Flores tubulosi anulo subglabro circumscripti. 
Achaenia obcompressa, glabra, anguste alata, epapposa vel raro 
marginibus in dentes breves vel aristas currentibus aegre papposa. 
Plantae e California, Baja-California, et proximis insulis Oceani 
Pacifici. (Type, Tuckermannia maritime, Nutt.=Coreopsis maritima.) 

Sect. IV. Pugiopappus (A. Gray) Blake, loc. cit.; Agarista DC. 
Prodr. 5: 569. 1836 (nee alior.); pro genere, A. Gray, Pacif. R. Rept. 
4: 104. 1857; etiam Proc. Amer. Acad. 8: 659. 1873; pro sect. Lep- 
tosynei A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 300. 1884. Nos. 10 and 11. 

Annuae, e basi ramosae, foliis 2-3-pinnatfidis plerumque basali- 
bus, capitulis mediocribus solitariis, pedunculis elongatis subnudis. 
Involucri bracteae exteriores 5-7, interiores circ. 8. Flores ligulati 
plerumque styliferi ac fertiles, interdum neutri vel cum stylis brevibus 
inclusis, ligula lati ac multistriati. Flores tubulosi anulati, anulo 
hirsute. Achaenia dimorpha; radii epapposa, cortice marginata et 
ad facies quidem plus minusve costata; disci marginibus longe 
villosa, apice jugo aristarum lineari-lanceolatarum sursum setulo- 
sarum munita. Plantae Californiae australis. (Type, Agarista 
Calliopsidea DC.=Coreopsis Calliopsidea.) 

Sect. V. Euleptosyne (A. Gray) Blake, op. cit. 341; pro sect. 
Leptosynei A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 299. 1884. Nos. 12 
and 13. 

Habitu involucroque sect. Pugiopappo similis sed bracteis exte- 
rioribus linearis. Flores ligulati glabri, fertiles. Flores tubulosi 
anulati ; styli ramis apice incrassatis, breviter appendiculatis. Achae- 
nia phelloptera, apice poculo minuto coronata. Plantae Arizonae, 
Californiae, et Baja-Californiae borealis. (Type, Leptosyne Doug- 
lasii DC.=Coreopsis Douglasii.') 

Sect. VI. Pseudo- Agarista A. Gray, Proc. Amer. Acad. 22: 428. 
ISGl; Epilepis Benth. PI. Hartweg. 17. 1839. Nos. 14-40. 

Plus minusve fruticosae, foliis saepius decompositis segmentis 
angustis. Receptaculi paleae cum fructu saepe deciduae, ad hujus 
faciem exteriorem applicatae, apice saepius eroso-denticulatae. 
Flores tubulosi exanulati, styli ramis apice plus minusve conicis vel 
caudato-appendiculatis. Achaenia plus minusve lineari-oblonga, 



284 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

exalata sed marginibus sursum villosa, apice biaristata aristis 
linearibus vel lanceolatis antrorsum hispidis. Plantae mexicanae et 
austro-americanae. (Type, Coreopsis petrophila.) 

Sect. VII. Eucoreopsis Nutt. Trans. Amer. Phil. Soc. n. ser. 1: 
357. lS4l;Coreopsoides Moench, Meth. PI. 594. 1794; Anacis Schrank, 
Denkschr. Akad. Munch. Math. Nat. 5: 5. 1817; Leachia Cass. 
Diet. Sci. Nat. 25: 388. 1822; Chrysomelea Tausch, Hort. Canal. 
1823; Chrysostemma Less. Syn. Gen. Compos. 227. 1832; sect. 
Chrysomelea (Tausch) Nutt. loc. cit.; sect. Gyrophyllum Nutt. op. 
cit. 358; sect. Leachia (Cass.) A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 
291. 1884. Nos. 41-91. 

Herbae, foliis oppositis integris vel pinnatim partitis plerumque 
petiolatis. Ligulae plus minusve cuneatae, apice 3-5-lobatae vel 
-dentatae. Receptaculi paleae basi latae apicem versus attenuato- 
filiformes. Styli rami apice abrupte cuspidati vel conici. Achaenia 
fere orbiculata, ad maturitatem incurvata, nonnulla vel omnia 
demum papillata vel muriculata, saepe ventris apice basique callum 
(hoc quidem in unico capitulo saepe valde polymorphum) ferentia; 
pappo 2 dentibus minutis paleaceis vel deficiente. Plantae boreali- 
americanae et africanae, rarissime (C. lanceolata) in Asia. (Type, 
Coreopsis coronata Hook.=Coreopsis nuecensis.) 

Sect. VIII. Silphidium Torr. & Gray ex A. Gray, Syn. Fl. N. 
Amer. 1, pt. 2: 294. 1884; pro parte sect. Eucoreopsidis Torr. & Gray, 
Fl.N. Amer. 2:341. 1843. No. 92. 

Folia opposita, magna, indivisa, serrata. Ligulae integrae. 
Styli rami apice ovato-triangulati, breviter mucronati. Achaenia 
obovato-oblonga, exalata, apice obsolete bidentata. Plantae e 
Georgia ad Boreali-Carolinam. (Type, Coreopsis latifolia.) 

Sect. IX. Calliopsis (Reichenb.) Nutt. Trans. Amer. Phil. Soc. 
n. ser. 7: 360. 1841; pro genere, Reichenb. Mag. Aesthet. Bot. No. 
70. 1823; etiam DC. Prodr. 5: 568. 1836; Diplosastera Tausch, Hort. 
Canal. 1823; cf. Flora 1: 83. 1824; sect. Coreoloma (pro minima parte 
ex exemplo C. Leavenworthii) et subsect. Calliopsidium (omnino) 
Torr. & Gray, op. cit. 346. Nos. 93-100. 

Herbae, foliis oppositis raro integris plerumque 1-2-pinnatim 
divisis lobis integris. Involucrum exterius parvum. Ligulae plerum- 
que bicolores vel quidem rosaceae, apice dentatae. Styli rami apice 
truncati ac subpenicillati. Achaenia apice calva vel biaristata, 
alata vel exalata. Plantae boreali-americanae, rarissime ex aliqua 
parte (C. lanceolata et C. tinctoria) sinenses. (Type, Calliopsis bicolor 
Reichenb. = Coreopsis tinctoria.} 



REVISION OF THE GENUS COREOPSIS 285 

Sect. X. Eublepharis Nutt. Trans. Amer. Phil. Soc. n. ser. 7: 
359. 1841; sect. Rabdocaulis Nutt. loc. cit.; sect. Coreoloma Torr. 
& Gray, Fl. N. Amer. 2: 346. 1843 (exclud. C. Leavenworthii) ; sect. 
Cosmella Torr. & Gray, op. cit. 348 (exclud. C. roseam). Nos. 101-111. 

Herbae, foliis oppositis vel alternis, plerumque integris. Ligulae 
flavae vel rosaceae, dentatae. Flores disci atro-purpurei vel flavi. 
Styli rami truncati vel obtuse conici. Achaenia alata alis fimbriatis 
vel pectinato-dissectis. Plantae boreali-americanae praesertim e 
parte austro-orientali. (Type, Coreopsis gladiata.) 

Sect. XI. Euprestinaria Schz. Bip. in Walp. Repert. 6: 163. 1846. 
Nos. 112-114. 

Herbae annuae vel perennes. Florum tubulosorum tubus glaber, 
styli ramis apice cono hispido superatis. Achaeniorum oblongorum 
vel oblongo-lanceolatorum aristae paleaceae lanceolato-lineares utrin- 
que ciliis adrectis hispidae. Plantae africanae praesertim abyssini- 
cae. (Type, Coreopsis Prestinaria.) 

CLAVIS 

a. Plantae boreali-americanae antillanaeque. 
6. Flores ligulati fertiles. 
c. Herbae annuae. 
d. Disci achaenia exaristata, phelloptera. 

e. Achaenia setis clavellatis numerosis obsita. 

12. C. Douglasii. 

e. Achaenia non clavellato-setosa 13. C. Stillmanii. 

d. Disci achaenia biaristata, longe ciliata. 

e. Involucri bracteae exteriores lineares . . 10. C. Bigelovii. 
e. Involucri bracteae exteriores late ovatae. 

11. C. Calliopsidea. 
c. Herbae perennes vel frutices. 
d. Folia simplicia vel ternatisecta, lamina vel foliolis sub- 

anguste lanceolatis vel latioribus. 

e. Capitula numerosa cymoso-paniculata; foliis vel foliolis 
nunc plus minusve lanceolatis nunc oblongo-ovatis. 

1. C. mutica. 

e. Capitula pauca vel solitaria; foliis simplicibus. 
/. Folia 2-6 cm. longa. 

g. Folia cuneate oblanceolata vel obovata. 

2. C. cuneifolia. 



286 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

g. Folia anguste lanceolato-oblonga .... 4. C. Buchii. 

f. Folia 1-2 cm. longa, ovalia 3. C. parvifolia. 

d. Folia vel segmenta linearia vel angustiora. 
e. Folia integra vel raro lateraliter 2-lobata, 2-6 cm. longa. 

5. C. cyclocarpa. 

e. Folia principalia pinnatim lobata. 
/. Flores ligulati usque ad 11 mm. longi. 

g. Involucrum circ. 6-8 mm. altum, foliorum lobis 

lateralibus plerumque 3 jugis ... 6. C. pinnatisecta. 

g. Involucrum circ. 4-5 mm. altum, foliorum lobis 

lateralibus saepius 1 jugo 7. C. insularis. 

/. Flores ligulati saltern 2 cm. longi. 

g. Pedunculi dispersi, plerumque 1.5-4 dm. longi. 

8. C. maritima. 

g. Pedunculi corymbose congregati plerumque 1-2 
(saepius usque ad 1.5) dm. longi. .9. C. gigantea. 
b. Flores ligulati steriles. 
c. Fruticosae. 

d. Achaenia glabra apice calva 51. C. cordylocarpa. 

d. Achaenia ciliata, biaristata. 

e. Folia simplicia 15. C. petrophiloides. 

e. Folia 1-3-pinnatisecta. 
/. Involucri bracteae interiores 5-6 mm. longae. 

14. C. petrophila. 

f. Involucri bracteae interiores 6-9 mm. longae. 
g. Plantae glabrae vel foliis vix pilosiusculae. 
h. Foliorum segmenta ultima 0.6-0.8 mm. lata; 
involucri bracteis exterioribus lineari-oblongis. 

18. C. Pringlei. 

h. Foliorum segmenta ultima 1-5 mm. lata; involu- 
cri bracteis exterioribus linearibus. 

17. C. rhyacophila. 

g. Plantae hispidae 19. C. rudis. 

c. Herbaceae. 1 
d. Stylorum rami termino acuti vel cuspidati. 

1 For convenience in making comparisons, the plants under c are keyed as 
closely as feasible according to the characters utilized by F. E Boynton in his 
treatment of the species of Coreopsis in the southeastern United States (J. K. Small, 
Fl. S. E. United States 1274. 1903). 



REVISION OF THE GENUS COREOPSIS 287 

e. Achaeniorum alae dissecte fimbriato-pectinatae. 

101. C. saxicola. 

e. Achaeniorum alae angustae vel latae, integrae (interdum 
fractae vel scissae) vel (C. congregata) supra quidem 
rotundo-lobatae sed numquam fimbriato-pectinatae. 

/. Foliorum laminae vel segmenta margine integra. 

g. Stylorum rami termino cuspidati; paleis basi latis 
superne attenuato-filiformibus; foliorum laminis 
simplicibus vel pinnatim lobatis. 

h. Perennes; ligulis omnino flavis. 

i. Achaenia alata alis patulis vel minime incur- 
vatis. 

j. Achaeniorum corpora (alis exclusis) circ. 2 
mm. longa et circ. 1 mm. lata. 43. C. debilis. 

j. Achaeniorum corpora (alis exclusis) 2.5-3.2 
mm. longa et saltern 1.4-2 mm. lata. 

k. Caules basim versus foliosi, gerentes ramos 
longos nudos pedunculoideos. 

L Involucri bracteae exteriores angustis- 
sime lineares, 5-11 mm. longae. 

46. C. heterolepis. 

L Involucri bracteae exteriores lanceolatae 
vel oblongo-lineares veloblongo-ovatae, 
3-8 mm. longae. 

m. Folia petiolis inclusis plerumque 3-8 
cm. longa, involucri bracteis exteri- 
oribus 3-5 mm. longis. 

42. C. corninsularis. 

m. Folia petiolis inclusis plerumque 

usque ad 1.5 dm. longis, involucri 

bracteis exterioribus 4-8 mm. longis. 

41. C. lanceolata. 

k. Caules fere usque ad summam foliosi; 
pedunculis moderate brevibus. 

I. Foliorum laminae latae, simplices vel 
lateraliter 1-5-lobatae. 

w. Involucri bracteae subaequales 7-10 
mm. longae 45. C. pubescens. 



288 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

m. Involucri bracteae exteriores 4-7 
(raro -8) mm. interiores 12-14 mm. 

longae 44. C. intermedia. 

I. Foliorum laminae in segmenta angusta 

pinnatim partitae. . . 47. C. grandiflora. 

i. Achaenia alata alis valde incurvatis integris 

demum calloso-incrassatis. . .48. C. auriculata. 

h. Annuae. 

i. Ligulae discolores 50. C. nuecensis. 

i. Ligulae omnino flavae 49. C. congregata. 

g. Stylorum rami termino acute conici; paleis lineari- 
bus vel termino vix dilatatis; foliorum laminis 
plerumque palmato-lobatis vel -divisis (pro C. 
major e var. Oemleri simplicibus). 
h. Foliorum laminae petiolatae. . . .91. C. tripteris. 
h. Foliorum laminae sessiles. 
i. Foliorum laminae infra medium non vel parce 

sectae 86. C. palmata. 

i. Foliorum laminae usque ad basim partitae. 
j. Flores tubulosi corollis flavi. 

k. Foliorum segmenta filiformia vel lineari- 
filiformia 0.3-1 mm. lata. 

87. C. verticillata. 

k. Foliorum segmenta latiora. . .90. C. major, 
j. Flores tubulosi corollis purpureo-brunnei. 
k. Foliorum principalium segmenta pauca, 
plerumque 1.5-3.5 mm. lata; internodiis 
quam foliis saepius longioribus. 

89. C. Delphinifolia. 

k. Foliorum principalium segmenta numero- 
sa, 0.5-1.5 mm. latis; internodiis quam 
foliis plerumque brevioribus. 

88. C. pulchra. 
f. Foliorum laminae regulariter dentatae. 

92. C. latifolia. 

d. Stylorum rami termino truncati vel obtuse conici. 
e. Achaenia exalata. 

/. Ligulae basi rubro-brunneae alibi flavae; disci floribus 
corolla atro-rubris. 



REVISION OF THE GENUS COREOPSIS 289 

g. Involucri bracteae exteriores quam interiores multo 
breviores; achaeniis anguste oblongis, tenuibus, 

subplanis , 93. C. tinctoria. 

g. Involucri bracteae exteriores quam interiores non 
plerumque breviores; achaenia obovata, crassa, 

convexa 94. C. basalis. 

f. Ligulae roseae; disci floribus corolla flavis. 

95. C. rosea. 
e. Achaenia alata. 
/. Achaeniorum alae integrae. 

g. Achaenia apice calva vel minute dentata. 
h. Ligulae discolores. 
i. Achaenia circ. 2 mm. longa, anguste vel late 

alata 96. C. Cardaminefolia. 

i. Achaenia 2.4-2.8 mm. longa, angustissime alata 
(vel interdum tantum marginata). 

99. C. Atkinsoniana. 

h. Ligulae omnino flavae 97. C. stenophylla. 

g. Achaenia perspicue aristata aristis interdum in 

maturitate deciduis. 

h. Ligulae discolores (specie texana) . . 98. C. similis. 
h. Ligulae omnino flavae (specie floridana). 

100. C. Leavenworthii. 

/. Achaeniorum alae dissecte fimbriato-pectinatae. 
g. Folia non juncoidea; ligulis flavis. 

h. Foliorum majorum laminae plus minusve oblongae 

vel lanceolatae. 

i. Folia inferiora quam internodia breviora vel 
paulo longiora, opposita. . .103. C. Linifolia. 
i. Folia inferiora quam internodia multo longiora. 
;'. Foliorum imorum laminae lineares vel lanceo- 
latae, apice plerumque mucronatae. 
k. Foliorum laminae simplices integraeque; 
achaeniis elliptico-oblongis, circ. 4 mm. 
longis, ala quam corpore multo angus- 

tiore 104. C. longifolia. 

k. Foliorum laminae saepe lobatae; achaeniis 
anguste oblongis, circ. 5 mm. longis, ala 
corpori latitudine aequali. 

105. C. falcata. 



290 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

y. Foliorum imorum laminae oblongae vel 

ellipticae, apice obtusae. . . 106. C. gladiata. 

h. Foliorum majorum laminae plus minusve ovatae. 

i. Folia opposita; petiolis (ac laminis juvenilibus) 

ciliatis 108. C. integrifolia. 

i. Folia alterna; petiolis (ac laminis quidem 

juvenilibus) eciliatis. . .107. C. Helianthoides. 

g. Folia juncoidea, laminis veris deficientibus; ligulis 

rosaceis 109. C. nudata. 

a. Plantae austro-americanae. 
6. Folia indivisa. 
c. Folia 0.5-2 cm. lata. 
d. Folia oblanceolata, 0.5-1.5 cm. lata. 

e. Folia integra, 5-7.5 mm. lata 25. C. oblanceolata. 

e. Folia supra medium acriter paucidentata, 0.8-1.5 cm. 
lata 25a. C. Woytkowskii. 

d. Folia anguste ovato-lanceolata, 1-2 cm. lata. 

16. C. Irmscheriana. 
c. Folia angustiora. 

d. Folia lineari-flagellaria tantum 0.5-1 mm. lata, totam 
longitudinem aequaliter angusta 26. C. longula. 

d. Folia angustissime spathulato-linearia, 1-2 mm. lata. 

27. C. venusta. 
b. Folia divisa. 
c. Folia primaria minuta, tantum 7-9.5 mm. longa. 

d. Folia cum fasciculis axillaribus, quam internodia plerum- 
que paulo brevioria; involucri bracteis exterioribus 
biserialibus circ. 18 29. C. senaria. 

d. Folia sine fasciculis axillaribus, quam internodia saltern 
duplo longiora; involucri bracteis exterioribus uniseri- 

alibus circ. 8 28. C. nodosa. 

c. Folia primaria majora. 
d. Capitula pansa ad anthesin circ. 1.1-2 cm. lata; foliorum 

laminis vel segmentis 1.5-4 mm. latis. 
e. Partes novellae glaucescentes; involucri bracteis exte- 
rioribus linearibus quam interioribus dimidio breviori- 

bus. 20. C. glaucodes. 

e. Nullae partes glaucescentes. 



REVISION OF THE GENUS COREOPSIS 291 

/. Involucri bracteae exteriores minutae, longitudine 

tantum circa tertiae interiorum. 
g. Involucri bracteae exteriores ovatae; foliorum seg- 
mentis lateralibus cuneatis vel elliptico-obovatis 
vel spathulatis, 2-6 mm. latis . . . 21. C. microlepis. 
g. Involucri bracteae exteriores lineari-oblongae; foli- 
orum segmentis lateralibus anguste linearibus, 

1-1.7 mm. latis 22. C. Macbridei. 

/. Involucri bracteae exteriores lineari-oblongae, quam 

interiores dimidio breviores 24. C. parviceps. 

d. Capitula pansa ad anthesin 2.2-5 cm. lata. 

e. Folia densissime congregata, ramis fere abditis. 

/. Foliorum segmenta lineari-oblanceolata, apicaliter 

subobtusa; involucri bracteis exterioribus circ. 8, 

oblongis, apice rotundatis, circ. 5 mm. longis et 

1.5 mm. latis, uniseriatim dispositis . . 30. C. foliosa. 

f. Foliorum segmenta lineari-subulata, apicaliter acuta; 

involucri bracteis exterioribus circ. 11, ovatis, 

circ. 6 mm. longis et 3.3-4.8 mm. latis, biseriatim 

dispositis 31. C. polyactis. 

e. Folia plerumque laxius (C. trilobae interdum dense) 

disposita, ramis plus minusve manifestis. 
/. Foliorum segmenta ultima lanceolata vel oblonga; 
involucri 6 bracteis exterioribus 2-3 mm. longis et 

1-1.6 mm. latis 32. C. notha. 

/. Foliorum segmenta ultima angustiora. 

g. Foliorum triloborum segmenta ultima flagellaria, 
apicaliter acuta, 1-3.5 cm. longa et plerumque 
0.3-0.6 mm. lata. 
h. Folia laxissime patentia vel etiam subreflexa. 

33. C. capillacea. 

h. Folia dense adgregata, suberecta. .34. C. triloba. 
g. Foliorum segmenta ultima diversa. 

h. Involucra glaberrima 35. C. spectabilis. 

h. Involucra (saltern infra) hispida. 

i. Kami foliaque hinc illinc resinosa, suaveolentia. 

36. C. suaveolens. 

i. Kami foliaque diversa (petiolis raro inferne 
resinosis). 



292 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

j. Involucri bracteae exteriores biseriales, circ. 

16 38. C. imbricata. 

j. Involucri bracteae exteriores uniseriales, 

multo pauciores. 
k. Foliorum primariorum multa 2.5-4.5 cm. 

longa. 

I. Achaenia villoso-ciliata dorso sub palea 
glabra sed ad ventrem villosissima. 

37. C. Pickeringii. 
1. Achaenia villoso-ciliata sed faciebus 

glabra 39. C. Townsendii. 

k. Folia primaria 1-2.5 cm. longa. 

1. Internodia superiora multo folia super- 
antia, ligulis siccis brunneo-flavis. 

23. C. Killipii. 

I. Internodia superiora non vel moderate 
folia superantia, ligulis siccis claro- vel 

subclaro-flavis 40. C. fasciculata. 

a. Plantae africanae. 

b. Achaenia valde obcompressa, marginibus saepissime tenuia 
alatave. 

c. Aristae achaeniorum filiformes ac corporibus breviores, vel 
deficientes. 

d. Plantae annuae (vel verisimiliter annuae). 
e. Folia superne hispida. 
/. Achaenia (alis inclusis) circ. 2-3.5 mm. lata. 
g. Involucri bracteae exteriores 6-11 mm. longae. 
h. Flores ligulati circ. 2.5-3 cm. longi. 

76. C. oligoflora (vide etiam 72. C. multifloram). 
h. Flores ligulati circ. 1-2 cm. longi. . . 79. C. vulgaris. 
g. Involucri bracteae exteriores 3-6 mm. longae. 

h. Gapitula numerosa, ad anthesin circ. 2.5-3.3 cm. 
lata, demum cum achaeniis maturis 1.2 cm. 
alta et plerumque paulo angustiora, achaeni- 
orum corporibus plerumque 7-10 mm. longis, 
aristis perspicue densissimeque erecto-hispidis 
supra paleas valde manifestis. 

80. C. Giorgii (vide etiam 78. C. Goffardii}. 



REVISION OF THE GENUS COREOPSIS 293 

h. Capitula pauca, ad anthesin 4 cm. lata, demum 
cum achaeniis maturis sub 1 cm. alta et 1.3 
cm. lata, achaeniorum corporibus 3.5-6 mm. 
longis, aristis minutissimis ac non supra paleas 

manifestis 77. C. Quarrei. 

h. Capitula subnumerosa, ad anthesin circ. 4-4.5 
cm. lata, demum cum achaeniis maturis circ. 
1 cm. alta et parce latiora, achaeniorum cor- 
poribus 4.5-6.5 mm. longis, aristis in paleis 

subabditis 82. C. injucunda. 

/. Achaenia (alis inclusis) 4-5 mm. lata. 

81. C. Mattfeldii. 
e. Folia superne glabra vel glabrata. 

/, Achaenia exaristata 56. C. Negriana. 

f. Achaenia biaristata. 

g. Foliorum caulinorum saepius 2-3-pinnatisectorurn 
segmenta anguste linearia; capitulis pansis ad 
anthesin 2.5-5 cm. latis. 
h. Foliorum segmenta plerumque circ. 1 mm. lata; 

involucro hispido 71. C. leptoglossa. 

h. Foliorum segmenta plerumque 2-4 mm. lata; 
involucro glabrato vel subglabrato. 

69. C. Feruloides. 

g. Foliorum 2-pinnatisectorum segmenta principalia 
oblongo-lanceolata; capitulis pansis ad anthesin 
2.5-4 cm. latis. 

61. C. Prestinariaeformis var. /3 incisa. 

g. Foliorum caulinorum indivisorum laminae vel 3-5- 
partitorum segmenta late linearia; capitulis pansis 
ad anthesin 5-7 cm. latis 83. C. Borianiana. 

g. Foliorum 3-5-lobatorum foliola ovata vel ovato- 

sublanceolata 61. C. Prestinariaeformis. 

d. Plantae perennes (vel verisimiliter perennes). 

e. Foliorum principalium laminae vel segmenta lanceolata 

vel latiora. 

/. Foliorum segmenta dentata dentibus perspicue elon- 
gatis (saepe 1-1.5 cm. longis) . . . 110. C. Mildbraedii. 
f. Foliorum laminae vel segmenta diversa. 
. Folia indivisa. 



294 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

h. Folia scabra, saepius 10-16 cm. longa; involucre 

hispido 85. C. scabrifolia. 

h. Folia glabra, 1.5-5 cm. longa; involucre glabro. 

62. C. Barteri. 
g. Folia divisa. 

h. Achaenia exaristata. 

i. Folia plerumque pinnata vel pinnatifida seg- 
mentis principalibus saepius late oblongo- 
lanceolatis vel ovatis, inter se saepe propin- 
quis, supra scaberulis subtus tomentellis. 
j. Caulis primo tomentellus mox glabratus. 

66. C. Whytei. 
y. Caulis perpetuo tomentellus. 

65. C. pinnatipartita. 

i. Folia 2-3-pinnatisecta, foliolis vel segmentis 
principalibus saepius anguste oblongo-lance- 

olatis, glabris 67. C. Lupulina. 

h. Achaenia biaristata. 
i. Capitula pansa ad anthesin 5.5-9 cm. lata. 
y. Folia pubescentia. 

73. C. bella (vide etiam. .72. C. multifloram). 

j. Folia faciebus glabra 84. C. togensis. 

i. Capitula pansa ad anthesin usque ad 4 cm. lata. 

y. Folia molliter denseque pubescentia; achae- 

niis plumbeo-nigris minutis corpore tantum 

3-4 mm. longis 68. C. Neumannii. 

y. Folia glabra vel subglabra. 

k. Pauca folia divisa. ...... .62. C. Barteri. 

k. Folia plerumque divisa. 

Z. Involucrum moderate vel valde hispidum. 

m. Involucrum moderate adpresso-his- 

pidum; pedunculo tenui glabra- 

toque; floribus tubulosis extrinsecus 

glabris 75. C. exilis. 

m. Involucrum valde hispidum; pedun- 
culo subrobusto hispidoque; floribus 
tubulosis inferne pilosis. 

64. C. oblonga. 
I. Involucrum non nisi basi hispidum. 



REVISION OF THE GENUS COREOPSIS 295 

m. Caulis rigidus superne circ. 2.5-3.5 
mm. crassus, foliis principalibus pe- 
tiolo adjecto circ. 4-5 cm. longis, 
capitulis pansis 4-5.5 cm. latis, 
planta kamerunensi . 63. C. monticola. 
m. Caulis gracilis superne 1-2 mm. 
crassus, foliis principalibus petiolo 
adjecto 5-8 cm. longis, capitulis 
pansis 3-3.5 cm. latis, planta ango- 

lensi 74. C. Curtisii. 

e. Foliorum principalium laminae vel segmenta late vel 

anguste linearia. 

/. Foliorum dentes saepe in setas elongatas desinentes; 
lamina indivisa, usque ad 2 dm. longa et 4-10 mm. 

lata 111. C. bracteosa. 

f. Foliorum dentes non setigerae. 

g. Folia inferiora 1-2 dm. longa; involucri bracteae 
exteriores circ. 12, lineari-subulatae, 1-1.5 cm. 

longae 57. C. camporum. 

g. Folia inferiora raro 1 dm. longa. 
h. Frutices. 

i. Achaenia exaristata. 
;'. Segmenta ultima 2-3 mm. lata. 

52. C. elgonensis. 
j. Segmenta ultima multo angustiora. 

53. C. Chippii. 

i. Achaenia biaristata 54. C. scopulorum. 

h. Herbae. 

i. Capitula pansa ad anthesin 4-8.5 cm. lata. 
j. Involucri bracteae exteriores elongatae, lat- 
eraliter lobatae; ligulis sulphureis. 

58. C. Ellenbeckii. 
j. Involucri bracteae exteriores normales; ligulis 

aurantiacis 70. C. ochracea. 

i. Capitula pansa ad anthesin circ. 2-2.5 cm. lata. 
;'. Folia atro-viridia, segmentis plerumque 

1.2-2.5 mm. latis 59. C. lineariloba. 

j. Folia pallida, segmentis plerumque circ. 1 
(raro 2) cm. latis 60. C. Schimperi. 



296 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

c. Aristae achaeniorum subulatae. 
d. Herbae annuae. 

e. Involucri bracteae interiores glabrae.112. C. Prestinaria. 
e. Involucri bracteae interiores pilis subplanis obtectae. 

61. C. Prestinariaeformis et var. /3 incisa. 

d. Herbae perennes 113. C. macranlha. 

b. Achaenia tantum moderate vel aegre obcompressa. 

c. Achaenia exaristata; foliis indivisis spathulato-oblanceolatis- 

55. C. Jacksonii- 

c. Achaenia biaristata aristis lanceolato-subulatis; foliis bipin<- 
natipartitis 114. C. pachyloma et var. inanis. 

ABBREVIATIONS USED FOR HERBARIA CITED 

Berl. Herb. Berlin Botanical Garden, Berlin-Dahlem. 

Boiss. Herb. Boissier, Geneva. 

Brit. Herb. British Museum of Natural History, South 

Kensington. 

Bruss. Herb. National Botanical Garden, Brussels. 

Calif. Herb. University of California, Berkeley. 

Del. Herb. Delessert, Geneva. 

Field Herb. Field Museum of Natural History, Chicago. 

Flor. Herb. Institute of Botany, University of Florence, 

Italy. 
Gray Herb. Gray, Harvard University, Cambridge, 

Massachusetts. 

Hamb. Herb. Hamburg Botanical Garden, Germany. 

Kew Herb. Royal Botanic Gardens, Kew. 

Linn. Herb. Linnaeus, Linnaean Society, London. 

Mo. Herb. Missouri Botanical Garden, St. Louis. 

Mun. Herb. Munich Botanical Garden, Germany. 

Mus. Cong. Herb. Musei Congoensis, Tervueren, Belgium. 

(Recently merged with the Herbarium of the 

National Botanical Garden, Brussels.) 
Mus. V. Herb. Museum of Natural History, Vienna. 
N. Eng. Herb. New England Botanical Club, Cambridge, 

Massachusetts. 
N. Y. Herb. New York Botanical Garden, New York City. 



REVISION OF THE GENUS COREOPSIS 297 

Oxf. Herb. University of Oxford, Oxford. 

Par. Herb. Museum of Natural History, Paris. 

Phila. Herb. Philadelphia Academy of Natural Sciences, 

Philadelphia. 

Pom. Herb. Pomona College, Claremont, California. 

Stanf. Herb. Stanford University, California. 

Stockh. Herb. National Natural History Museum, Stock- 

holm. 

U. S. Herb. United States National Museum, Wash- 

ington. 

U. V. Herb. University of Vienna, Austria. 

1. Coreopsis mutica DC. Prodr. 5: 571. 1836. ElectraGaleottii 
A. Gray, PI. Wright. 110. 1852. C. Galeottii (A. Gray) Hemsl. Biol. 
Centr. Amer. Bot. 2: 195. 1881. C. mutica var. genuina Blake, 
Proc. Amer. Acad. 52: 55. 1-917. 
Foliorum segmenta vel laminae tenuiter lanceolatae vel etiam lineari- 

lanceolatae var. 7 leptomera. 

Foliorum segmenta vel laminae latiores. 

Folia petiolis adjectis tantum circ. 2.5-4 cm. longa. 

var. 8 holotricha. 
Folia 4-17 cm. longa. 

Folia utrinque glabrata C. mutica sensu stricto. 

Folia infra dense pubescentia var. /3 subvillosa. 

Fruticosa, plus minusve glabra, 0.3-1.3 (etiam usque ad 2) m. 
alta. Folia petiolata petiolis anguste alatis 0.6-3 cm. longis, petiolo 
adjecto 4-17 cm. longa, simplicia vel trisecta vel saepe tripartita, 
lamina segmentisve oblonge lato-lanceolatis vel lanceolato-ovatis 
(terminali apice saepe acuminato) utrinque glabratis vel inferne 
praecipue ad venas sparsim hispidis, acriter serratis, superne saepe 
subcoriaceis. Capitula paniculato-corymbosa, nunc pauca (5-15 
pro unica inflorescentia) nunc numerosa (usque ad circ. 60), tenuiter 
pedicellata pedicellis saepe glabris 1-4 cm. longis, radiata, pansa 
ad anthesin 2-5 cm. lata et 8-17 mm. alta. Involucrum saepe 
glaberrimum, bracteis exterioribus circ. 5, nunc oblongis nunc late 
angusteve spathulato-oblanceolatis, apice plus minusve mucronatis, 
5-12 mm. longis, quam interioribus ovato-oblongis vel rotundato- 
ovatis demum saepe dimidio brevioribus. Flores ligulati circ. 5, 
lutei, fertiles, ligula oblongi, tubo pilosi, apice 2-4-denticulati, 1-2 
cm. longi. Paleae glabrae, oblongo-lineares. Florum tubulosorum 



298 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

stylorum rami apice subulati hispidique. Achaenia valde obcom- 
pressa, corpore nigra alls stramineo-brunnea, exteriora late oblanceo- 
lata alia angustiora, glaberrima, 6-13 mm. longa et 1-3 mm. lata, 
apice emarginato calva vel interiora rarissime biaristata. 

Type specimen: Collected by F. W. Keerl at Tlapujahua (Tlapa- 
coya), northern Puebla, Mexico (Bruss.). DeCandolle cited Martius' 
herbarium for his type specimen, but a good-sized and representative 
fragment is in DeCandolle's Prodromus Herbarium (Del.). 

Distribution: State of Hidalgo, Mexico, southeastwardly through 
Oaxaca and Chiapas into Guatemala. 

Specimens examined : Baites, Mexico, 1864 (Gray) ; Gust . Bernoulli 
196, in thickets, Bano de los Padres, Guatemala, November, 1865 
(N.Y.) ; Cassiano Conzattii 1431 , alt. 1,650 meters, Cerro San Antonio, 
State of Oaxaca, Mexico, June 26, 1906 (Gray) ; Ehrenberg 354 pro 
parte, Mexico (Pom.); Galeotti 2086, Cordilleras of Mexico (Gray, 
fragment; one of two types of Electro, Galeottii A. Gray); Ghies- 
bregkt 133, State of Chiapas, Mexico (Gray); idem 539, growing 5-6 
ft., cold and temperate region, November, 1864-1870 (Gray; Mo.); 
Sutton Hayes, large shrub, near Amatitlan, Department of Amatitlan, 
Guatemala, July 20, 1860 (Gray); Heyde & Lux 3792, alt. 2,400 
meters, Laguna de Ayarza, Department of Jalapa, Guatemala, 
1892 (Berl.; Gray; N.Y.); E. W. D. Holway 5, alt. 1,500 meters, 
Guatemala City, Guatemala, January 1, 1915 (Gray) ; idem 52, alt. 
2,100 meters, San Rafael, Department of Guatemala, Guatemala, 
January 9, 1915 (Gray); idem 154, alt. about 1,950 meters, Solola, 
Guatemala, January 30, 1915 (Gray); idem 613, Guatemala City, 
December 21, 1916 (Gray); W. A. Kellerman 6296 , alt. 1,465 meters, 
San Rafael, Department of Guatemala, Guatemala, February 3, 1907 
(Field) ;C. G. Pringle 8218, alt. 2,850 meters, Sierra de Pachuca, State 
of Hidalgo, Mexico, September 14, 1899 (Berl.; Field; Gray; Mo.; 
N.Y. ; Pom.) ; idem 13041 , alt. 2,490 meters, dry, calcareous bluffs near 
Metepec Station, State of Hidalgo, September 20, 1904 (Berl. ; Field ; 
Gray); C. A. Pur pus 1550, sunny, rocky slopes, Pachuca, State of 
Hidalgo, September, 1905 (Field; Gray; Mo.; N.Y.); J. N. Rose, 
J. H. Painter, & J. S. Rose 8837, near Tulancingo, State of Hidalgo, 
July 22, 1905 (Gray; Mo.; N.Y.; U.S.); Osbert Salvin, alt. 1,590 
meters, Volcan del Fuego, Guatemala, August, 1873 (Berl.); H. Von 
Tuerckheim II. 2043, alt. 1,400 meters, San Cristobal, Department 
of Alta Verapaz, Guatemala, December, 1907 (Field, 2 sheets; Gray; 
Mo.; N.Y.). 



REVISION OF THE GENUS COREOPSIS 299 

Gray wrote, concerning material referred here to this one species 
(PL Wright. 110. 1852): "We appear to have two species, which, 
however, are not as well marked as could be desired, viz.: 1. Electro, 

Mexicana 2. E. Galeottii." Gray's use of "foliis lato- 

lanceolatis" for Electro, mexicana indicates that he mistook simple, 
broad-leaved forms of Coreopsis mutica (E. Galeottii) for DeCandolle's 
Electro, mexicana, which latter had narrow leaves and would best be 
designated today as a variety (vide C. mutica var. 7 leptomera, infra). 

Coreopsis mutica var. subvillosa DC. Prodr. 5: 571. 1836. 
C. mexicana var. hyperdasya Blake, Proc. Amer. Acad. 49: 338. 1913. 

Folia infra ubique dense pubescentia, supra venis exceptis glabrata 
vel scabriuscula, caulibus et gemmis et inflorescentia piloso-tomen- 
tosis lente subglabratis. 

Type specimen: Collected by Wilhelm Friedrich von Karwinski 
in Mexico (Mun.). 

Distribution: State of Hidalgo and southeastwardly through 
Puebla into State of Oaxaca, Mexico. 

Specimens examined: C. Conzatti 1955, alt. 1,700 meters, Alturas 
de San Pablo Hintzo, State of Oaxaca, August 25, 1907 (Field) ; idem 
2074, alt. 1,900 meters, Las Sedas, State of Oaxaca, October 20, 1907 
(Field) ; idem & V. Gonzalez 545 et 546, alt. 2,000 meters, Cerro de San 
Felipe, State of Oaxaca, September 1, 1897 (Gray) ; Ehrenberg 354 pro 
parte, between Mineral del Monte and Pachuca, State of Hidalgo 
(Berl., 2 sheets); E. W. Nelson 1718, alt. 1,800-2,250 meters, near 
Reyes, State of Oaxaca, October 17, 1894 (Gray) ; Nicolas, State of 
Puebla, October, 1905 (Field) ; C. G. Pringle 4896, alt. 1,800 meters, 
ravines of hills near City of Oaxaca, State of Oaxaca, September, 
1894 (Field; Gray, 2 sheets; Mo.; N.Y.; Pom.; U.S.); Charles L. 
Smith, alt. 2,100-2,400 meters, Sierra de San Felipe, State of Oaxaca, 
October 11, 1894 (Mo.; N.Y.); Lucius C. Smith 25, San Juan del 
Estado, State of Oaxaca, June 18, 1894 (Gray); idem 808, alt. 
2,130 meters, La Carbonera, State of Oaxaca, September 20, 1895 
(Gray). 

Coreopsis mutica var. 7 leptomera Sherff, Bot. Gaz. 88: 300. 
1929. Electro, mexicana DC. Prodr. 5: 630. 1836. C. mexicana 
(DC.) Hemsl. Biol. Centr. Amer. Bot. 2: 196. 1881. C. chrysantha 
Sess & Moc. Fl. Mex. ed. 2. 194. 1894 (non alior.). Silphium 
ternatum Sess & Moc. op. cit., 195. 

Glabrata; e specie foliorum glabrorum laminis vel segmentis 
tenuiter lanceolatis vel etiam lineari-lanceolatis differt. 



300 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: Collected by Cyrus Guernsey Pringle, No. 9895, 
at altitude of 2,040 meters, clay banks, Dublan, State of Hidalgo, 
Mexico, October 15, 1902 (Field). 

Distribution: State of Guanajuato eastwardly through Quere*taro 
into State of Hidalgo and thence southward into Federal District, 
Mexico. 

Specimens examined: commun. Alfredo Duges sub num. 472, 
Guanajuato, State of Guanajuato, 1895 (Gray); Hort. Hulier 8, 
December 5, 1866 and 101, January 9, 1867 (N.Y.); Mendez, about 
Villalpando beyond City of Guanajuato, State of Guanajuato (Gray; 
cotype oi Electro, mexicana DC.); Pringle 9895 (type, Field; cotypes, 
Berl.; Gray; Mo.; N.Y.; U.S.); idem 13547, growing 5 to 6 feet high 
at alt. 2,550 meters, barranca above Santa Fe, Federal District, 
September 1, 1905 (U.S.); C. A. Purpus 1339, Ixmiquilpan, State of 
Hidalgo, September, 1908 (Berl.); J. N. Rose, J. H. Painter, & 
J. S. Rose 9717, near Cadereyta, State of Quere"taro, August 22, 
1905 (Gray; N.Y.; U.S.); Uhde 627, Mexico. 

DeCandolle's Electro, mexicana came from Villalpando, near the 
City of Guanajuato, and was a form with simple, narrow leaf blades. 
The specimen sent by Duges from the same general locality had 
mainly tripartite leaves, a distinction of no diagnostic value in this 
species, and, what was indeed significant, the segments were likewise 
narrow. A study of these and other collections shows that the form 
with narrow leaf blades or leaflets constitutes a rather well-marked 
variety occupying a range of greater western but lesser southern 
extent than does the species proper. 

Coreopsis mutica var. 5 holotricha Blake, Proc. Amer. 
Acad. 52: 55. 1917. C. mexicana var. hyperdasya f. holotricha Blake, 
op. cit. 49: 338. 1913. 

Folia parva circ. 2.5-4 cm. longa, utrinque cinerea pube densa 
subscabra. 

Type specimen: Collected by Carl Albert Purpus, No. 3099, 
vicinity of San Luis Tultitlanapa, State of Puebla, Mexico (Gray). 

Distribution: Known only from type locality in State of Puebla, 
Mexico. 

Specimens examined: Purpus 3099 (type, Gray; cotypes, Berl.; 
Field; Mo.; U.S.). 

2. Coreopsis cuneifolia Greenm., Proc. Amer. Acad. 40: 43. 
1904. 



REVISION OF THE GENUS COREOPSIS 301 

Fruticosa, verisimiliter 5-9 dm. alta, caulibus rubris griseisve 
aegre angulatis, ramulis primo antrorsum subadpresso-pubescentibus 
demum nodis exceptis glabratis. Folia sessilia vel subpetiolata, 
simplicia, cuneata vel oblanceolato-cuneata, dimidio superiore 
dentibus patentibus mucronatisque paucidentata, infra medium 
sensim cuneateque usque ad basim angustata, utrinque glabra vel 
infra parce pubescentia, glandulo-punctata, 1.5-5 cm. longa et 
0.6-2 cm. lata. Capitula solitaria vel paniculato-cymosa pedunculis 
tenuibus saepe 1-2 dm. longis, radiata, pansa ad anthesin 1-2.5 cm. 
lata et 8-11 mm. alta. Involucri glabri bracteae exteriores circ. 5, 
lineari-oblongi, adpressae vel parce patentes, 2-5 mm. longae; 
interiores oblongo-ovatae, 5-8 mm. longae. Flores ligulati 5 vel 6, 
fertiles, flavi, ligula late oblongi vel cuneato-obcordati, apice nunc 
vix nunc profunde dentati, 0.6-1.3 cm. longi, tubo brevi parce 
pubescenti. Paleae glabrae, apice obtusae. Florum tubulosorum 
(circ. 12) stylorum rami terminaliter angustato-appendiculati. 
Achaenia valde obcompressa, circumambitu naviculoidea, glaber- 
rima, 7-10 mm. longa et 3-4 mm. lata, fusca vel alis viridi-brunnea, 
nitida, apice emarginato calva. 

Type specimen: Collected by Joseph Nelson Rose, No. 2344, 
State of Durango, Mexico, August 16, 1897 (Gray). 

Distribution: State of Durango southward into State of Jalisco, 
Mexico. 

Specimens examined: C. R. Barnes & W. J. G. Land 165, alt. 
1,680 meters, Sierra de San Esteban, State of Jalisco, September 28, 
1908 (Field); iidem 313, alt. 1,800 meters, dry hills along road to 
Santo Domingo Mine, Etzatlan, State of Jalisco, October 6, 1908 
(Berl.; Field); C. G. Pringle 8781, dry, rocky mountains above 
Etzatlan, State of Jalisco, October 2, 1903 (Field; Gray; N.Y.); 
idem 11900, alt. 1,800 meters, Sierra de San Esteban, near Guada- 
lajara, State of Jalisco, October 21, 1903 (Berl.; Field; Gray); 
Rose 2344 (type, Gray; cotype, U.S.). 

3. Coreopsis parvifolia Blake, Proc. Amer. Acad. 49: 338. 1913. 

Fruticosa, juventate adpresse pubescens denique glabrata, veri- 
similiter 5-9 dm. alta. Folia subpetiolata, rigida, parva, tantum 
circ. 1-2 cm. longa, ovalia, supra adpresse pubescentia infra paulum 
crinita vel glabrata, supra partem inferiorem integram late cuneatam 
utroque circ, 5-mucronato-dentata, summa imminuta subintegra. 
Capitula solitaria ramos elongatos superne subnudos terminantia, 
radiata, pansa ad anthesin 2.5-3 cm. lata et 1-1.5 cm. alta. Involu- 



302 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

cri bracteae exteriores circ. 5, subcrinitae, oblongo-spathulatae, 
obtusae, 4-6 mm. longae; interiores oblongo-ovatae, obtusae, apice 
fimbriatae, 8-11 mm. longae. Flores ligulati circ. 5, lutei, ligula 
ovales vel oblongi, apice denticulati, circ. 1.2-1.5 cm. longi. Paleae 
lineari-oblongae, nitidae. Flores tubulosi 4-dentati, stylorum ramis 
apice tenuiter hispido-caudatis. Achaenia immatura plana, oblance- 
olata, nitida, alato-marginata, glaberrima, apice calvo emarginata, 
8 mm. longa et 2.2 mm. lata. 

Type specimen: Collected by Carl Albert Purpus, No. 2581, on 
dry, rocky hillsides, Esperanza, State of Puebla, Mexico, August, 
1907 (Gray). 

Distribution: Known only from type locality in State of Puebla. 
Mexico. 

Specimens examined: Purpus 2581 (type, Gray; cotypes, Field; 
U.S.); idem 2581a, eodem loco, October, 1911 (Berl.). 

4. Coreopsis Buchii (Urb.) Blake, Contr. U. S. Nat. Herb. 
22: 641. 1924. Selleophytum Buchii Urb. in Fedde, Repert. Sp. Nov. 
13: 484. 1914. 

Fruticosa, erecta, glabra, 3-10 dm. alta, ramis teretibus obscure 
brunneo-rubris plus minusve pruinatis, internodiis 2-4 cm. longis. 
Folia opposita, sessilia, 4-7 cm. longa et 1-1.5 cm. lata, anguste 
lanceolato-oblonga, glabra, subcoriacea, pallide viridia, subnitida, 
margine integra sed interdum aegerrime undulata, basi lata leviter 
cordata, supra medium sensim angustata, apice acuta, nervo medio 
supra prominente, lateralibus utroque latere 12-16, minoribus inter- 
mixtis, utrinque reticulato-conjunctis et prominentibus. Capitula 
pauca (solitaria vel terna umbellata), radiata, recte pedunculata 
pedunculis circ. 4-6 cm. longis, in juventute globosa nigrescentiaque, 
fructifera circ. 2 cm. lata et circ. 1.5 cm. longa. Involucri glabrati 
bracteae exteriores 4-5, laxe accumbentes, subcoriaceae, e basi circ. 
2-2.5 mm. lata sensim angustatae, apice indurate subacutae, tergo 
tenuiter vel obsolete 3-5-nervatae, circ. 8-10 mm. longae; interiores 
lanceolato-oblongae, inferne coriaceae superne tenuiores, circ. 12-13 
mm. longae et circ. 3-4 mm. latae, apice obtusiusculae. Flores 
ligulati flavi, tubo 2.5 mm. longi et (extrinsecus) pilosuli, limbo 8 
mm. lati (apice non viso) tergo breviter pilosi. Paleae apice acumi- 
natae, pleraeque 3-nervatae, circ. 11-12 mm. longae et circ. 1-1.5 
mm. latae. Achaenia oblongo-linearia, plano-obcompressa vel sub- 
tetragona, griseo-brunnea, exalata, apice et marginibus et minus 
costa interiore breviter denseque setosa, unaquaque facie 8-sulcu- 
lata, corpore marginalia 7 mm. longa et usque ad 1.3 mm. lata alia 



REVISION OF THE GENUS COREOPSIS 303 

usque ad 10 mm. longa et 0.6-0.7 mm. lata, apice biaristata aristis 
filiformibus antrorsum hispidis circ. 4-5 mm. longis, 1-2 squamellis 
interdum interjectis. 

Type specimen: Collected by WilhelmBuch, No. 1137, at altitude 
of 1,800 meters, in open forests, Morne la Selle, Haiti, January 
(Berl.). 

Distribution: Known only from Haiti. 

Specimens examined: Emery C. Leonard 4103, on cliff near Fond 
Parisien, Etang Saumatre, Haiti, May 5-13, 1920 (U.S.). 

5. Coreopsis cyclocarpa Blake, Proc. Amer. Acad. 49: 339. 
1913. Leptosyne mexicana A. Gray in Wats. Proc. Amer. Acad. 
22: 429. 1887. 

Herba perennis, 6-7 dm. alta; caulibus pluribus e radice lignea, 
infra parce pubescentibus et 1-2 mm. crassis. Folia opposita, 
integra vel prope medium rarissime trilobata, lamina (vel segmentis) 
lineari-filiformia, basi ciliata, apice acuta, 2-6 cm. longa. Capitula 
corymbose disposita, non numerosa, tenuiter pedunculata pedunculis 
glabris vel superne sparsim hispidis plerumque 2-7 cm. longis, pansa 
ad anthesin 1.5-2.5 cm. lata et circ. 6-8 mm. alta. Involucri bracteae 
exteriores 5 vel 6, glabrae, late lineari-oblongae vel ovato-lanceolatae, 
crassiusculae, apice acutae vel mucronatae, 2-3 (-3.5) mm. longae; 
interiores ovatae vel ovato-lanceolatae, glabratae vel apice fimbriato- 
ciliatae, 5-7 mm. longae. Flores ligulati 8-10, flavi, fertiles, ligula 
oblongi, apice subprofunde dentati, circ. 1.1-1.3 cm. longi. Paleae 
oblanceolatae, apice rotundato-obtusae, 4-5 mm. longae. Flores 
tubulosi exanulati, styli ramis apice incrassatis breviter appendi- 
culatis. Achaenia valde obcompressa, epapposa; extima (ex floribus 
ligulatis) obovata, purpureo-nigrescentia, anguste alato-marginata, 
circ. 3.5 mm. longa et circ. 2.2-2.8 mm. lata; alia minora. 

Type specimen: Collected by Edward Palmer, No. 568, along a 
rivulet in a grassy bottom, Rio Blanco, State of Jalisco, Mexico, 
September, 1886 (U.S.). 

Distribution: State of Jalisco, Mexico. 

Specimens examined: Palmer 568 (type, U.S.; cotype, Gray); 
C. G. Pringle 3570, State of Jalisco, September 10, 1890 (Gray); 
idem 3841, by streams, etc., near Guadalajara, State of Jalisco, 
September 24, 1891 (Berl.; Field, 2 sheets; Gray; N.Y.); idem 
11546, alt. 1,500 meters, dry bluffs near Guadalajara, State of 
Jalisco, October 4, 1903 (Field; Gray); J. N. Rose & J. H. Painter 
7480, near Guadalajara, September, 1903 (N.Y.). 



304 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

6. Coreopsis pinnatisecta Blake, Proc. Amer. Acad. 49: 339. 
1913. Leptosyne Pringlei Robins. & Greenm. Amer. Journ. Sci. 
3. 50: 155. 1895. 

Perennis, e radice brevi-oblonga crassiuscula lignosaque, 3-6 dm. 
alta, caulibus pluribus, adscendentibus, simplicibus vel subsimplici- 
bus, infra foliosis supra subnudatis, striato-angulatis, glabratis vel 
infra moderate hispidis. Folia opposita, petiolata petiolis 0.5-1.2 
cm. longis, petiolo adjecto 2-3 cm. longa, subrigida, pinnata, adscen- 
dentia, sparsim hispida; rhachi et (plerumque 5 vel 7) foliolis linea- 
ribus, acriter apiculatis, 0.4-1 mm. latis, foliolis 6-12 mm. longis 
inferioribus sensim longioribus. Capitula solitaria vel 1-4-congregata 
inflorescentia corymbosa, tenuiter pedunculata pedunculis 5-15 cm. 
longis, radiata, pansa ad anthesin 1.5-1.9 cm. lata et circ. 6-7 mm. 
alta. Involucri glabrati bracteae exteriores 5 vel 6, late oblongae, 
supra saepe latiores, apice obtusae, 3-4 mm. longae; interiores 
ovato-oblongae, apice puberulenti subobtusae, circ. 6 mm. longae. 
Flores ligulati circ. 6, flavi, ligula oblongi, apice 3-denticulati dente 
mediano minore, fertiles, circ. 9-11 mm. longi. Paleae glabrae, 
lineari-oblongae, apice subobtusae. Flores tubulosi 5-dentati, infra 
anulo hispido instructi, stylorum ramis apice incrassatis breviter 
appendiculatis. Achaenia obovata, atra, glabra, circ. 4 mm. longa 
et circ. 1.3 mm. lata, apice epapposo rotundata atque emarginata. 

Type specimen: Collected by Cyrus Guernsey Pringle, No. 4871, 
at altitude of 2,100 meters, on the Sierra de San Felipe, State of 
Oaxaca, Mexico, August 7, 1894 (Gray). 

Distribution: State of Oaxaca and reaching northward barely 
into State of Puebla, Mexico. 

Specimens examined: Pringle 4871 (type, Gray; cotypes, Berl.; 
N.Y.; Pom.; U.S.); C. A. Purpus 4098, Cerro de Paxtle, vicinity 
of San Luis Tultitlanapa, State of Puebla, September, 1909 (Field; 
Gray; N.Y.). 

7. Coreopsis insularis (Brandeg.) Blake, Proc. Amer. Acad. 
49: 340. 1913. Leptosyne insularis Brandeg. Erythea 7: 5. 1899. 

Herbacea, probabiliter perennis, decumbens et diffuse ramosa, 
fere glabra, forsitan 3-6 dm. alta. Folia opposita, petiolata petiolis 
6 mm. longis, petiolo adjecto 1-2.5 cm. longa, pinnatim plerumque 
3- (rarius 5-) lobata, lobis anguste linearibus 0.4-1 mm. latis. 
Capitula pauca, tenuissime pedunculata pedunculis glabris saepius 
0.8-2 dm. longis, radiata, pansa ad anthesin circ. 1.3 cm. lata et circ. 
5-6 mm. alta. Involucri bracteae exteriores circ. 5 vel 6, lineares 



REVISION OF THE GENUS COREOPSIS 305 

vel lineari-lanceolatae, subglabrae, apice obtusae vel rotundatae, 
circ. 2.5-3.5 mm. longae; interiores ovatae vel oblongo-acuminatae, 
4-5 mm. longae. Flores ligulati 5, flavi, fertiles, ligula elliptico- 
oblongi, apice 2-lobulati, circ. 7 mm. longi. Paleae lineares. Flores 
tubulosi minute 2-5 mm. longi, exanulati, stylorum ramis termino 
anguste elongato-conicis. Achaenia extima (ex floribus ligulatis 
orta) valde obcompressa, elliptico-obovata, late alata, glaberrima, 
3.5-4.2 mm. longa, epapposa; interiora angustiora, anguste marginata. 

Type specimen: Collected by A. W. Anthony, No. 394, on Socorro 
Island, off coast of Baja California, Mexico, March-June, 1897 
(Calif.). 

Distribution: Known only from Socorro Island, off coast of Baja 
California. 

Specimens examined: Anthony 394 (type, Calif.; cotypes, Gray; 
U.S.); F. S. Bachelew 223, Socorro Island, May 27-July 3, 1903 
(Gray; N.Y.). 

8. Coreopsis maritima (Nutt.) Hook. f. Bot. Mag. pi. 6241. 
1876. Tuckermannia maritima Nutt., Trans. Amer. Phil. Soc. 2. 
7: 363. 1841. Leptosyne maritima (Nutt.) A. Gray, Proc. Amer. 
Acad. 7: 358. 1868; etiam Noblet, Rev. Hort. 1873: 330 et pi. (sine 
num.). 1873. L. maritima var. gigantea Hort. ex Noblet, loc. cit. 
Leptogyne maritima (Nutt.) A. Gray ex Hook. f. loc. cit. (sphalm). 

Caules carnoso-herbacei, e radice crassa profunda recta sublig- 
neaque, patentes, ramosi, glabri, 3-8 dm. alti. Folia nunc alterna 
nunc opposita, petiolata petiolis latis vittatis striatis basi dilitatis 
1-7.5 cm. longis, petiolo adjecto saepius 1-1.5 cm. longa, 2-3- 
pinnata, segmentis oblonge linearibus, planis, obtusis, 1-2.5 mm. 
latis et usque ad 3 cm. longis. Capitula non numerosa, speciosa, 
elongate pedunculata pedunculis robustis glabris plerumque 1.5-4 
dm. longis, radiata, pansa ad anthesin 6.5-8 (raro usque ad 13.5) 
cm. lata et 1.5 cm. alta. Involucri bracteae exteriores 6-8, oblongae 
vel ovato-lanceolatae, obtusae, glabrae vel apice minutissime pubes- 
centes, 1-2 cm. longae; interiores oblongae, apice pubescentes, saepius 
paulo longiores. Flores ligulati 14-20 (plerumque 16-18), flavi, 
ligula elliptico-oblanceolati, apice circ. 3-denticulati, 2-3.5 (-5.5) 
cm. longi. Paleae glaberrimae, oblongo-lineares, apice acutae, 
demum circ. 1-1.2 cm. longae. Florum tubulosorum stylorum rami 
abrupte conico-incrassati vix appendiculati, anulo glabro. Achaenia 
anguste oblongo-obovata, plana, nigra nisi alis atro-brunneis, circ. 
6-7 mm. longa et alis inclusis 2-3.5 mm. latis, apice glabra vel 
interdum alis productis breviter 1 -2-aristata. 



306 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: Collected by Edward Tuckerman, Jr., on shelving 
rocks, near sea, San Diego, California. Type material not examined, 
perhaps still extant among the Nuttall plants (Brit., Kew, or Phila.). 

Distribution: South westernmost California; rare northwest- 
wardly as far as Monterey County and southwardly in Baja (Lower) 
California as far as All Saints' Bay and San Martin Island. 

Specimens examined: A. W. Anthony 216, San Martin Island, 
Baja (Lower) California, March-June, 1897 (Field; Gray; Pom.); 
Margaret Armstrong 741, San Diego, California (N.Y.); T. S.Brande- 
gee, San Martin Island, Baja California, April, 1897 (Pom.); idem, 
San Diego, California, June, 1900 (Pom.); idem, eodem loco, May, 
1906 (N.Y.); Harley P. Chandler 5123, La Jolla, California, April 
16, 1904 (N.Y.);F. E. &E. S. Clements 226, cliff edge, dune strand, 
eodem loco, March 14, 1914 (Field; Gray; N.Y.); D. Cleveland, San 
Diego, California (Gray); R. B. Cowles, moist slope, Coronado 
Islands, Baja California, March 29, 1921 (Pom.); G. W. Dunn, 
eodem loco, April 17, 1891 (Field) ; Alice Eastwood 2565, Point Loma, 
California, April 6, 1913 (Gray) ; Miss F. E. Fish, All Saints' Bay, 
Baja (Lower) California, May, 1882 (Gray) ; G. L. Fleming & W. W. 
Eggleston 19676, alt. 50 meters, Torrey Pines Reservation, San Diego 
County, California, April 10, 1924 (Gray; Pom.); Hort. Bot. Harvard 
Univ. cult, annis 1851 et 1852 e sem. a Fremontio lect. (Gray) ; Marcus 
E. Jones, Todos Santos Bay, Baja California, April 11, 1882 (Pom., 
2 sheets) ; idem, ranch 29 miles southwest of Tia Juana, Baja Cali- 
fornia, April 13, 1925 (Pom., 2 sheets); idem, Coronado Islands, 
Baja California, June 10, 1926 (Pom.); idem 3134, Soledad, Cali- 
fornia, March 29, 1882 (N.Y., 2 sheets; Pom., 2 sheets); A. J. 
McClatchie, San Diego, California, April, 1893 (N.Y.); Philip A. 
Munz 7955, sea bluffs, Cardiff, California, May 9, 1924 (Gray; Pom.) ; 
C. R. Orcutt, Point Loma, California, March 24, 1884 (Field) ; idem 
& W. S. Bowne, San Diego, 1884 (N.Y.) ; C. C. Parry, eodem loco, 
1850 (N.Y., 2 sheets, one from the Torrey Herbarium and constitut- 
ing the basis of a drawing by Riocreux); idem, eodem loco, 1883 
(Gray) ; C. G. Pringle, bluffs of the seashore, eodem loco, April 25, 
1882 (Field); F. M. Reed 6147, Potrero Canyon, Baja California, 
February 5, 1929 (Pom.); M. S. Snyder, seashore, La Jolla, Cali- 
fornia, February 25, 1895 (Field, 2 sheets) ; Mary F. Spencer 86, in 
beach sand, vicinity of San Diego, California, April 16, 1916 (Gray; 
N.Y.); H. Strauss, San Diego and nearby islands, September 24, 
1908 (Berl.) and February 3, 1912 (Berl.) ;L. Street, San Elejo Lagoon, 
Baja California, May 12, 1917 (Pom.). 



REVISION OF THE GENUS COREOPSIS 307 

9. Coreopsis gigantea (Kell.) H. M. Hall, Univ. Calif. Publ. 
Bot. 3: 142. 1907; cf. S. B. Parish, Muhlenbergia 8: 133-134. 1913; 
etiam C. F. Millspaugh, Field Mus. Bot. 5: 284, pi. 9 and pi. 13, f. 1. 
1923. Leptosyne gigantea Kellogg, Proc. Calif. Acad. 4: 198. 1872. 
Tuckermannia gigantea (Kell.) Jones, Contrib. West. Bot. No. 15: 
74. 1929. 

Perennis, erecta, e radice magna plus minusve lignosa, caule 
robusto carnoso vix lignoso 5-12.5 cm. crasso et plerumque 3-12 
rarius usque ad 20 rarissime usque ad 30 dm. alto, durante per 3-8 
annos, ramis glabris, primariis distantibus horizontalibus vel adscen- 
dentibus terminos versus foliosis. Folia saepius alterna, petiolata 
petiolis glabris carnosis basi plus minusve dilatatis 2-9 cm. longis, 
petiolo adjecto saepius 0.8-2 dm. longa, glabra, plerumque 2-3- 
pinnata, segmentis nunc moderate (0.8-1.3 mm. latis) nunc 
angustissime (0.25-0.4 mm. latis) lineari-filiformibus, integris vel 
lateraliter subemarginatis, saepe 3-nervatis, usque ad +5 cm. 
longis. Capitula pedunculata pedunculis (ramulis ultimis) sub- 
robustis stramineis nudis vel subnudis corymbose congregatis 
plerumque 1-2 dm. longis, plerumque radiata, pansa ad anthesin 
4-8 cm. lata et circ. 10-13 mm. alta. Involucri glabri bracteae 
exteriores 5-7, oblongo-lineares vel lineari-lanceolatae, apice trun- 
catae, 1-2 cm. longae, quam interiores 12-15 ovato-oblongae vel 
obovatae apice pubescentes nunc paulo breviores nunc vix longiores. 
Flores ligulati 10-16, flavi vel interdum apice pallido-flavidi, ligula 
anguste vel late oblanceolati, apice circ. 3-dentati, 2-3.8 cm. longi. 
Paleae oblongo-oblanceolatae, glaberrimae, apice obtusae, demum 
7 mm. longae. Florum tubulosorum stylorum rami abrupte 
incrassati non appendiculati, anulo setis capitatis hispido. Achaenia 
subplana, obovato-oblonga vel oblanceolato-oblonga, glabrata, nigra 
vel marginibus latis subincrassatis atro-brunnea, omnino (alis 
inclusis) 5.5-6.8 mm. longa et 2.5-3.5 mm. lata, apice calva vel alis pro- 
ductis obsolete bidentata, saepe poculo minutissimo centrali coronata. 

Type specimen: Collected by William George Washington Har- 
ford (under auspices of Captain S. Forney, United States Coast Sur- 
vey}, near Cuyler Harbor, San Miguel Island, about 40 miles off 
coast of Santa Barbara, California. 

Distribution : San Luis Obispo to Ventura counties, southwestern 
California; also on islands off shore and extending southwardly 
as far as Guadalupe Island, off coast of Baja (Lower) California. 

Specimens examined (all from California or islands nearby except 
where otherwise noted [Palmer 41])' Anon., mouth of Santa Maria 



308 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

River, February 7, 1880 (Gray) ; T. S. Brandegee, Island Santa Cruz, 
April, 1888 (Gray); Dr. Coulter 316, San Miguel to Santa Barbara 
(Gray); M. B. Dunkle 1857, alt. 4.5 meters, shallow soil on rock, 
Bird Rock, Santa Catalina Island, April 11, 1928 (Pom.); Alice 
Eastwood, Point Sal (near boundary between Santa Barbara and 
San Luis Obispo counties), May 12, 1896 (Gray); W. W. Eggleston 
19577, alt. 30 meters, Concepcion to Jalama, Santa Barbara County, 
March 29, 1924 (Gray; Pom.); A. D. E. Elmer 3634, Surf, Santa 
Barbara County, May, 1902 (Field; Gray; Pom.); Roxana S. Ferris 
7572, ocean side of sand dunes, 5 miles south of Surf, Santa Barbara 
County, April 4, 1929 (Pom.); Grant & Wheeler 680 and 2228, 
Catalina Island, April 21-26, 1904 (Field; N.Y.; Pom.); John 
Thomas Howell 8220, San Nicolas Island, March 13, 1932 (Pom.); 
Marcus E. Jones, Santa Cruz Island, April 13, 1924 (Pom., 2 sheets) ; 
idem, Malibu Hills, north of Santa Monica Mountains, April 26, 
1926 (Pom.); idem, Santa Cruz Island, March 25, 1929 (Pom.); 
Philips Mills Jones, Santa Rosa Island, April, 1901 (Calif. };Ezra C. 
Knopf 56, in rocky soil on hillsides, Pebble Beach Road, Santa 
Catalina Island, March 26, 1921 (Field) ; idem 258, Bird Rock, Santa 
Catalina Island, November 20, 1921 (Field) ; C. F. Millspaugh 4630, 
eodem loco, January 22, 1920 (Field); Philip A. Munz 11378, sand 
dunes, Surf, May 17, 1929 (Pom.); idem & E. Crow 11553, Santa 
Cruz Island, April 7, 1930 (Pom.); iidem 11691, seedlings on shaded 
wall of Water Canyon, Santa Rosa Island, April 8, 1930 (Pom.); 
iidem 11750, shaded canyon wall northwest of ranch house, Santa 
Rosa Island, April 9, 1930 (Pom.); Munz & J. Voss 11877, high 
exposed cliff, San Miguel Island, April 10, 1930 (Pom.); L. W. 
Nuttall 571, on sea cliffs, near Avalon, Santa Catalina Island, May 
14, 1920 (Field); Edward Palmer 41, Guadalupe Island, off Baja 
(Lower) California, 1875 (Berl.; Gray); Parry, San Diego (Gray); 
Miss Plummer, mountains above Santa Barbara, 1878 (Gray); 
Huron H. Smith 5034, road to Pebble Beach, Santa Catalina Island, 
May 30, 1912 (Field) ;Blanch Trask 76, San Nicolas Island, April, 1901 
(Gray, 2 sheets) ; W. G. Wright, Ventura County, March, 1894 (Field). 

10. Coreopsis Bigelovii (A. Gray) Voss in Sieb. & Voss, Vil- 
morin Blumeng. ed. 3. 1: 488. 1894 (where spelled C. Bigelowii). 
Pugiopappus Bigelovii A. Gray, Pacif. R. Rept. 4: 104. 1857. 
P. Breweri A. Gray, Proc. Amer. Acad. 8: 660. 1873. Leptosyne 
Bigelovii A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 300. 1884. L. Hamiltonii 
Elmer, Bot. Gaz. 41: 323. 1906. Coreopsis Bigelovii (A. Gray) 
H. M. Hall, Univ. Calif. Publ. Bot. 3: 141. 1907. 



REVISION OF THE GENUS COREOPSIS 309 

Herba annua, erecta, 1-4 dm. alta, caulibus saepius pluribus vel 
numerosis, simplicibus vel inferne ramosis, basim versus foliosis, 
supra multum elongatis ac nudis pro pedunculis, glabris. Folia 
saepe dense congregata, plus minusve alterna, petiolata petiolis 
usque ad 6 cm. longis, petiolo adjecto 5-11 cm. longa, plerumque 
1-2-pinnata, segmentis ultimis linearibus, planis, obtusiusculis, sub- 
membranaceis, glaberrimis vel rarius vix scabrido-hispidis, plerum- 
que 0.6-1.7 mm. latis. Capitula caules (pedunculos) terminantia, 
radiata, pansa ad anthesin 2-4.2 cm. lata et 0.8-1 cm. alta. Involucri 
glabri bracteae exteriores circ. 8, lineares, apice obtusiusculae ac 
nitido-induratae, 4-9 mm. longae; interiores oblongo-ovatae, 
moderate longiores. Flores ligulati 8, flavi (interdum infra intense 
supra aegre), ligula elliptico-oblanceolati vel anguste obovati, apice 
dentati, fertiles, 1-2 cm. longi. Paleae lineari-oblongae vel navi- 
culoideae, obtusae vel subacutae, glabrae, apice minute eroso-ciliatae, 
demum circ. 7-9 mm. longae. Flores tubulosi anulo glabri, stylorum 
ramis abrupte terminantibus et vix appendiculatis. Achaenia 
dimorpha exteriora (e floribus ligulatis) valde obcompressa, cuneato- 
obovata, rubro-nigra, glabrata, faciebus valde rugosa, marginibus 
anguste calloso-alata, apice exaristata, 4-5 mm. longa et 2-3 mm. 
lata; interiora (e floribus tubulosis) angustiora, oblongo-linearia, 
faciebus nitidis nigra, tergo glabrata ventre erecte albido-villosa, 
marginibus densissime erecto- et albido-villosa, corpore 4-6.2 mm. 
longa et 1.5 mm. lata, apice biaristata aristis lanceolatis planis 
densissime erecteque albido-ciliatis circ. 2-3 mm. longis, inter aristas 
poculo membranaceo vix manifesto coronata. 

Type specimen: Collected by John Milton Bigelow, Mojave 
Creek, California, 1853-1854 (Gray). 

Distribution : California. 

Specimens examined: R. Bacigalupi 1209, alt. 540 meters, just 
above Oak Grove on the Mineral King Road, Tulare County, April 
23, 1925 (Pom. ; forma elatior) ; Bigelow (type, Gray) ; W. H. Brewer 
241, San Buenaventura, 1860-1862 (Gray; sub nom. Pugiopappo 
Breweri} ; Ira W. Clokey & B. Templeton 4520, alt. 350 meters, dry 
open hillside, Mandeville Canyon, Santa Monica Mountains, March, 
1929 (Pom.) ; Frederick V. Coville & Frederick Funston 742, alt. 800- 
2,000 meters, Shepherd Canyon, Argus Mountains, April 30, 1891 
(Berl.; Field; Gray); Craig, Newsom, & Hiland 80, at 35 miles south 
of Randsburg, Mojave Desert, April 5, 1927 (Pom.); W. R. Dudley 
340, region of Tehachopi Peak, Tehachopi Mountains, June 25, 
1895 (N.Y.); Alice Eastwood, Kaweah, April 27, 1895 (Gray); H. C. 



310 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Ford, Santa Barbara County, 1889 (Gray) ; G. B. Grant 5424, Saugus, 
April 23, 1903 (Pom., 2 sheets) ; H. M. Hall 3074, alt. 1,050 meters, 
vicinity of Elizabeth Lake, Antelope Valley, southern California, 
May 1-3, 1902 (Pom.); A. A. Heller 7662, Kern Canyon, Kern 
County, April 12, 1905 (Field; Gray); C. L. Hitchcock 12340, near 
brush, open desert, 10 miles southwest of Shoshone, April 17, 1932 
(Pom.); Marcus E. Jones, alt. 1,380 meters, Darwin, April 28, 1897 
(Pom.); idem, alt. 1,380 meters, Shepherd's Canyon, Argus, Moun- 
tains, April 30, 1897 (N.Y.; Pom.); idem, alt. 1,800 meters, Pleasant 
Canyon, Panamint Mountains, May 6, 1897 (Pom.); idem, alt. 
1,140 meters, Cajon Pass, May 16, 1903 (Pom., 2 sheets); idem, 
Cave Spring, March 12, 1924 (Pom.); William L. Kennedy, Fort 
Tejon, spring of 1876 (Field) ; J. G. Lemmon, top of Mount Diablo 
(Field); idem 138, Mojave Desert, 1880 (Gray); Philip A. Munz 
4466, alt. 1,140 meters, dry slope near south side of summit, Cajon 
Pass, southern California, April 27, 1921 (Pom.); idem 9136, alt. 
900 meters, dry slope, Tule River, Tulare County, March 22, 1925 
(Pom.); idem 9186, alt. 900 meters, disintegrated lava, north of 
Parkfield, Monterey County, March 24, 1925 (Pom.); idem 10042, 
sandy desert, 5 miles south of Willow Springs, April 4, 1926 (Pom.); 
idem, R. D. Harwood & L M. Johnston 4077, alt. 1,200 meters, 
clearing in chaparral, north side of Cajon Pass, May 25, 1930 (Pom.); 
Munz & D. Keck 7842, gravelly wash, 7 miles east of Daggett, April 6, 
1924 (Pom.); iidem 7879, sandy wash, 10 miles southwest of Garlic 
Springs, Mojave Desert, April 8, 1924 (Pom.); Samuel B. Parish, 
Marengo, April, 1882 (Field) ; idem & W. F. Parish 226, Whitewater 
River, March, 1881 (Berl. ; Field) ; C. C. Parry, Pacific Coast of North 
America, 1880 (Field); idem 115, eodem loco, 1881 (Field); idem & 
J. G. Lemmon 184, southern California, etc., 1876 (Field); C. G. 
Pringle, hills bordering the Mojave Desert, May 11, 1882 (Field; 
Gray); C. A. Purpus 5063, alt. 300-600 meters, sunny hillsides, 
Middle Tule River, April-September, 1897 (Gray); idem 5531, 
alt. 1,200-1,500 meters, hillsides near Erskin Creek, southeastern 
California (Gray); Marjorie Shaw 1941, Upper Cajon Pass, May, 
1917 (Pom.); eadem, Edna Spaulding & Mrs. C. L. Walton, alt. 
900 meters, Antelope Valley, Mojave Desert, April 6-8, 1917 (Pom.); 
Mary F. Spencer 407, summit of Cajon Pass, San Bernardino 
County, April 27, 1917 (Gray; Pom.); eadem 1954, alt. 960 meters, 
Mojave Desert, May 1, 1922 (Gray; Pom.); eadem 2040, rocky 
places, Maillard Canyon, near Banning, April 23, 1922 (Pom.); 
William Trelease, Hesperia, April 10, 1892 (Mo., 2 sheets) ;G. R. Vasey 



REVISION OF THE GENUS COREOPSIS 311 

290, Branch of Mojave River, 1880 (Field; Gray); W. G. Wright, 
San Bernardino, 1880 (BerL); idem 172, Colorado Desert, March, 
1881 (Gray). 

11. Coreopsis Calliopsidea (DC.) A. Gray in Torr. Bot. Mex. 
Bound. 90. 1858. Agarista Calliopsidea DC. Prodr. 5: 569. 1836; 
Regel, Gartenflora 23: 356, pL 816. 1874. Pugiopappus Calliopsidea 
(DC.) A. Gray, Proc. Amer. Acad. 8: 660. 1873; Bot. Calif. 1: 355. 
1876. Leptosyne Calliopsidea (DC.) A. Gray, Syn. Fl. N. Amer. 1, 
pt. 2: 300. 1884. L. Calliopsidea var. nana A. Gray, loc. cit. 

Herba annua, erecta, glabra, 1-6 dm. alta, caule tereti simplici 
vel ramoso nunc usque ad medium (vel ultra) nunc tantum ad basim 
folioso. Folia alterna, petiolata petiolis alatis usque ad 2 (raro -5) 
cm. longis, petiolo adjecto 1-7 cm. longa, 1-2-pinnatifida rhachi 
lobisque planis linearibusque plerumque 0.6-1.7 (raro -5) mm. latis. 
Capitula ramos elongates aphyllos terminantia, solitaria, radiata, 
pansa ad anthesin 2-5 cm. lata et circ. 9-12 mm. alta. Involucri 
bracteae exteriores 4-6, oblongo-ovatae vel saepius ovato-cordatae, 
apice obtusae, angustissime pallido-marginatae, tergo longitudinaliter 
4-13-striatae, 4-7 mm. longae; interiores circ. 8, oblongo-ovatae, 
apice acutae vel pubescentes, 6-13 mm. longae. Flores ligulati 
plerumque 8, flavi, tubo clavo-setosi, ligula late cuneato-obovati, 
apice plus minusve truncati et irregulariter multidenticulati, 1-2.5 
cm. longi. Paleae lineari-oblanceolatae, longitudinaliter nitido- 
striatae striis interrupte brunneis, supra minutissime eroso-ciliatae, 
plerumque 4-7 mm. longae, cum achaenio deciduae, ad faciem 
hujus exteriorem applicatae. Flores tubulosi corolla 5-dentati, 
basim versus clavo-setosi, anulo pubescentes, stylorum ramis termi- 
naliter brevi-conici. Achaenia plana dimorpha, exteriora (ex floribus 
ligulatis) obovato-oblonga faciebus nitidis purpureo-nigrisque glabra 
vel sparsim pulverulenta, margine subanguste alata ala parce in- 
crassata, apice calva, omnino 4.5 mm. longa et 2.2 mm. lata; 
interiora (ex floribus tubulosis) lineari-oblanceolata vel cuneato- 
oblonga, nigra, tergo nitida glaberrimaque, marginibus ventreque 
erecte elongate- villosa, corpore 4-5 mm. longa, apice biaristata 
aristis circ. 3-4 mm. longis densissime antrorso-hispidis. 

Type specimen: Collected by David Douglas in California (Del.). 
Distribution: Southern half of California. 

Specimens examined: Anon., loco ignoto (BerL); Townsend S. 
Brandegee, Alcalde, Fresno County, April, 1891 (Field); Craig, 
Newsom & Hiland 129, Randsburg, April 5, 1927 (Pom.); David 



312 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Douglas, California (Gray); idem (distrib. Herb. Soc. Hort. Lond.), 
California, 1833 (Berl.; doubtless a cotype) ; Alice Eastwood, Cuyama, 
White Hills, May 2, 1896 (Gray) ; eadem 3187, Mojave, Kern County, 
May 12, 1913 (Gray) ; J. B. Feudge 1382, alt. 750 meters, open desert, 
Kramer, Mojave Desert, May 25, 1926 (Pom.); Harvard Univ. 
Bot. Gard., cult., 1871 (Gray); H. E. Hasse, Mojave Station, Kern 
County, March 15, 1889 (Field); A. A. Heller 7723, gravelly hill- 
sides, Sunset, April 20, 1905 (Field; Gray); Ivan M. Johnston, 
rather common, sandy soil, near Mojave, May 5, 1920 (Pom.); 
idem, common, 1 mile south of Crutts Post Office, Mojave Desert, 
May 14, 1922 (Pom.); Marcus E. Jones, Cuyama Canyon, April 28, 
1926 (Pom.); idem, Adelanto, May 12, 1927 (Pom.); idem, alt. 900 
meters, Mojave, May 20, 1903 (Pom.); A. Kellogg & W.G. W. Har- 
ford436, California (Gray) ; J. G. Lemmon (137?), California (Gray; 
type of var. nana A. Gray) ; idem 4520, Lemmon's Ranch, Cholame, 
June, 1887 (Gray); S. B. Parish 9771, Mojave Station, Kern County, 
April 26, 1915 (Gray); Parry, California (Gray); W. M. Pierce, 
desert, 20 miles east of Victorville, April 12, 1922 (Pom.); C. G. 
Pringle, Mojave Desert, May 14, 1882 (Field; Gray). 

12. Coreopsis Douglasii (DC.) H. M. Hall, Univ. Calif. 
Publ. Bot. 3: 140. 1907. Leptosyne Douglasii DC. Prodr. 5: 531. 
1836. L. calif ornica Nutt. Trans. Amer. Phil. Soc. ser. 2, 7: 363. 1841. 
L. Newberryi A. Gray, Proc. Amer. Acad. 7: 358. 1868. 

Herba annua, glabra, 1.5-4 dm. alta, caule brevi erectoque 
ramoso ramis (pedunculis) paucis basim versus foliosis aliter nudis 
vel raro folio valde imminuto instructis, apice monocephalicis. 
Folia (subradicalia) petiolata petiolis plerumque 2-5 cm. longis, 
petiolo adjecto 3-10 (-15) cm. longa, filiformi-linearia integraque 
vel saepius 2-5-partita lobis lineari-filiformibus apice obtusis. 
Capitula (solitaria, elongatissime pedunculata) radiata, pansa ad 
anthesin 2-3.3 cm. lata et 6-8 mm. alta. Involucri glabrati bracteae 
exteriores 5-16, lineari-attenuatae vel lineari-oblongae, apice obtusis- 
simae, 4-8 mm. longae; interiores ovatae, apice acutiusculae, paulo 
longiores. Flores ligulati circ. 8, fertiles, flavi (vel apicem versus 
saepe aibidi), ligula oblongi vel ovato-subrotundati, apice circ. 
3-dentati dentibus saepe magnis rotundatisque, plerumque 1-1.8 
cm. longi. Paleae lineares vel lineari-oblongae, apice obtusae, 
demum 3-6.5 mm. longae. Flores tubulosi anulo plerumque hispido 
instructi, stylorum ramis cono hispido lato brevique terminatis. 
Achaenia obcompressa, cuneato-obovata, diverse rubro-brunnea vel 
nigro-rubra, primo (sed non demum) nitida, marginibus crasso- 



REVISION OF THE GENUS COREOPSIS 313 

alata, duabus faciebus setis clavo minimo similibus instructa, 2-3 
mm. longa et 1.2-1.5 mm. lata, apice exaristata sed poculo minuto 
membranaceoque coronata. 

Type specimen: Collected by David Douglas in California (Del.). 

Distribution: Southern Arizona westward into southern Cali- 
fornia and reaching southward slightly into Baja (Lower) California, 
Mexico. 

Specimens examined (all from California except where noted to 
the contrary) : LeRoy Abrams 3342, Moro Hills, Fallbrook, April 28, 
1903 (Gray); idem 3628, dry hills between Campo and Jacumba, 
May 27, 1903 (Gray; Pom., 2 sheets); idem 11788, sandy areas in 
hills between Rosamond and Mojave, April 30, 1927 (Pom.); idem 
11915, between Red Rock Canyon and Walker Pass Road, El Paso 
Range, May 1, 1927 (Pom.); C. F. Baker 4090, Claremont, April 15, 
1903 (Field; Gray; Pom.); H. N.Bolander, Julian City near Golden 
Cherry Mine (Gray); Katharine Brandegee, San Diego, April 25, 
1900 (Calif.) ; T. S. Brandegee'3368, common on the mesas, San Diego, 
April 26, 1903 (Field; Gray; Pom., 2 sheets);* 1 . E. & E. S. Clements 
224, La Jolla, March 7, 1914 (Field; Gray); iidem 225, eodem loco, 
March 17, 1914 (Calif.; Gray); D, Cleveland, San Luis Rey, 1877 
(Gray); Mrs. A. L. Coombs, Warner's Springs, San Diego County, 
spring of 1911 (Gray) ; Dr. Coulter 339, California (Gray) ; Douglas, 
California (cotype, Gray) ; AliceEastwood 2574, Warner's Hot Springs, 
San Diego County, April 9, 1913 (Gray); eadem 3194, Mojave, Kern 
County, May 13, 1913 (Gray); eadem 4377, Blochman's Ranch, 
Mariposa County, April 21, 1915 (Gray); W. W. Eggleston 19748, 
alt. 850-950 meters, Jacumba Springs, San Diego County, April 
11-16, 1924 (Pom.) ; Roxana S. Ferris 7466, sandy flat between Coso 
Hot Springs and Coso Junction, Kern County, April 30, 1928 
(Pom.); eadem 7710, about 5 miles from Mojave or the Willow 
Springs Road, Kern County, May 3, 1929 (Pom.); George B. Grant 
681, Altadena, April, 1902 (Field, 2 sheets) ; idem 682, eodem loco et 
tempore (Gray); Harvey M. Hall 3973, northwest corner of Baja 
California, Mexico, April, 1903 (Field) ; Harvard University Botan- 
ical Garden, cult., 1882 (Gray) ; H. E. Hasse, Los Angeles, April 22, 
1888 (Field); A. A. Heller 7674, Kramer, April 13, 1905 (Field; 
Gray); C. L. Hitchcock 12224, sandy wash, 10 miles southeast of 
White Tanks, East Riverside County, April 9, 1932 (Pom.) ; Edmund 
C. Jaeger 140, Cabazon, April 11, 1920 (Pom.); Ivan M. Johnston, 
among creosote bushes in open sandy areas, near Mojave, May 5, 
1920 (Pom.); idem, on hillside, Victorville, May 15, 1920 (Pom.); 



314 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

idem, alt. 870 meters, in sandy wash, 5 miles southeast of Victorville, 
May 17, 1920 (Pom.); idem 1251, alt. 600 meters, field above 
Claremont, March 29, 1917 (Pom.); Marcus E. Jones, Pasadena, 
February 22, 1882 (Pom.); idem, Fallbrook, March 25, 1882 (Pom.); 
idem, San Diego, April 3, 1882 (Pom.) ; idem, alt. 300 meters, White- 
water, May 11, 1903 (Pom., 2 sheets) ; idem, alt. 880 meters, Victor, 
May 17, 1903 (Pom., 2 sheets); idem, San Diego, April 14, 1906 
(Pom., 2 sheets); idem, alt. 1,200 meters, Julian, April 17, 1906 
(Pom.); idem, near Santa Ana Canyon, March 10, 1926 (Pom.); idem 
3361 pro parte, Pasadena, May 2, 1882 (N.Y.; Pom., 2 sheets); 
idem 3373, eodem loco et tempore (Berl.); William Kellogg, Rancho 
Encinal, Monterey County, April 15, 1903 (Gray) ; J. G. Lemmon, 
San Bernardino, March, 1880 (Gray) ; Philip A. Munz 2148, foothills, 
Corona, April 27, 1918 (Pom.); idem 2538, alt. 750 meters, desert, 
Barstow, April 12, 1919 (Pom.) ; idem 3814, alt. 300 meters, dry slope, 
Santa Ana River Canyon near Rincon, April 11, 1920 (Pom.); idem 
9357, alt. 480-510 meters, dry bottom of Cobal Canyon, north of 
Claremont (Gray; Pom.); idem 12415, open desert, 20 miles north 
of Box S Ranch, San Bernardino County, April 21, 1932 (Pom.); 
idem & D. Keck 7930, sandy desert 20 miles west of Barstow, April 
10, 1924 (Gray; Pom.); Philip A. Munz & R. D. Harwood 3385, alt. 
360 meters, gravelly wash, Yemescal Canyon, March 20, 1920 (Pom.) ; 
iidem 3475, alt. 900 meters, sand, Victorville, March 28, 1920 
(Pom.); Philip A. Munz, L. Street, & G. Williams 2362, alt. 535 
meters, roadside, San Gorgonio Pass, near Cabazon, March 28, 
1919 (Pom.) ; iidem 2388, eodem loco et tempore (Pom.) ; Newberry, 
Sitgreaves Pass, southern Arizona, March 26 (Gray; type of Lep- 
tosyne Newberryi A. Gray) ; Thomas Nuttall, sine loco vel tempore 
(Gray) ; Charles R. Orcutt, Point Loma, San Diego, March 12, 1884 
(Calif.); idem, northern Baja California, April 17, 1886 (Field); 
Edward Palmer 126, Camp Grant, May 10, 1867 (Gray; the specimen 
cited by A. Gray for his Leptosyne Newberryi); idem 217, Mojave 
River, southeastern California, May, 1876 (Field); idem 677, 
San Quentin Bay, Baja California, January, 1889 (Field; Gray); 
S. B. Parish 2211, Whitewater Canyon, April, 1880 (Gray); idem 
5191, alt. 300-750 meters, vicinity of San Bernardino, April 13, 
1903 (Field); idem 10415, Mojave Station, Kern County, March 
25, 1915 (Gray) ; idem & W. F. Parish, San Bernardino, April, 1881 
(Field); iidem, foothills of San Bernardino Mountains, May, 1888 
(Field); C. C. Parry, San Bernardino (Gray); idem 116 (Gray); 
idem & J. G. Lemmon 185, southern California, etc., 1876 (Field; 
Gray); W. M. Pierce, desert, 20 miles east of Victorville, April 12, 



REVISION OF THE GENUS COREOPSIS 315 

1922 (Pom.); M. S. Snyder, La Jolla, March 15, 1895 (Field); Mary 
F. Spencer 1 72, in hollows of granite rocks, San Diego County, April 
8, 1917 (Gray; Pom.); eadem 563, desert sand, Hesperia, Mojave 
Desert, May 4, 1917 (Gray; Pom.); eadem 653, alt. 750 meters, in 
rocky places, Maillard Canyon, April 22, 1922 (Gray) ; eadem 653a, 
in rocky places, Snow Creek near White Water, Mojave Desert, 
April 10, 1922 (Gray); eadem 2123, alt. 750 meters, in rocky places, 
Maillard Canyon near Banning, April 23, 1922 (Gray); Wallace, 
Los Angeles, 1854 (Gray). 

The exterior involucral bracts, when few and more oblong, would 
seem to connote a variety. Too many apparently transitional forms 
have been encountered in herbaria, however, to permit as yet 
definite conclusions to be reached. 

13. Coreopsis Stillmanii (A. Gray) Blake, Proc. Amer. Acad. 
49: 342. 1914. Leptosyne Stillmanii A. Gray in E. Durand, Journ. 
Acad. Nat. Sci. Phila. 3: 91. 1855; Torr. Bot. Mex. Bound 2: 92. 1859. 
Involucri bracteae exteriores 5-8, oblongae, 4-8 mm. longae; achae- 

niis 3.5-4 mm. longis. .-.' C. Stillmanii. 

Involucri bracteae exteriores 8-16, tenuissime lirieares superne usque 
ad apicem sensim angustatae, 7-12 mm. longae; achaeniis 

4-4.5 mm. longis var. /8 Jonesii. 

Herba annua, erecta, glabra, 1.5-4.5 dm. alta, infra ramosa ac 
foliosa, ramis (interdum numerosissimis) in pedunculos nudos 
erectos monocephalicos elongatis. Folia petiolata petiolis usque ad 
5 cm. longis saepe latis planisque, petiolo adjecto 4-10 cm. longa, 
pinnatim 3-7-partita foliolis membranaceis anguste lineari-spathu- 
latis apice obtusis supra medium plerumque 1-2.5 mm. latis interdum 
rursus lobatis. Capitula (solitaria, longe pedunculata) radiata, pansa 
ad anthesin 2-3 cm. lata et 7-9 mm. alta. Involucri bracteae exteri- 
ores 5-8, oblongae, glabrae, obtusae, 4-8 mm. longae; interiores 
late ovatae, apice pubescentes, interdum paulo breviores. Flores 
ligulati circ. 8, flavi, ligula oblongi vel obovati, apice grosse circ. 
3-dentati, fertiles, usque ad 1.5 cm. longi et 1 cm. lati. Paleae glab- 
rae, oblongae, plus minusve acutae, 4-6 mm. longae. Flores tubulosi 
anulo plus minusve glabri, stylorum ramis termino abrupte crasso- 
conicis et vix apiculatis. Achaenia cuneato-orbiculata vel anguste 
cuneato-obovata, plana, dorso glabrata ventre plus minusve papil- 
losa vel aegre setosa setis interdum clavellatis, duabus faciebus nigra, 
margine crasso-alata rugosaque, circ. 3.5-4 mm. longa et (alis 
idjectis) 1.4-2.3 mm. lata, apice exaristata sed poculo membranaceo 
coronata. 



316 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: Collected by Dr. Stillman in valley of the upper 
Sacramento River, northern California (Gray). 

Distribution: California. 

Specimens examined: H. N. Bolander 4520, hillsides, Auburn, 
April 10, 1865 (Calif.; Gray; N.Y.; U.S.); Thomas Bridges 260, 
California (N.Y.); Mrs. C. C. Bruce 1987, Iron Canyon, Butte 
County, May, 1897 (Pom., 2 sheets); J. W. Congdon, Mariposa 
Water Ditch, Mariposa County, March 28, 1901 and April 4, 1903 
(U.S.); Harvard University Botanical Garden, cult., 1873 and 1882 
(Gray); A. Kellogg & W. G. W. Harford 439, dry sandhills, Antioch, 
April 16, 1868-1869 (Gray) and April 8, 1869 (N.Y.); Dr. Stillman 
(type, Gray; cotype, N.Y.). 

Coreopsis Stillmanii var. /3 Jonesii Sherff, Bot. Gaz. 97: 605. 
1936. 

Herba 2 dm. alta, basi multum ramosa. Capitula ad anthesin 
3.5-4 cm. lata et circ. 1 cm. alta. Involucrum glabratum; bracteis 
exterioribus 8-16, angustissime linearibus, sensim usque ad apicem 
obtuse truncatum angustatis, 7-12 mm. longis, demum saepe 
patentibus subreflexisve, quam interioribus oblongo-ovatis paulo 
brevioribus. Achaenia 4-4.5 mm. longa; corpore ipso nigro, circ. 
1 (alis stramineis inclusis circ. 2) mm. lato, glaberrimo vel secundum 
costam medianam sparsim papillato-hispidulo et alibi interdum 
paucissimis setulis armato, his apice nunc acribus nunc capitatis. 

Type specimen: Collected by Marcus E. Jones, No. 3361 pro 
parte, Pasadena, California, May 2, 1882 (Pom.). 

Distribution: Known only from type locality in California. 
Specimens examined: Jones 3361 pro parte (type, Pom.). 

14. Coreopsis petrophila A. Gray, Proc. Amer. Acad. 22: 428. 
1887. 

Fruticosa, 6 dm. forsitan quidem 1 m. alta, ramosissima, 
glabrata, caulibus gracilibus obscure tetragonis, saepe rubescentibus. 
Folia tenuiter petiolata petiolis 1-2.5 cm. longis, petiolo adjecto 3-6 
cm. longa, pinnatipartita, segmentis 5-9 subulato-linearibus, acriter 
indurato-apiculatis, inaequalibus, majoribus saepius trifidis vel 
1-2-dentatis, ultimis 0.6-5 mm. latis. Capitula graciliter brevius- 
culeque pedicellata (pedicellis pubescentibus) et subcymosa, radiata, 
pansa ad anthesin 2-2.5 cm. lata et circ. 7-8 mm. alta. Involucri 
campanulati subcriniti bracteae exteriores lineari-oblongae, minutae, 
tantum circ. 1-2 (-3) mm. longae; interiores oblongae vel oblongo- 



REVISION OF THE GENUS COREOPSIS 317 

ovatae, 5-6 mm. longae. Flores ligulati 5, flavi, integri, ligula 
oblongi, 8-12 mm. longi, tubulo hispido. Paleae scariosae, angusto- 
oblongae vel oblanceolatae, dorso longissime villosissimae, apice 
trilobatae, quam achaeniorum corpora paulo longiores. Flores 
tubulosi circ. 6-10, stylorum ramis apice tenuiter hispido-appendi- 
culatis. Achaenia recta, subturgida, obovato-oblonga, nigra, dorso 
glabra, marginibus intusque longissime villosissima, corpore 3-4.5 
mm. longa et 0.7-0.9 mm. lata, apice biaristata aristis tenuibus 
antrorsum barbellatis 3-4.5 mm. longis. 

Type specimen: Collected by Edward Palmer, No. 530, hanging 
loosely about rocks at the entrance of canyons, Rio Blanco, State of 
Jalisco, Mexico, September, 1886 (U.S.). 

Distribution: State of Durango and southward into State of 
Jalisco, Mexico. 

Specimens examined: C. R. Barnes & W. J. G. Land 158, alt. 
1,680 meters, Sierra de Esteban, State of Jalisco, September 28, 1908 
(Field); Edward Palmer 489< Tejame"n, State of Durango, August 
21-27, 1906 (Gray; N.Y.); idem 530 (type, U.S.; cotypes, Gray; 
N.Y.); C. G. Pringle 2197, cool ledges of barrancas, near Guadala- 
jara, State of Jalisco, November 20, 1888 (Gray) ; idem 2303, rocky 
bluffs of Rio Grande de Santiago, near Guadalajara, September 30, 
1889 (Field; Gray; N.Y.) idem 11507, alt. 1,500 meters, Sierra de 
Esteban, near Guadalajara, September 30, 1903 (Berl.; Field; Gray). 

15. Coreopsis petrophiloides Robins. & Greenm. Proc. Amer. 
Acad. 29: 388. 1894. 

Fruticosa, forsitan 7-9 dm. alta, caulibus plus minusve glabratis, 
superne foliosis; ramulis angulatis, paulo pubescentibus. Folia 
subsessilia vel petiolata petiolis marginatis usque ad 2 cm. longis, 
omnino 4-9 cm. longa, rigida, simplicia, rhomboideo-lanceolata, 
acuta, dimidio superiore inciso-serrata, glandulo-punctata, supra 
paulo pubescentia, infra glabra ac pallidissima. Capitula in inflo- 
rescentias terminales corymbosas congregata, tenuiter pedicellata 
pedicellis bracteatis plerumque 2-6 cm. longis, radiata, pansa ad 
anthesin 1.5-2.5 cm. lata et 8-10 mm. alta. Involucri bracteae 
adpresse hispidae, exteriores circ. 6-8, lineari-oblongae, apice 
rotundatae ac cartilagineae, tergo 3-nervatae, circ. 2-3 mm. longae; 
interiores ovali-oblongae, 5-7 mm. longae. Flores ligulati 8, 
flavi, ligula oblongi, apice denticulati, 1 cm. longi. Paleae angus- 
tissime rhomboideo-lanceolatae, tergo valde erecto-villosae ventre 
glaberrimae, apicem acrem versus lateraliter acriterque 3-lobatae 



318 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

lobo terminal! multo longiore, 6-7 mm. longae. Flores tubulosi 
5-dentati, stylorum ramis abrupte breviterque angusto-appendi- 
culatis. Achaenia oblongo-linearia, plana, brunneo-nigra, nitida, 
longitudinaliter obscuro-sulcata, tergo glabrata, marginibus et 
ventris nervo mediano perspicue erecto-villosa pilis brunneo-albidis, 
corpora circ. 5-6.5 mm. longa et pilis exclusis 0.5-1 mm. lata, apice 
biaristata aristis tenuissimis stramineis antrorsum hispidis circ. 
4-5.5 mm. longis. 

Type specimen: Collected by Cyrus Guernsey Pringle, No. 5508, 
at altitude of 2,400 meters, Nevado de Colima, State of Jalisco, 
Mexico, May 22, 1893 (Gray). 

Distribution: Nevado de Colima of southernmost Jalisco, south- 
eastwardly through Michoacan into State of Guerrero, Mexico. 

Specimens examined: Brother G. Arsene, alt. 2,100 meters, Cam- 
panario, vicinity of Morelia, State of Michoacan, December, 1910 
(U.S.); idem 3185, alt. 2,100 meters, Cerros San Miguel, vicinity of 
Morelia, November 18, 1909 (Gray; Kew; Mo.; N.Y.; Stockh.; 
U.S., 2 sheets); idem 5295, eodem loco, December, 1910 (Gray; 
Mo.; U.S.); idem 6026, alt. 2,200 meters, Cerro Azul, vicinity of 
Morelia, 1910 (Gray; Mo.; N.Y.; U.S.);tem 6060, alt. 2,200 meters, 
Cerros San Miguel, vicinity of Morelia, November 15, 1911 (Gray; 
Kew; Mo.; U.S.); E. W. Nelson 2185, alt. 2,100-2,550 meters, north- 
east slope of Sierra Madre, near Chilpancingo, State of Guerrero, 
December 24, 1894 (U.S.); idem 2249, alt. 2,700-3,060 meters, top 
of Sierra Madre, eodem loco et tempore (U.S.); Parkinson, Mexico 
(Kew; Oxf.); Pringle 5508 (type, Gray). 

16. Coreopsis Irmscheriana Bruns, Mitteil. Inst. Allgem. Bot. 
Hamburg 8: 81. f. 11, nos. 6-9. 1929. 

Herba (ex auctore perennis) 2-3 dm. alta, e radice primaria 
parce fibrosa 5-15 cm. longa; caule erecto, satis dense folioso, 
rubescenti, tereti, pilis strictis brevibus aspero, laxe ramoso; ramis 
decumbentibus vel adscendentibus, teretibus vel quadrangularibus, 
vix sulcatis, rubescentibus, hirtis, 1-2 dm. longis. Folia caulina 
opposita, brevissime petiolata vel sessilia, anguste ovato-lanceolata, 
irregulariter crenato-serrata, usque ad 4 cm. longa et 1.5 cm. lata; 
ramulorum alternantia, saepius sessilia, utrinque pilis rigidis sparse 
adpresseque hirsuta, plerumque sub 2.3 cm. longa et 0.7 cm. lata. 
Capitula solitaria ad apices ramorum superne subnudorum dense 
hirsutorum, radiata, pansa ad anthesin 3-3.5 cm. lata et saltern 
1 cm. alta. Involucri bracteae 16 irregulariter biseriales, lineari- 



REVISION OF THE GENUS COREOPSIS 319 

oblongae, dense hirsutae, usque ad 8 mm. longae et 2 mm. latae. 
Flores ligulati 12, aurei, ligula oblongi, circ. 1.4-1.6 cm. longi et 
db 4 mm. lati. Paleae margine serratae, apice longe acutae, circ. 
6 mm. longae. Flores tubulosi lutei, 7 mm. longi; tubo inferne 
piloso; limbi lobis intus pilosis. Achaenia obcompressa, nigra, pilosa, 
biaristata aristis ciliato-pilosis. 

Type specimen : Collected by Ernesto Guenther and Otto Buchtien, 
No. 55, at altitude of 30 meters, in the Ravine ("Schlucht") of 
Chule, Mejia, Lomas, Peru, November 12, 1923 (Hamb., 2 sheets). 

Distribution: Known only from type locality in southern Peru. 

Specimens examined: Guenther & Buchtien 55 (2 type sheets, 
Hamb.; cotype, Mun., from its photograph in Field). 

17. Coreopsis rhyacophila Greenm. Proc. Amer. Acad. 35: 
313. 1900. 

Suffruticosa, erecta, glabra, 3-10 dm. alta, caulibus teretibus, 
griseo-brunneis vel brunneo-rubescentibus, foliosis. Folia opposita 
petiolata petiolis tenuibus usque ad circ. 6 cm. longis, petiolo adjecto 
usque ad 11 cm. longa, plerumque 2-3-pinnatisecta, utrinque glabra, 
glanduloso-punctata, segmentis linearibus vel anguste oblongo- 
lanceolatis, apice acutis vel acerrimis, membranaceis, saepius 2-5 
mm. latis. Capitula corymbosa, non numerosa, tenuiter pedicellata 
pedicellis glabris nudis vel 1-3-bracteatis saepius 5-7 cm. longis, 
radiata, pansa ad anthesin 2.5-3 (rarius -4.5) cm. lata et circ. 8-11 
mm. alta. Involucri bracteae exteriores circ. 8, patentes vel reflexae, 
lineares, apice subacutae, basi saepe sparsim hispidae, 4-6 mm. 
longae; interiores late oblongo-ellipticae, apice pubescentes, 7-9 mm. 
longae. Flores ligulati 7-10, flavi, ligula elliptico-oblongi, apice 
plerumque integri, 1.4-2.3 cm. longi. Paleae lineari-oblongae, cum 
fructu deciduae, ad hujus faciem exteriorem applicatae, apice acres 
et saepe 1-2-lobatae, dorso ac margine erecto-villosissimae, ventre 
glabrae, 5-6 mm. longae. Florum tubulosorum styli ramosi, ramis 
conico-appendiculatis. Achaenia obcompressa, cuneate lineari- 
oblanceolata, nigra, dorso glabra, margine ac ventris costa mediana 
erecto-villosissima, corpore circ. 4 mm. longa et circ. 0.8 mm. lata, 
apice tenuiter biaristata aristis albidis antrorsum hispidis circ. 3-4 
mm. longis. 

Type specimen: Collected by Cyrus Guernsey Pringle, No. 7866, 
at altitude of 2,500 meters, in lava fields near Cuernavaca, State of 
Morelos, Mexico, February 9, 1899 (Gray). 



320 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Distribution: States of Michoacan and Morelos, southern 
Mexico. 

Specimens examined : Brother G. Arsene, Cerro Azul, District of 
Morelia, State of Michoacan, October, 1909 (Field); Pringle 7866 
(type, Gray); idem 8332, alt. 2,400 meters, lava fields above Cuer- 
navaca, State of Morelos, May 5, 1900 (Berl.; Field; Gray; N.Y.; 
Pom.); idem 8431, alt. 2,550 meters, eodem loco, October 13, 1900 
(Field; Gray; N.Y.; Pom.); idem 9886, eodem loco, November 19, 
1902 (Berl.; Field; Gray; N.Y.). 

18. Coreopsis Pringlei Robins. Proc. Amer. Acad. 43: 41. 1907. 

Fruticosa, ramosa, ramis glabris teretibus cortice ochraceo-griseo 
obtectis, ramulis striatis viridibus plus minusve 6-angulatis foliosis. 
Folia opposita petiolata petiolis 1 cm. longis ac saepe marginatis, 
petiolo adjecto 2-4 cm. longa et 1-3 cm. lata, bipinnatifida, pallide 
viridia, glaberrima vel vix pilosiuscula, segmentis patentibus angus- 
tissime linearibus leviter acutatis integris vel cum lobis secundariis 
paucis instructis 4-18 mm. longis et 0.6-0.8 mm. latis. Capitula 
ramulos terminantia solitaria vel corymbose 3-5-aggregata, tenuiter 
pedunculata pedunculis 1-5 cm. longis nudis vel in mediana parte 
cum bractea unica lineari instructis, erecta vel nutantia, radiata, 
pansa ad anthesin 2-3 cm. lata et 7-9 mm. alta. Involucri campa- 
nulati bracteae exteriores circ. 8, lineari-oblongae, apice rotundatae 
basi pilosiusculae, 3-5 mm. longae et 1 mm. latae; interiores oblongo- 
ovatae, circ. 6-8 mm. longae. Flores ligulati circ. 8, juventate supra 
aurei subtus flavidi maturitate laete flavi, ligula oblongi, apice 
integri, circ. 1.2 cm. longi et 4-6 mm. lati. Paleae lineari-oblongae, 
cum fructu deciduae, ad hujus faciem exteriorem applicatae, apice 
eroso-ciliatae, circ. 5 mm. longae. Stylorum rami pro floribus 
tubulosis anguste conico-appendiculati. Achaenia vix matura 
obcompressa, cuneate lineari-oblanceolata, dorso obscure circ. 
4-costata et glabra vel sparsim erecto-villosa, facie ventrali et 
marginibus erecte villosissima, corpore nigro circ. 5 mm. longa, apice 
tenuiter biaristata aristis albidis antrorsum hispidis circ. 3-4 mm. 
longis. 

Type specimen: Collected by Cyrus Guernsey Pringle, No. 10050, 
at altitude of 2,040 meters, on dry ledges, San Juan del Rio, State 
of Quere*taro, Mexico, September 8, 1905 (Gray). 

Distribution: Known only from type locality in State of Quere*- 
taro, Mexico. 

Specimens examined: Pringle 10050 (type, Gray; cotypes, Berl.; 
Field; N.Y.; U.S.). 






REVISION OF THE GENUS COREOPSIS 321 

19. Coreopsis rudis (Benth.) Benth. & Hook, ex Hemsl. Biol. 
Centr. Amer. Bot. 2: 196. 1881. Epilepis rudis Benth. PL Hartweg. 
17. 1839. 

Fruticosa, erecta, hispida, foliosa, forsitan 7-9 dm. alta. Folia 
opposita petiolata petiolis anguste alato-marginatis circ. 1-2 cm. 
longis, petiolo adjecto circ. 4-7 cm. longa, vix rigidula, pinnatisecta, 
segmentis cuneatis lanceolatisve mine integris mine grosse dentatis 
pinnatifidisve, saepius 3-10 mm. latis. Capitula corymboso-pani- 
culata pedicellis tenuibus plerumque 0.5-1.5 cm. longis, radiata, 
pansa ad anthesin circ. 2.5 cm. lata et circ. 8-10 mm. alta. Involucri 
campanulati bracteae exteriores circ. 6, oblongae, obtusae, circ. 3-4 
mm. longae, interioribus late oblongo-obovatis dimidio breviores. 
Flores ligulati circ. 5, flavi, ligula ovali-oblongi, neutri, apice sub- 
integri vel plus minusve denticulati. Paleae cum fructu deciduae, 
ad hujus faciem exteriorem applicatae, dorso valde longeque erecto- 
villosae, ventre glaberrimae, apice trifidae, circ. 5-6 mm. longae. 
Flores tubulosi 5-dentati, stylorum ramis cono superatis. Achaenia 
obcompressa, anguste lineari-oblanceolata, dorso circ. 4-costato 
glabra, marginibus villosissima, ventre praeter lineam villosam 
glabra, corpore subnigro 4-5 mm. longa et circ. 0.7-0.8 mm. lata, 
apice tenuiter biaristata aristis stramineis antrorsum hispidis circ. 
2-3 mm. longis. 

Type specimen: Collected by Theodor Hartweg, No. 116, in pine 
forests, Bolanos, Canton of Colotlan, State of Jalisco, Mexico, 1836- 
1838 (Kew). 

Distribution: Known only from type locality in Canton of Colot- 
lan, State of Jalisco, Mexico. 

Specimens examined: Hartweg 116 (type, Kew; cotypes, Berl.; 
Gray). 

Bentham described the ray flowers as neuter, but Bentham and 
Hooker (Gen. PI. 2: 386, sub Epilepide. 1873) described them as 
fertile. In the fruiting heads before me they are completely sterile. 

20. Coreopsis glaucodes Blake & Sherff ex Sherff, Bot. Gaz. 
80: 369. 1925. 

Frutex glaberrimus, 1 m. altus, partibus novellis glaucescenti- 
bus, caule tereti subgriseo, ramis ramulisque striatis, parce angulatis, 
subbrunneis, glaucescentibus, internodiis 1-6 cm. longis. Folia 
opposita tenuiter petiolata petiolis basi in poculum circ. 1 mm. altum 
connatis, 4-10 mm. longis, petiolo adjecto circ. 1.5-3.5 cm. longa 
et 6-18 mm. lata, circumambitu saepius cuneata vel rhombica 



322 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

rarius deltoidea, ad medium vel ultra 3- (vel etiam 4-) lobata et 
in petiolum cuneate decurrentia, lobis cuneatis vel oblanceolatis 
vel lineari-lanceolatis vel lanceolatis, acutis, integris (interdum 
terminal! rarius etiam lateralibus 2-3-dentatis), membranaceis, pal- 
lido-viridibus, glaucescentibus, plerumque 6-16 mm. longis et 1.5- 
3.5 mm. latis. Capitula parva 1-4 ad apices ramorum, tenuiter 
pedunculata pedunculis 1.5-5.5 cm. longis, radiata, pansa ad anthesin 
1.2-2 cm. lata et 5-7 mm. alta. Involucri glabri glaucescentisque 
bracteae exteriores 8, anguste triangularibus, obtusae, 3-nerviae, 
2-2.5 mm. longae et 0.2-0.5 mm. latae; interiores oblongae, obtusae, 
apice obscure ciliolatae, multinerviae, flavo-marginatae, glauces- 
centes, 5-7 mm. longae et circ. 1.8 mm. latae. Flores ligulati 8, 
aurei, ligula oblongi, subintegri, circ. 8-nervii, 7.5 mm. longi et 
3.2 mm. lati. Paleae oblongae, subacutae vel obtusae, circ. 
5-nerviae, ad lineam medianam pilosae, apice eroso-spinulosae, 
4.5 mm. longae. Disci florum stigmata deltoidea, hispidula, api- 
culata. Achaenia oblanceolato-oblonga, obcompressa, griseo-nigra, 
dorso glabra, marginibus piloso-ciliata, apice ciliata, ventre ad 
costam medianam sparsim pilosa, corpore 3.5-4 mm. longa et 1.8 
mm. lata, biaristata aristis sursum ciliatis, 1.5-2 mm. longis. 

Type specimen: Collected by August Weberbauer, No. 4048, in 
the open formation, at altitude of 2,600 meters, below Hacienda La 
Tahona, near Hualgayoc, Department of Cajamarca, Peru, May 15, 
1904 (Berl., 2 sheets). 

Distribution: Known only from type locality. 

Specimens examined: Weberbauer 4048 (2 type sheets, Berl.). 

21. Coreopsis microlepis Blake & Sherff ex Sherff, Bot. Gaz. 
80: 370. 1925. 

Frutex + 4 dm. altus, caule tenui sub inflorescentia plerumque 
simplici raro ramis paucis erectis simplicibus ramoso, usque ad 1.8 
mm. crasso, hexagono, glabro vel in lineis posita sub nodis plus 
minusve hispidulo, internodiis 1.3-5 cm. longis, quam foliis plerum- 
que longioribus. Folia opposita tenuiter petiolata petiolis superne 
submarginatis inferne hirsuto-ciliatis 4-10 mm. longis basi in poculum 
connatis, lamina circumambitu deltoidea paene ad costam trilobata, 
8-21 mm. longa et 7-22 mm. lata, segmentis membranaceis, glabra, 
margine saepe revolutis, acriter apiculatis, lateralibus cuneatis vel 
elliptico-obovatis vel spathulatis, integris vel 2-3-dentatis, 5-11 
mm. longis et 2-6 mm. latis, terminali 3-4-lobato saepius usque circ. 
ad medium lobis integris vel 2-3-dentatis. Capitula mediocria 3-15 



REVISION OF THE GENUS COREOPSIS 323 

cymam vel paniculam nudam efficientia rarius solitaria, pedicellata 
pedicellis pilosis 1.5-5 cm. longis, radiata, pansa ad anthesin circ. 
1.5-2 cm. lata et 6-7 mm. alta. Involucri glabri vel basi ipso his- 
piduli bracteae exteriores 8-9, ovatae vel oblongo-ovatae, sub- 
obtusae, 1.5-2 mm. longae et 0.6-0.8 mm. latae; interiores elliptico- 
oblongae, obtusae, facie subnigrae margine angusta flavidae, 6-7 mm. 
longae et circ. 2.2 mm. latae. Flores ligulati verisimiliter 8, aurei, 
ligula oblongo-ovati, subintegri, circ. 7-nervii nervis principalibus 
dorsaliter sparsimque hirsutulis, db 6.7 mm. longis et 3 mm. latis. 
Paleae obovatae, obtusae vel acutae, circ. 6-nerviae, ad lineam 
medianam pilosae, circ. 4.5 mm. longae. Disci florum stigmata 
incrassata, hispidula, abrupte apiculata. Achaenia obovata vel 
oblonga, griseo-nigra, longe piloso-ciliata et duabus faciebus pilosa, 
corpore 2.8-3.5 mm. longa et circ. 1.2 mm. lata, biaristata aristis 
subpaleaceis, sursum piloso-ciliatis, 0.8-1.2 mm. longis. 

Type specimen: Collected by Alexander Mathews, No. 1418, 
Province of Chachapoyas, Department of Amazonas, Peru, 1835- 
1836 (Kew, 2 sheets). 

Distribution: Known only from type locality. 

Specimens examined: Mathews 1418 (2 type sheets, Kew); A. 
Raimondi 1774, road between Chacapoyas and Cheto, Peru, March, 
1869 (Berl.). 

22. Coreopsis Macbridei Sherff, Bot. Gaz. 89: 369. 1930. 

Frutex glabratus, ramosus, caulibus subteretibus. Folia laxe 
disposita, opposita, petiolata petiolis tenuibus circ. 1-1.5 cm. longis, 
primaria petiolo adjecto etiam 4-4.5 cm. longa, pinnatim 5-7-secta, 
segmentis anguste linearibus, glaberrimis, apicaliter acribus, 1-2.5 
cm. longis et 1-1.7 mm. latis. Capitula corymboideo-paniculata, 
numerosa, tenuissime pedunculata (vel pedicellata) pedunculis 
saepius 3-7.5 cm. longis glaberrimis vel apicem versus vix pubes- 
centibus, radiata, pansa ad anthesin tantum circ. 2 cm. lata et 
6-9 mm. alta. Involucri glabrati sicci subnigri bracteae exteriores 
circ. 8, lineari-oblongae, obtusae, tantum circ. 1.5-2 mm. longae; 
interiores oblongo-ovatae circ. 5-6 mm. longae. Flores ligulati circ. 
8, fusco-flavi, ligula lineari-elliptici, apicaliter integri vel rarius 
denticulati, tantum circ. 8-10 mm. longi et 1.5-3 mm. lati. Paleae 
lineari-oblongae, circ. 5-striatae, tergo villosissimae ventre glab- 
rae cum achaenio deciduae. Disci florum stigmata incrassata et 
minutissime caudato-appendiculata. Achaenia lineari-oblanceolata, 
valde obcompressa, dorso sub palea glabra, ventre ad costam medi- 



324 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

anam villosa, marginibus longe perspicueque villoso-ciliata, apice 
biaristata aristis sursum villosis circ. 2 mm. longis. 

Type specimen: Collected by J. Francis Macbride, No. 3504, neat 
compact half-shrub (or shrub) of southwestern rock outcrops, alt. 
about 2,100 meters, Huanuco, Peru, April 26, 1923 (Field). 

Distribution: Peru. 

Specimens examined: Macbride 3152, ragged shrub, river canyon 
slopes, alt. about 2,100 meters, Ambo, April 4, 1923 (Field; U.S.); 
idem 3504. (type, Field; cotype, U.S.). 

The specimens examined had been determined by Dr. S. F. Blake 
as a form of Coreopsis Townsendii Blake, with more leaf lobes than 
on the type. A careful comparison with the type of that species, 
however, shows many striking differences. C. Macbridei has a more 
scraggly habit; its leaves are fewer, their petioles narrower, and their 
divisions 5-7 rather than usually 3; the capitula are more numerous 
(more than 30 on one sheet), their peduncles and involucres almost 
completely glabrous (not conspicuously tomentose), their external 
bracts about 1.5-2 mm. (not 4-4.5 mm.) long, their diameter at 
flowering about 2 cm., not 2.5-3 cm., the ligules deep yellow (not 
light yellow) and only about 8-10 mm. long and 1.5-3 mm. wide, 
not 15-18 mm. long and 5-7 mm. wide, etc. 

23. Coreopsis Killipii Sherff, Bot. Gaz. 94: 594. 1933. 

Fruticosa, glabra, 6-12 dm. alta, ramosa ramis tenuibus angulatis 
internodiis quam foliis longioribus. Folia petiolata petiolis moderate 
ciliatis circ. 4-13 mm. longis basi in poculum 1.5 mm. altum 
connatis, petiolo adjecto circ. 1.5-2.5 cm. longa, pinnatim vel sub- 
bipinnatim divisa, segmentis ultimis saepius oblongo-linearibus 
crassiusculis eciliatis planis vel vix revolutis apice acutis subacutisve. 
Capitula pauca, in pedunculis ramos elongates superne nudatos 
terminantibus 1-1.2 cm. longis disposita, radiata, pansa ad 
anthesin 2.6-2.8 cm. lata et 0.8-1 cm. alta. Involucrum 
tantum ad basim ipsam hispidum, bracteis exterioribus circ. 8, 
oblongis, eciliatis, apice obtusis vel rotundatis, 3-5 mm. longis et 
1-1.7 mm. latis; interioribus ovato-oblongis, siccis plus minusve 
atris, apicem versus vix ciliatis, 7-10 mm. longis. Flores ligulati 
forsitan 5 vel 6, brunneo-flavi, ligula elliptico-oblongi, apice integri 
vel parce denticulati, circ. 1.2 cm. longi. Paleae spathulato-oblance- 
olatae, dorso (praecipue ad costam medianam) hispidae ventre 
glaberrimae, apicem acutum versus ciliatae, circ. 5-6 mm. longae ad 
corpus achaenii strictim applicatae. Flores tubulosi subbrunneo- 
flavi, stylorum ramis incrassatis apice subito breviterque conico- 



REVISION OF THE GENUS COREOPSIS 325 

appendiculatis. Achaenia valde obcompressa, cuneato-oblanceolata, 
nigra vel fere purpurascenti-nigra, dorso glaberrima, ventre margini- 
busque perspicue erecto-pilosa pilis elongatis sericeis, corpore 4-5 
mm. longa, apice biaristata aristis linearibus stramineis antrorsum 
hispidis 1.5-2.5 mm. longis. 

Type specimen: Collected by Ellsworth P. Killip and Albert C. 
Smith, No. 21823, straggling shrub 2-4 feet high, open hillside at 
altitude of 3,000-3,200 meters, Tarma, Department of Junin, Peru, 
April 20-22, 1929 (Field). 

Distribution: Known only from type locality in Peru. 

Specimens examined: Killip & Smith 21823 (type, Field). 

24. Coreopsis parviceps Blake & Sherff ex Sherff, Bot. Gaz. 80: 
368. 1925. 

Frutex glaberrimus, caule tereti, striatulato, griseo, ramis ramu- 
lisque tenuibus striato-angulatis subbrunneis, internodiis 1.5-4 cm. 
longis. Folia opposita petiolata petiolis tenuibus basi in pocula 
brevia et vix ciliolata connatis 3-8 mm. longis; laminis inferioribus 
circumambitu deltoideis, circ. 2.2 cm. longis et usque ad 2.8 cm. latis, 
irregulariter 2-4-partitis, crassiusculis, minutissime punctulatis, 
segmentis linearibus vel lineari-ellipticis 9-16 mm. longis et 1.8-3 
mm. latis perlucidule pinnato-venosis apice acutis; superioribus 
integris vel interdum trilobatis, lineari-ellipticis, 8-19 mm. longis 
et 1.5-3 mm. latis. Capitula parva, solitaria, tenuiter pedunculata 
pedunculis glabris db 2.7 cm. longis, radiata, pansa ad anthesin 
circ. 1.8 cm. lata et circ. 6 mm. alta. Involucri bracteae exteriores 
6-8, lineari-oblongae, glabrae, apice obtusae, circ. 3 mm. longae et 
0.8 mm. latae, interiores late lanceolato-oblongae, apice obtusae et 
eroso-denticulatae, circ. 6 mm. longae et 2.5 mm. latae. Flores 
ligulati circ. 8, aurei, ligula late oblongo-ovati, circ. 12-nervii, apice 
subintegri ac subemarginati, circ. 9 mm. longi et 4 mm. lati. Paleae 
oblongo-obovatae, dorso sparsim pubescentes, circ. 5-nerviae, apice 
truncatae vel emarginatae plus minusve spinuloso-ciliatae, circ. 3.5 
mm. longae. Disci florum stigmata deltoideo-triangularia, acuminata, 
hispidula. Achaenia valde immatura oblonga, piloso-ciliata, ventre 
ad costam medianam sparsim pilosa, apice ciliata ac biaristata aristis 
sursum piloso-ciliatis, 2 mm. longis. 

Type specimen: Collected by Const. De Jelski (distrib. Ign. 
Szyszylowicz), No. 765, Tambillo, Peru, August 19, 1878 (Berl., 2 
sheets). 

Distribution: Known only from type locality. 

Specimens examined: De Jelski 765 (2 type sheets, Berl.). 



326 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

25. Coreopsis oblanceolata Blake, Contr. U. S. Nat. Herb. 
22: 642. 1924. 

Frutescens, caule gracili, glabro, internodiis inferioribus mod- 
erate brevioribus superioribus multo longioribus quam foliis. Folia 
opposita, oblanceolata, Integra, coriacea, glabra, apice acutiuscula 
obtusave et obtuse calloso-mucronulata, de loco diametri maximi 
sensim in basim petiolo saepe similem angustata, parce revoluta, 
manifeste unicostata venis lateralibus obscuris, utrinque viridia, 
circ. 2.5-3.8 cm. longa et circ. 5-7.5 mm. lata. Capitula 
solitaria vel perpauca, pedunculata pedunculis subtenuibus usque 
ad 11 cm. longis, radiata, pansa ad anthesin 4 cm. lata et 
1.2 cm. alta. Involucri bracteae exteriores circ. 8, anguste ob- 
longae, basi parce angustatae et sparsissime pubescentes, apice 
rotundatae vel obtusae, marginibus breviter hispidae, 3-vittatae, 
circ. 5.5 mm. longae et circ. 1.8 mm. latae; interiores oblongo- 
obovatae, acutae acuminataeve, apice eroso-ciliatae, circ. 7.5 mm. 
longae. Flores ligulati 8, flavi, ligula elliptico-oblongi, apice 
bidentati, 2 cm. longi et 6 mm. lati. Paleae elliptico-ovatae, 
acuminatae, inferne ciliatae, apicem versus eroso-ciliatae, 5 mm. 
longae. Flores tubulosi flavi, tubi apice pilosi; styli ramis termino 
breviter triangulatis et acutis. Achaenia immatura valde obcom- 
pressa, oblongo-elliptica, ventre pubescentia, dense longo-ciliata, 
apice setosa, corpore circ. 4 mm. longa, biaristata aristis lanceolato- 
linearibus, subaequalibus, dense antrorsumque hispidis, circ. 2.2 
mm. longis. 

Type specimen: Collected by Charles Henry Tyler Townsend, No. 
A211, at altitude of 2,400-2,850 meters, on top of the western 
Cordillera opposite Huancabamba, Peru, September 26, 1911 (Field). 

Distribution: Known only from type locality in northernmost 
Peru. 

Specimens examined: Townsend A211 (type, Field). 

25a. Coreopsis Woytkowskii sp. nov. 

Fruticosa, usque ad 7 dm. alta; ramulis sulculatis albido-tomen- 
tosis. Folia opposita subtenuiter petiolata petiolis marginatis con- 
duplicatis hispido-ciliatis et faciebus plus minusve setosis usque ad 
circ. 1.5 cm. longis, petiolo adjecto usque ad 5.5 cm. longa et 0.8-1.5 
cm. lata, oblongo-oblanceolata, supra medium vel apicem versus 
pauciserrata (1-3 dentibus pro utroque latere), apice acuta rarius 
subtruncata sub medio usque ad petiolum sensim vel curvescente 
angustata, submembranacea, utrinque minutissime numerosissime- 



REVISION OF THE GENUS COREOPSIS 327 

que glandulo-papillata sed non vel secundum venulas hispidula, 
suaveolentia. Capitula solitaria ad ramulorum apices disposita, 
pedunculata pedunculo nitide albo-lanato circ. 2.5-4 cm. longo, 
radiata, pansa ad anthesin 4-4.5 cm. lata et 1.3-1.8 cm. alta. In- 
volucrum extus basi perspicuissime alibi moderate sparsimve albo- 
lanatum; bracteis exterioribus 6, oblongis, supra medium dilatatis, 
apice rotundis, tergo 3-nervatis et minutissime glandulo-papillatis, 
circ. 8 mm. longis et 2-2.6 mm. latis; interioribus oblongo-ovatis 
rfcdimidio longioribus. Flores ligulati 8, intense lutei, ligula obovati, 
apice obscure 2- vel 3-denticulati, 2 cm. longi, tubo ovarioque plus 
minusve albo-pilosis. Paleae oblongo-lineares, dorso longissime 
erecteque sericeo-pilosae, ventre glabrae, apice eroso-emarginatae, 
7 mm. longae. Achaenia matura non visa; valde immatura plana, 
cuneate linearia, dorso glabra, ventre marginibusque elongate 
erecteque sericeo-pilosa, apice biaristata aristis tenuissimis antror- 
sum hispidis corollis subaequalibus. 

Type specimen : Collected by Felix Woytkowski, No. 24, on rocky 
hills at altitude of 2,625 meters, vicinity of Celendin, Department of 
Cajamarca, Peru, June 5, 1936 (Field). 

Distribution: Known only from type locality in Peru. 
Specimens examined: Woytkowski 24- (type, Field). 

The collector's notes state that this species "grows in profusion 
in some places." It is very easily distinguishable in its foliage from 
all other South American species of Coreopsis. Fortunately, the 
type, though collected but very recently, was received in time to be 
treated in this work. To Mr. Paul C. Standley, Associate Curator 
of the Herbarium of Field Museum of Natural History, I am in- 
debted for the privilege of examining it. 

26. Coreopsis longula Blake, Contr. U. S. Nat. Herb. 22: 642. 
1924. 

Frutex glaber, verisimiliter circ. 1 m. altus, caule ramisque 
tenuibus et multiangulato-striatis, internodiis quam foliis saepius 
brevioribus. Folia opposita interdum cum fasciculis axillaribus, 
simplicia, flagellari-linearia, basi in poculum circ. 1 mm. altum 
connata, crassiuscula, subrevoluta, sicca minutissime numerosis- 
simeque albido-punctulata, eciliata, apice obtusiusculo cartilagineo- 
indurata, saepius 4-7.5 cm. longa et 0.6-1.1 mm. lata. Capitula 
non pauca, pedunculata pedunculis tenuibus 1-4 cm. longis, radiata, 
pansa ad anthesin 2.4 cm. lata et circ. 8 mm. alta. Involucri 
glabrati bracteae exteriores circ. 8, oblongo-lineares, obtusae, 



328 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

2.5-4 mm. longae et 0.4-0.6 mm. latae; interiores oblongo-lanceolatae, 
flavo-marginatae, apice obtuso subtruncatae et eroso-denticulatae, 
7-8.5 mm. longae et 2-2.5 mm. latae. Flores ligulati circ. 8, flavi, 
ligula elliptico-oblanceolati, apice integri vel obscure denticulati, 
circ. 1.2 cm. longi et 2.5-3.5 mm. lati. Paleae oblongo-lineares, 
obtusiusculae, 5-7-striatae, glabrae vel dorsaliter ad costam medi- 
anam hispidae, 7-8.5 mm. longae et 0.6-1 mm. latae. Disci florum 
stigmata incrassata, breviter obscureque crasso-apiculata. Achaenia 
lineari-oblonga, obcompressa, atra, corpore circ. 7 mm. longa et circ. 1 
mm. lata, dorso leviter ventre marginibusque graviter sursum pilosa, 
apice sursum pilosa et vestigio floris minutissimo coronata et bia- 
ristata aristis sursum hispidis 2.5-3 mm. longis. 

Type specimen: Collected by Alexander Mathews, in the Province 
of Chachapoyas, Peru, 1835-1836 (Gray). 

Distribution: Known only from type locality. 

Specimens examined: Matheivs, Province of Chachapoyas, Peru 
(type, Gray); idem, eodem loco, sine anno (Kew) et 1835 (Kew) et 
1836 (Kew, 2 sheets). 

27. Coreopsis venusta HBK. Nov. Gen. et Sp. 4: 180 (229). 
1820. 

Frutex ramosus, erectus vel procumbens, ramis teretibus glabris 
striatis, ramulis tetragonis, striatis; omnibus internodiis plerumque 
0.6-3 cm. longis. Folia opposita, indivisa, anguste spathulato- 
linearia, apice acutiuscula, basi angustata, integerrima, exsiccata 
subcoriacea, subuninervia, eciliata, supra glabra, infra ad costam 
medianam hispida, 1.5-4 cm. longa et 0.8-1.5 mm. lata. Capitula 
solitaria, tenuiter pedunculata pedunculis glabratis 5-15 cm. longis, 
radiata, pansa ad anthesin circ. 2.5 cm. lata et 6-8 mm. alta. Invo- 
lucri plus minusve hispidi bracteae exteriores circ. 8, oblongae, 
obtusae, circ. 3.5-4 mm. longae et 1-1.5 mm. latae; interiores ovato- 
ellipticae, obusae, multinerviae, margine angusto flavidae aliter atro- 
brunneae, 5-6 mm. longae. Flores ligulati 6-8, aurantiaci, ligula 
oblonga, apice obsolete tridentata vel subintegri, circ. 12 mm. longa 
et 5-6 mm. lata. Paleae oblanceolatae, tenuissime membranaceae, 
flavi, perspicue atronervatae circ. 6 nervis, supra laciniato-ciliatae, 
circ. 6 mm. longae. Disci florum stigmata incrassata, hispida, 
apicaliter brevi-caudata. Achaenia lineari-oblonga, obcompressa, 
brunneo-atra, tergo sub palea glabra, margine piloso-ciliata, ventre 
hispida saepe plus minusve tuberculata, corpore circ. 5 mm. longa 
et 1-1.2 mm. lata, apice biaristata aristis sursum piloso-ciliatis 
circ. 3 mm. longis. 



REVISION OF THE GENUS COREOPSIS 329 

Type specimen: Collected by Alexander Humboldt and Aime 
Bonpland, near Loja (Loxa), Province of Loja, Ecuador (Par.). 

Distribution: Ecuador and Peru. 

Specimens examined: A. S. Hitchcock 21564, alt. 2,200-3,100 
meters, Paramo between San Lucas and Ona, Province of Loja, Ecua- 
dor, September 7, 1923 (Gray; N.Y.; U.S.); Humboldt & Bonpland, 
near Loja, Ecuador (cotype, Berl.); iidem, Paramo de Saraguru, 
Ecuador (Berl.); A. Raimondi 6463, Callacate, Department of 
Cajamarca, Peru, May, 1879 (Berl.); idem 12060, Huambo, Province 
of Chota, Department of Cajamarca, Peru, June, 1868 (Berl.). 

28. Coreopsis nodosa Sherff, Bot. Gaz. 94: 595. 1933. 

Fruticosa, forsitan 6-8 dm. alta, ramosissima ramulis nunc sub- 
teretibus nunc valde angulatis glanduloso-puberulis internodiis 
numerosissimis plerumque 1-4 rarius usque ad 10 mm. longis et 
circ. 1-1.5 mm. crassis. Folia numerosissima opposita, petiolata 
petiolis plus minusve hispido-ciliatis basi in poculum minutum 
connatis 1-5 mm. longis (parte inferiore ad ramulos juncta manente 
post casum foliorum), petiolo adjecto tantum circ. 7-9.5 mm. longa, 
glabra, pinnatim 3-7-partita segmentis crassiusculis oblongis vel 
ovato-oblongis sparsim hispidulis apice subobtuso-mucronatis 
plerumque tantum 2-3.5 mm. longis et 0.4-0.7 mm. latis. Capitula 
tenuiter pedunculata pedunculis (supra summa folia) circ. 1 cm. 
longis, radiata, pansa ad anthesin 1.8 cm. lata et 7-8 mm. alta. 
Involucri basaliter tomentuloso-hispidi bracteae exteriores circ. 8, 
cuneato-oblongae, longitudinaliter 3-vittatae, glabratae, apice ob- 
tusissimae vel rotundatae, 2-3 mm. longae; interiores cuneato- 
oblongae apice obtusissimae 4-7 mm. longae. Flores ligulati 6-8, 
claro-flavi, ligula elliptico-oblongi, apice integri, 8 mm. longi. 
Paleae lineari-oblongae, dorso marginibusque erecto-hispidae ventre 
glabratae, apice acutae, 5-6 mm. longae, ad corpus achaenii strictim 
applicatae. Achaenia valde obcompressa, cuneate lineari-oblonga, 
atra, duabus faciebus praesertim ventrali ac marginibus valde 
erecto-setosa setis longis albidis sericeis, corpore circ. 4 mm. longa, 
apice biaristata aristis tenuibus stramineis antrorsum setosis 1-1.8 
mm. longis, inter aristas poculo minuto coronata. 

Type specimen: Collected by Francis W. Pennell, No. 13646, 
open, rocky slope, at altitude of 2,900-3,100 meters, Ollantaitambo, 
Department of Cuzco (Cusco), Peru, April 26, 1925 (Field). 

Distribution: Known only from type locality in Peru. 

Specimen examined: Pennell 13646 (type, Field). 



330 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

29. Coreopsis senaria Blake & Sherff ex Sherff, Bot. Gaz. 80: 
367. 1925. 

Frutex ramulis hexagonis, lineatim (2 lineis) hispidulis, infra 
foliorum basibus persistentibus vestitis, internodiis usque ad 10 mm. 
longis. Folia opposita cum fasciculis axillaribus, brevissime petiolata 
petiolis tantum circ. 1-1.5 mm. longis glabris omnino connatis; 
laminis 3-partitis, segmentis anguste lineari-oblanceolatis, apice 
acutis callosisque, margine integris, carnosis, atro-viridibus, glabris, 
1-nerviis, circ. 6-8 mm. longis et 0.6-0.9 mm. latis. Capitula solitaria 
in pedunculis brevissimis (usque ad 1 cm. longis) hirtellis insidentia, 
radiata, pansa ad anthesin circ. 2 cm. lata et 7 mm. alta. Involucri 
bracteae exteriores manifeste biseriales, omnino circ. 18, subaequales, 
herbaceae, lanceolatae vel lineari-lanceolatae, obtusae vel subacutae, 
saepe apiculatae, 3-nerviae, infra medium ciliatae, dorso glabrae, 
4-5 mm. longae et 1 mm. latae; interiores 8-9, membranaceae, 
oblongae, obtusae, margine angusto flavidae aliter brunneae, cilio- 
latae sed faciebus glabrae, circ. 2.5 mm. latae. Flores ligulati circ. 
8, aurei, ligula oblongo-ovati, circ. 11-nervati, circ. 6 mm. longi et 
3 mm. lati. Paleae lanceolatae, acuminatae, ciliatae, ad lineam 
medianam pilosae, 5-nerviae, circ. 4 mm. longae. Achaenia valde 
immatura obovato-oblonga, margine apiceque piloso-ciliata, facie 
exteriore glabra interiore aegre pilosa, biaristata aristis sursum 
piloso-ciliatis, circ. 2.2 mm. longis. 

Type specimen: Collected by A. Stubel, No. 35 pro parte, on 
punas, on an excursion from Pacasmayo to Moyobamba, Peru, 
April to June, 1875 (Berl.). 

Distribution: Peru. 

Specimens examined: A. Raimondi 436, heights between Balsas 
and Celendin, departments of Amazonas and Cajamarca, Peru, April, 
1869 (Berl.); Stubel 35 pro parte (type, Berl.); Felix Woytkowski 23, 
growing up to 6 dm. tall, common, on rocky hills, alt. 2625 m., vicinity 
of Celendin, Department of Cajamarca, Peru, June 5, 1936 (Field). 

The leafy branchlets have a juniperous aspect. The numerous 
outer bracts of the involucre also afford a ready means of determi- 
nation. 

30. Coreopsis foliosa A. Gray, Proc. Amer. Acad. 5: 125. 1861. 

Frutex ramosus, caule ramisque subquadrilateralibus striatis 
hispidis fere occultis. Folia numerosissima opposita glabra petiolata 
petiolis planis circ. 3-9 mm. longis, petiolo adjecto 1.2-2 cm. longa, 
lamina nunc digitato-ternata nunc pinnatim 4-7-partita, segmentis 



REVISION OF THE GENUS COREOPSIS 331 

oblanceolatis, crassiusculis, apice obtusiusculis, basim versus angus- 
tatis, faciebus minutissime densissimeque albido-punctulatis, mar- 
gine saepe revolutis, 4-8 mm. longis et 1-1.6 mm. latis. Capitula 
terminalia, breviter pedunculata pedunculis hispidis tenuibus 0.5-2 
cm. longis, radiata, pansa ad anthesin circ. 2.8-3.4 cm. lata et 1.2 
cm. alta. Involucri hispidi bracteae exteriores circ. 8, oblongae, 
apice rotundatae, circ. 5 mm. longae et 1.5 mm. latae; interiores 
lanceolatae apicaliter obtusae et interdum suberosae circ. 8 mm. 
longae. Flores ligulati circ. 8, ligula anguste obovati, apice plus 
minusve denticulati, 1.5-1.9 cm. longi. Paleae late lineari-oblongae, 
apice obtuso minute erosae, circ. 6 mm. longae et 1 mm. latae. 
Disci florum stigmata rotundo-incrassata, breviter apiculata. 
Achaenia late oblanceolata, nigra, piloso-ciliata, una facie perspicue 
altera leviter pilosa, corpore circ. 5 mm. longa et 1.8 mm. lata, apice 
biaristata aristis sursum hispidis circ. 3 mm. longis. 

Type specimen: Collected by Alexander Mathews, No. 1376, 
Taulia, Peru, in 1835 (Kew). 

Distribution: Known only from type locality. 

Specimens examined: Mathews 1376 (type and cotype, Kew). 

31. Coreopsis polyactis Blake & Sherff ex Sherff, Bot. Gaz. 
80:372. 1925. 

Frutex densissime foliosus, 1 m. altus, ramis subteretibus ac 
sordido-hirsutulis, internodiis 4-14 mm. longis. Folia opposita 
petiolata petiolis plus minusve ciliatis ad basim in poculum hirsutu- 
lum 1 mm. altum connatis, petiolo adjecto 1-1.6 cm. longa, in speci- 
minibus siccis plerumque subflavescentia, tripartita, segmentis 
integris vel terminali tripartite, linearibus vel anguste lineari- 
oblanceolatis, 3-6 mm. longis et 0.5-1 mm. latis, acutis, crassis, 
sparsim hirsutulis praecipue ad margines; axillis plerumque fasci- 
culis obsitis. Capitula saepius solitaria, brevissime pedunculata 
pedunculis crassiusculis dense flavescenteque subtomentoso-pilosis 
1-1.8 cm. longis demum fistulosis, radiata, pansa ad anthesin 3-3.5 
cm. lata et 8-10 mm. alta. Involucri bracteae dense flavescenteque 
subtomentoso-pilosulae exteriores circ. 11, manifeste biseriales, 
foliaceae, ovatae, obtusae, multinerviae, subaequales, 5-7 mm. 
longae et 3.3-4.8 mm. latae; interiores circ. 13, crasso-membranaceae, 
elliptico-oblongae, supra usque ad apicem subobtusum vel subacutum 
angustatae, anguste pallido-marginatae, multinerviae, quam ex- 
teriores dimidio longiores. Flores ligulati circ. 12, aurei, ligula 
obovati, subintegri, circ. 10-nervii, circ. 1.4-1.6 cm. longi et 6.5 mm. 



332 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

lati. Paleae lineari-oblongae, acutae, circ. 7-nerviae, supra sparsim 
ciliolatae, ad lineam medianam pilosae, circ. 6 mm. longae. Disci 
florum stigmata deltoidea, hispidula, apiculata. Achaenia immatura 
lineari-oblonga, obcompressa, piloso-ciliata, dorso glabra, ventre 
pilosa, apice eciliata sed biaristata aristis sursum ciliatis, 3-4 mm. 
longis. 

Type specimen: Collected by August Weberbauer, No. 6995, on 
grass steppes with scattered shrubs, at altitude of 3,500-3,600 meters, 
between Hacienda Llague*n and Succhabamba, Province of Otuzco, 
Department of Libertad, Peru, June 28, 1914 (Berl., 2 sheets). 

Distribution: Known only from type locality. 

Specimens examined: Weberbauer 6995 (2 type sheets, Berl.). 

Distinct from all other South American species of Coreopsis in 
its broadly oval-ovate, biseriate outer involucral bracts. 

32. Coreopsis notha Blake & Sherff ex Sherff, Bot. Gaz. 80: 
373. 1925. 

Folia plerumque 3-4 cm. longa C. notha sensu stricto. 

Folia circ. 1.5 cm. longa var. /3 parvulifolia. 

Frutex 1 m. altus, caule striatulo subtereti glabrescente, 
ramis striato-angulatis sparsim hispidulis, internodiis plerumque 
1-4.5 cm. longis. Folia opposita, tenuiter petiolata petiolis sparsim 
ciliatis 1-1.5 cm. longis ad basim in poculum plus minusve hispidulum 
1.5 mm. altum connatis, petiolo adjecto plerumque 3-4 cm. longa, 
ternatisecta lamina circumambitu deltoidea, foliolis lateralibus 
breviter stipitatis vel subsessilibus 3-5-sectis terminali longiusculo 
stipitato pinnatim 5-secto, segmentis ultimis lanceolatis vel oblongis, 
acutis, membranaceis, obscure pubescentibus et ciliatis, plerumque 
3-8 mm. longis et 1-2.5 mm. latis. Capitula longe tenuiterque 
pedunculata pedunculis monocephalicis sordide crispo-pilosulis 
praecipue apicem versus 7-21 cm. longis, radiata, pansa ad anthesin 
3.4-3.8 cm. lata et circ. 7-8 mm. alta. Involucri bracteae exteriores 
6, oblongae, apice obtusae vel rotundatae, basi plus minusve ciliatae, 
dorso non nisi ad basim pilosulae ventre sordido-pilosulae, 2-3 mm. 
longae et 1-1.6 mm. latae; interiores 8, oblongo-ellipticae, obtusae, 
anguste flavescenti-marginatae, apice minute ciliolatae aliter glabrae, 
9-12 mm. longae et 3-4.6 mm. latae. Flores ligulati, aurantiaci, 
ligula oblongo-obovati, apice subintegri, 13-15-striati, 1.7-2.3 cm. 
longi et circ. 9 mm. lati. Paleae oblongae, apice truncate hispidulo- 
ciliatae, ad dorsi lineam medianam pilosae, circ. 12-nerviae. Disci 
florum stigmata breviter deltoidea, hispidula, apiculata. Achaenia 



REVISION OF THE GENUS COREOPSIS 333 

lineari-oblanceolata, obcompressa, brunneo-nigra, marginibus piloso- 
ciliata, apice ciliolata, ad ventris costam medianam pilosa, corpore 
circ. 6 mm. longa et 1.7 mm. lata, bi- (raro tri-) aristata aristis 
sursum piloso-ciliatis, 2.7-3.4 mm. longis. 

Type specimen: Collected by August Weberbauer, No. 3812, at 
altitude of 2,400-2,700 meters, above San Pablo, Province of Caja- 
marca, Department of Cajamarca, Peru, April 26, 1904 (Berl., 2 
sheets). 

Distribution: Known only from Department of Cajamarca, Peru. 

Specimens examined: A. Raimondi 7732 and 7914, alt. 1,920 
meters, Cerro de Catache, Cascas, Province of Contumaza, Depart- 
ment of Cajamarca, Peru, May 21, 1875 (Berl.); Weberbauer 3812 
(2 type sheets, Berl.). 

Leaves similar to those of the Mexican C. rhyacophila Greenm., 
but smaller and somewhat less dissected. Vernacular name is given 
as "pul," and the heads are said to be used for dyeing. 

Coreopsis notha var. ft parvulifolia Sherff, Amer. Journ. Bot. 
22: 707. 1935. 

A specie foliis tantum circ. 1.5 cm. longis differt. 

Type specimen: Collected by A. Raimondi, No. 6695, Con- 
tumaza, Province of Contumaza, Department of Cajamarca, Peru, 
1875 (Berl.). 

Distribution: Known only from type locality in northern Peru. 

Specimen examined: Raimondi 6695 (type, Berl.). 

Sharply separable from C. notha by the smaller leaves and perhaps 
smaller stature of the plant. The more or less filiform leaf petioles 
offer a ready distinction from the otherwise deceivingly similar C. 
fasciculata var. laevigata of southern Peru. 

33. Coreopsis capillacea HBK. Nov. Gen. et Sp. 4: 180 
(230). 1820. 

Fruticosa, adscendens vel erecta, caule glabro demum 8 mm. 
crassa, forsitan 6-8 dm. alta, ramosa ramis lignosis glabris tetragonis- 
que, ramulis glabratis vel ad terminos hispidulis subdense foliaceis 
foliis quam internodiis multo longioribus. Folia opposita, petiolata 
petiolis usque ad 2 cm. longis, petiolo adjecto, 3-6 cm. longa, pin- 
natim 3-5-partita segmentis filiformibus 0.5-1 mm. latis, glabratis 
vel rarius minute hispidulis subplanis integerrimis apice acerrimis 
lateralibus quam terminali paulo brevioribus. Capitula solitaria 
vel perpauca, tenuiter pedunculata pedunculis pubescentibus circ. 



334 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

5-10 cm. longis, radiata, pansa ad anthesin circ. 2.5-3.8 cm. lata 
et 8-11 mm. alta. Involucri bracteae exteriores circ. 8, lanceolato- 
lineares vel oblongo-lineares, facie dorsali longitudinaliter valde 
3-nervatae, apice subobtusae, marginibus et facie ventrali albido- 
hispidae; interiores late oblongo-lanceolatae, longiores, glabratae, 
acutiusculae. Flores ligulati circ. 8, aurantiaci, ligula oblongo- 
elliptici vel oblongo-oblanceolati, apice denticulati, 1.5 cm. longi. 
Paleae lanceolatae vel sursum abrupte ad apicem et deorsum sensim 
ad basim angustatae, apice saepe eroso-dentatae, dorso hispidae 
ventre glabrae, 5 mm. longae. Florum tubulosorum styli rami 
apice breviter acriterque conici. Achaenia atra, oblonge lineari- 
lanceolata, valde obcompressa, longitudinaliter multistriata, margini- 
bus et ventre (saepe etiam ad tergi costam medianam) dense erecto- 
hispida pilis albidis elongatis sericeis et basi papillatis, corpore circ. 
4.5-5.5 mm. longa et circ. 1.2-1.5 mm. lata, apice plus minusve 
setoso biaristata aristis linearibus antrorsum hispidis circ. 2 mm. 
longis. 

Type specimen: Collected by Alexander Humboldt and Aime 
Bonpland, locality uncertain but suspected by Kunth as being in 
the Peruvian Andes. Type should be among the Humboldt and 
Bonpland specimens at Paris. 

Distribution: Southernmost Ecuador and perhaps adjacent Peru. 

Specimens examined: Ed. Andre 1^387, cold region, Catamayo, 
Province of Loja, Ecuador, October 31, 1876 (Kew); William Jame- 
son, hills near Loja, Ecuador, commun. anno 1865 (Kew) ; J. N. Rose, 
A. Pachano, & George Rose 23146, vicinity of Zaragura, Ecuador, 
September 28, 1918 (Gray; N.Y.; U.S.);Berthold Seemann 675, Loxa 
(Loja), Ecuador, August, 1847 (Kew). 

34. Coreopsis triloba Blake, Contr. U. S. Nat. Herb. 22: 
643. 1924. 

Fruticosa, erecta, caule ramisque gracilibus dense foliatis terminos 
versus primo piloso-tomentosis mox glabratis, internodiis plerumque 
2-8 mm. longis interdum longioribus (quidem 2 cm.). Folia opposita, 
petiolata petiolis 1-2 cm. longis basi in poculum 0.5-1 mm. altum 
connatis, petiolo adjecto 2-4 cm. longa, glabra, raro integra lineari- 
filiformiaque plerumque tripartita segmentis coriaceis lineari-fili- 
formibus apice subulatis integris vel mediano interdum 3-partito, 
ultimis 0.3-0.8 mm. latis. Capitula pauca (plerumque 1-3 ad 
unicum ramulum), tenuiter pedunculata pedunculis primo piloso- 
tomentosis demum glabratis, 1-6 cm. longis, radiata, pansa ad 



REVISION OF THE GENUS COREOPSIS 335 

anthesin circ. 1.5 cm. lata et 6-8 mm. alta. Involucri bracteae circ. 8, 
lanceolatae vel lanceolato-ovatae, obtusiusculae, crasso-herbaceae, 
adpressae vel parce laxae, longitudinaliter 3-vittatae, glabrae, circ. 
2.5-3 mm. longae et 0.8-1.2 mm. latae; interiores oblongo-ellipticae, 
obtusae, glabrae, circ. 5-6 mm. longae. Flores ligulati forsitan 8, 
flavi, ligula elliptico-oblongi, apice denticulati, 6 mm. longi. 
Paleae immaturae acutiusculae, apice spinuloso-ciliatae. Achaenia 
(valde immatura) obcompressa, ciliolata, biaristata aristis paleolis 
similibus antrorsum hispido-ciliatis 0.8 mm. longis. 

Type specimen: Collected by William Jameson in Ecuador (U.S., 
2 sheets). 

Distribution: Ecuador and Peru. 

Specimens examined: Const. De Jelski 705, Cutervo, Peru, May, 
1879 (Berl., 2 sheets); Jameson, Ecuador (2 type sheets, U.S.); 
E. P. Killip & A. C. Smith 22171, open rocky hillside, alt. 3150 
meters, Mantaro Canyon, south of Huancayo, Department of Junin, 
April 29, 1929 (Field; forma foliis plus minusve atypicis) ; A. Raimondi 
3022, Cutervo, Province of Cutervo, Department of Cajamarca, Peru, 
May, 1879 (Berl.). 

35. Coreopsis spectabilis A. Gray, Proc. Amer. Acad. 5: 125. 1861. 

Suffrutex saltern 3-4.5 dm. altus, caulibus glaberrimis, teretibus, 
striatis, internodiis 2-6 cm. longis et circ. 1.5-2 mm. crassis. Folia 
opposita, tenuiter petiolata petiolis 0.5-2 cm. longis basaliter in 
poculum hyalinum circ. 1-1.3 mm. altum connatis, petiolo adjecto 
3-6 cm. longa, 2-3-pinnatisecta, segmentis ultimis planis, glabris, 
acriter indurato-apiculatis, plerumque 0.5-0.8 mm. latis. Capitula 
perspicue pedunculata pedunculis glaberrimis (inferne 1-bracteatis 
vel saepius foliatis 2-3 jugis foliorum) 2.2-2.7 dm. longis, radiata, 
pansa ad anthesin 4 cm. lata et 6-8 mm. alta. Involucri bracteae 
glaberrimae, exteriores 6-8, late lineari-oblongae, apice obtusae, 
4-5.5 mm. longae et 0.9-1.5 mm. latae; interiores anguste ovatae, 
1-1.2 cm. longae. Flores ligulati (forsan 8), flavi, ligula obovati, 
apice obscure denticulati, circ. 1.5-1.8 cm. longi et 1 cm. lati. 
Paleae oblanceolatae, apicaliter obtusae, dorsaliter (ad margines 
glabrae aliter) sursum sericeo-pilosae, ventraliter glabrae, 6.5-8 mm. 
longae, unaquaque cum achaenio decidente. Achaenia valde obcom- 
pressa, nigra, oblanceolata, dorsaliter glaberrima, ventraliter (secun- 
dum costam medianam) ac marginaliter pilis sericeis albidisque 
perspicue surso-pilosa, ad latera subtilia, corpore 5.5-6.5 mm. longa 
et 2-2.3 mm. lata, apice biaristata aristis linearibus, stramineis, 
sursum sericeo-hispidis, parce patentibus, 2.5-3 mm. longis. 



336 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: Collected by McLean, in the Andes Mountains 
of Peru (Kew). 

Distribution: Peru. 

Specimen examined: McLean, Andes Mountains, Peru (type, 
Kew). 

36. Coreopsis suaveolens Sherff, Bot. Gaz. 89: 369. 1930. 
Frutex ramosus, verisimiliter 5-8 dm. altus, ramis subteretibus 

plerumque glabratis sed hac iliac glutinosis odore Covilleae divari- 
catae (Cav.) Vail suaveolentibus. Folia opposita fasciculata, pri- 
maria 1.2-1.6 cm. longa, pinnatim 3-5-secta, plus minusve glutinosa, 
segmentis petiolis similibus vel rursus sectis, ultimis linearibus, 
glanduloso-hispidis, carnosis, apicaliter mucronatis, tantum 0.3-0.6 
mm. latis. Capitula ramos superne nudos (pedunculos tenues) 
terminantia, solitaria, radiata, pansa ad anthesin 2.3 cm. lata et 
7-9 mm. alta. Involucri hispidi bracteae exteriores circ. 8, oblongae 
vel late lanceolatae, obtusae, 3-5 (-7) mm. longae, quam interiores 
oblongo-lanceolatae circ. 8 mm. longae saepius dimidio breviores. 
Flores ligulati circ. 6, lutei, ligula oblongo-elliptici, apice integri vel 
(etiam profunde) bifidi, 7-11 mm. longi. Disci florum stigmata 
brevia, abrupte incrassata truncataque. Achaenia ignota. Habitu 
C. fasciculatae Wedd. valde similis. 

Type specimen: Collected by Erich Werdermann, No. 1114, at 
altitude of about 3,800 meters, Cordillera de Lallinca, Province of 
Tarapaca, Department of Tarapaca, Chile, March, 1926 (Gray). 

Distribution: Known only from type locality in Chile. 

Specimens examined: Werdermann 1114 (type, Gray; cotypes, 
Field; Stockh.; Calif.). The label on the Field Museum sheet 
says "Cord. Co. Columfusca, Apacheta, Prov. Tarapaca. . . ." 

37. Coreopsis Pickeringii A. Gray, Proc. Amer. Acad. 5: 124. 
1861. Coreopsis Boliviana Blake, Contr. U. S. Nat. Herb. 22: 644. 
1924. 

Frutex fere glaberrimus vel praecipue caule ramisque plus minusve 
hispidus, 6-10 dm. altus, internodiis plerumque 1-4 cm. longis. 
Folia opposita petiolata petiolis tenuibus usque ad 2 cm. longis, basi 
in poculum membranaceum saepe 4 mm. altum connatis, petiolo 
adjecto 2-3.5 cm. longa, ternatisecta, foliolis 3-5-sectis, segmentis 
glabris, crassiusculis, linearibus, eciliatis, acriter apiculatis, usque 
ad 1 cm. longis et 0.6-1.5 mm. latis, quam rhachi tenui vix latioribus. 
Capitula longe tenuiterque pedunculata pedunculis usque ad 1.5 



REVISION OF THE GENUS COREOPSIS 337 

(rarius 2.3) dm. longis, radiata, pansa ad anthesin 3.5-4 (-5) cm. 
lata et 0.9-1.1 cm. alta. Involucri hispidi bracteae exteriores circ. 8, 
late lineari-oblongae, apice obtusae, dorso glabrae hispidaeve, 
marginibus longe ciliatae, circ. 5 mm. longae et 1-2 mm. latae; 
interferes oblongo-lanceolatae, flavo-marginatae, multistriatae, circ. 
1 cm. longae et 3-3.5 mm. latae. Flores ligulati 8 vel 9, flavi, ligula 
anguste obovati, apice rotundati et integri vel obscure denticulati, 
1.5-2.4 cm. longae et 4-9 mm. lati. Paleae oblongae, apice obtusis- 
simae vel truncatae et saepe eroso-dentatae, exteriores dorso villosae, 
circ. 7-8 mm. longae. Disci florum stigmata ovoidea, breviter 
apiculata. Achaenia oblongo-oblanceolata, plana, atra, dorso sub 
palea plus minusve glabra ventre et praesertim marginibus sursum 
villosissima, corpore 5.5-6.5 mm. longa et circ. 1.1-1.3 mm. lata, 
apice biaristata aristis sursum hispidis circ. 2 mm. longis. 

Type specimen : Collected by the United States Exploring Expe- 
dition under Captain Wilkes, high in the Andes above Obrajillo, 
Department of Lima, Peru (U.S.). 

Distribution: Peru and Bolivia. 

Specimens examined: Otto Buchtien 97, alt. 3,750 meters, dry 
slopes, La Paz, Bolivia, June 18, 1906 (Field) ; J. Francis Macbride 
2932, alt. about 2,400 meters, on southeastern rock slide, Matucana, 
Peru, March 14-18, 1923 (Field; U.S.); idem & Feather stone 210, alt. 
about 2,400 meters, rock outcrop on northern slope, Matucana, Peru, 
April 12-May 3, 1922 (Field) ; Francis W. Pennell 14342, forming 
large clumps on open, rocky slope, alt. 2,700-2,900 meters, Canta, 
Department of Lima, Peru, June 11-19, 1925 (Field) ; H. H. Rusby 
1685, alt. 3,000 meters, near La Paz, Bolivia, October, 1885 (type 
collection of C. boliviana Blake; Field; Gray; N.Y.); U. S. Explor. 
Exped. under Capt. Wilkes, above Obrajillo, Peru (type U.S.). 

38. Coreopsis imbricata Sherff, Bot. Gaz. 89: 370. 1930. 

Frutex 6-12 dm. altus, caule ramisque glabris, striatis, ramulorum 
internodiis numerosis saepius tantum 2-10 mm. longis. Folia 
opposita, petiolata petiolis tenuibus basi vix connatis circ. 1 cm. 
longis, petiolo adjecto circ. 2-2.5 cm. longa, ternatim bipinnatisecta, 
segmentis linearibus, crassiusculis, glabris, eciliatis, acriter apiculatis, 
2-8 mm. longis et circ. 0.3-0.6 mm. latis. Capitula ramulos termi- 
nantia (vix manifeste pedunculata), radiata, pansa ad anthesin 
3.5-4.5 cm. lata et 1-1.2 cm. alta. Involucri villosi bracteae exteri- 
ores circ. 16, biseriales, late lineari-oblongae, apice obtusae, 6-7 
mm. longae et 1.1-1.5 mm. latae ; interiores ovatae, flavo-marginatae, 



338 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

obtusae, 1-1.1 cm. longae. Flores ligulati circ. 8, flavi, ligula oblongo- 
elliptici, apice integri vel subintegri, 1.8-2.4 cm. longi et 6-7 mm. 
lati. Paleae lineares, circ. 5-striatae, apice obtusae et denticulatae, 
8 mm. longae et 0.5 mm. latae. Disci florum stigmata caudato- 
apiculata. Achaenia lineari-oblanceolata, plana, atra, dorso glabra, 
ventre ac marginibus ac apice sursum villosa, corpore 5 mm. 
longa et 1.4 mm. lata, biaristata aristis sursum hispidis 2.7 
mm. longis. 

Type specimen: Collected by R. Pearce, altitude 3,600 meters, 
Cordillera Huanta (midway between Lima and Cuzco), Peru, 
February, 1867 (Kew). 

Distribution: Known only from type locality. 

Specimens examined: Pearce, Cordillera Huanta (type, Kew). 

Close to Coreopsis Pickeringii Gray, from which it differs in its 
numerous, much shorter internodes, its lack of elongate peduncles, 
the caudate-tipped stigmas of its disc florets, its double outer in- 
volucre the bracts of which are larger, etc. 

39. Coreopsis Townsendii Blake, Contr. U. S. Nat. Herb. 
22: 643. 1924. 

Fruticosa, forsitan 4-7 dm. alta, caule gracili, tetragono, sub- 
viridi, glabro, ramis erectis. Folia opposita, petiolata petiolis basi 
connatis 1-2.5 cm. longis, petiolo adjecto 2-5 cm. longa, pinnatim 
3-5-partita segmentis integris linearibus acutis coriaceis unicostatis 
glabris vel inferne ad costam sparsim obscureque pubescentibus 
7-23 mm. longis et 1-1.8 mm. latis. Capitula solitaria vel perpauca, 
tenuiter pedunculata pedunculis laxe (ad capitulum dense) pilosis 
circ. 1-5.5 cm. longis, radiata, pansa ad anthesin 2.5-3.4 cm. lata 
et 7 mm. alta. Involucri bracteae exteriores circ. 6, lineari- 
oblongae, obtusae vel rotundatae, 3-vittatae, inferne ciliatae aliter 
glabrae, circ. 4-4.5 mm. longae et circ. 1-1.2 mm. latae; interiores 
elliptico-oblongae, apice obtuso eroso-ciliatae, 8 mm. longae et 2.5 
mm. latae. Flores ligulati 8, flavi, ligula anguste oblongo-obovati, 
14 mm. longi et 7 mm. lati. Paleae oblongae, apice eroso-ciliatae, 
dorso infra mediane erecto-pilosae, circ. 3.3-4 mm. longae. Flores 
tubulosi flavi, tubi apice glandulo-puberuli, gutture campanulato- 
infundibuliformi superne sparsim pilis multiloculatis piloso; stylorum 
ramis termino deltoideis et obtusiusculis. Achaenia submatura 
oblonga, valde obcompressa, atra, ventris costa mediana plus minusve 
erecto-pilosa, tergo glabra, dense longeque erecto-ciliata, corpore 
circ. 3.5 mm. longa, apice biaristata aristis (immaturis) lanceolatis, 
trigonis, dense antrorsumque hispidis, 1.3 mm. longis. 






REVISION OF THE GENUS COREOPSIS 339 

Type specimen: Collected by Charles Henry Tyler Townsend, 
No. A192, at altitude of 1,950-2,250 meters, Huascaray, Peru, 
September 10, 1911 (Field). 

Distribution: Bolivia and Peru. 

Specimens examined: K. Fiebrig 3476, alt. 3,200 meters, Jaipa, 
Bolivia, February 20, 1904 (Berl., 2 sheets); Townsend A192 (type, 
Field). 

40. Coreopsis fasciculata Wedd. Chlor. And. 1: 71. 1855. 
Coreopsis Matthewsii A. Gray, Proc. Amer. Acad. 5: 124. 1861. 

Involucra plus minusve tomentosa C. fasciculata sensu stricto. 

Involucra glaberrima vel basi vix sparso-hispida . . .var. laevigata. 

Frutex circ. 7.5 cm. altus, caule ramisque glaberrimis vel inter- 
dum pubescentibus, angulato-striatis, parce subtetragonis, internodiis 
saepius 1-3 cm. (ramulorum saepius 2-7 mm.) longis. Folia opposita 
(ilia ramulorum sterilium subfasciculata), petiolata petiolis tenuibus 
6-10 mm. longis, inferne saepe ciliatis raro resinosis basi in poculum 
1-3 mm. altum connatis, petiolo adjecto 1.3-2 cm. longa, tripartita, 
foliolis 2-3-fidis, segmentis linearibus, vix membranaceis, plerumque 
glabratis, apice apiculatis, 1-7 mm. longis et 0.4-1.2 mm. latis. 
Capitula ramulos (in pedunculos superne tomentosos 1-12 cm. longos 
productos) terminantia, radiata, pansa ad anthesin 2.3-2.8 cm. 
lata et 8-10 mm. alta. Involucri plus minusve tomentosi bracteae 
exteriores 6-8, spathulato-lanceolatae vel lineari-lanceolatae, apice 
obtuso saepe rotundatae, 1.3-5 mm. longae et 0.5-2 mm. latae; 
interiores lanceolatae, saepius flavo-marginatae, circ. 1 cm. longae. 
Flores ligulati circ. 7 vel 8, flavi, ligula elliptico-oblongi vel anguste 
obovati, apice subintegri, 1-1.5 cm. longi. Paleae lineari-oblance- 
olatae, apice laciniato-denticulatae, dorso villosae, circ. 6 mm. 
longae. Disci florum stigmata parce caudato-apiculata. Achaenia 
lineari-oblanceolata, plana, dorso sub palea glabra, ventre margini- 
busque sursum villosa, corpore circ. 5 mm. longa et circ. 1 mm. lata, 
biaristata aristis sursum villosis circ. 2 mm. longis. 

Type specimen: Collected by Claude Gay in Peru (Par.). 

Distribution: Ecuador (where apparently rare) to Peru and 
Bolivia. 

Specimens examined: J. Ball, alt. 3,600-3,900 meters, rocky places 
near Chicla, Peru, April 21-23, 1882 (Kew); A.Bonpland, equatorial 
America (Par., sub nom. Coreopside Matthewsii A. Gray); Otto Buch- 
tien 595, alt. 3,300 meters, lower half of Obrajes, La Paz, Bolivia, 
May 13, 1919 (Field, 2 sheets; Gray); A.E. Douglas, Arequipa, Peru, 



340 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

1892 (Gray); Mr. & Mrs. F. E. Hinkley 51, ravines at about 2,700 
meters, southern slopes of Chachani Mountain, north of Arequipa, 
Peru, March, 1920 (Gray); Hrdlicka, alt. 2,400-3,000 meters, 
western Cordillera, vicinity of San Damian, Peru, January-February, 
1913 (U.S.); E. P. Killip & A. C. Smith 21536, straggling shrub, 
2-4 ft. high, alt. 3,000-3,500 meters, open hillside, Rio Blanco, 
Department of Lima, Peru, April 15-17, 1929 (Field) ; iidem 23304, 
up to 3 ft. high, alt. 4,200 meters, open hillside between Huanta and 
Hacienda Pargora, Department of Ayacucho, Peru, May 2-19, 
1929 (Field) ; Macbride & Featherstone 436, alt. about 2,400 meters, 
rocky, grassy summit slope, Matucana, Peru, April 12-May 3, 1922 
(Field) ; iidem 690, alt. about 3,600 meters, rocky places, Rio Blanco, 
Peru, May 8-19, 1922 (Field); iidem 1237, alt. about 3,000 meters, 
rocky southwestern canyon slope, Yanahuanca, Peru, June 16-22, 
1922 (Field); Alexander Mathews 571, Purrochuca and Obrajillo, 
Peru, April to July (Kew; the basis of C. Matthewsii A. Gray); 
F. W. Pennell 13244, alt. 2,800-2,900 meters, rocky ravine above 
Arequipa, Peru, April 7-16, 1925 (N.Y.); idem 14396, alt. 2,800- 
3,200 meters, open, rocky slopes, along Rio Chillon, above Obrajillo, 
Department of Lima, Peru, June 13-23, 1925 (N.Y.) ; idem 14710, alt. 
3,300-3,500 meters, Huaros, Department of Lima, Peru, June 23, 1925 
(N.Y.) ;C.S. Sargent 27, above Chicla, Peru, December 26, 1905 (U.S.) ; 
C. Troll 3157, Paychama, Chile, March 9, 1927 (Berl.); Warszewicz 
37, Ecuador (Berl.); August Weberbauer 203, rocky places above 
3,000 meters alt., Huillacachi, southwest from Matucana, Peru, 
December 28, 1901 (Berl.); idem 2394 pro parte, alt. 3,300-3,500 
meters, Tarma, Department of Junin, Peru, February 10, 1903 
(Berl.); idem 5764, alt. 3,100 meters, between the Rio de Lomas and 
the Rio Yauca (15 10'-15 20' S. Lat.), May 15, 1911 (Berl.; Field); 
idem 5782, alt. 3,300 meters, south from Chavina, Province of Tarina- 
cochas, Department of Ayacucho, Peru, May 16, 1911 (Berl.; Field). 

Coreopsis fasciculata var. ft laevigata Sherff, Amer. Journ. 
Bot. 22: 707. 1935. 

Foliorum segmenta saepius lineari-spathulata vel anguste ob- 
longo-oblanceolata, involucris glaberrimis vel basi vix sparso-hispidis. 

Type specimen: Collected by Fortunato L. Herrera, No. Ill, at 
altitude of 3,400-3,600 meters, Cuzco, Peru (Berl.). 

Distribution: Southern Peru. 

Specimens examined: Herrera 111 (type, Berl.); idem 391, alt 
3,200-3,600 meters, environs of Cuzco, November, 1924 (Berl.; 



REVISION OF THE GENUS COREOPSIS 341 

planta tinctoria, fide lectoris); idem 1186, alt. 3,500-3,600 meters, 
heights of Cuzco, November, 1926 (Berl.); idem 1481 pro parte, 
alt. 3,350 meters, environs of Cuzco, March, 1927 (Gray; cum 
Bidente andicola var. decomposita 0. Ktze. commixta). 

41. Coreopsis lanceolata L. Sp. PI. 908. 1753; Sims in Curtis, 
Bot. Mag. pi. 2451 . 1823. Bidens Caroliniana, florum radiis latissimis, 
insigniter dentatis, etc. Martyn, Hist. PI. Rar. 26 and plate. 1728. 
Bidens succisae folio, radio amplo laciniato Dillen. Hort. Eltham. 
PL Rar. Icon, et Nom. 55, pi. 48, /. 56. 1732. Coreopsoides lanceolata 
(L.) Moench, Meth. 594. 1794. Coreopsis lanceolata var. glabella 
Michx. Fl. Bor. Amer. 2: 137. 1803. Leachia lanceolata (L.) Cass. 
Diet. Sci. Nat. 25: 389. 1822. Chrysomelea lanceolata (L.) Tausch, 
Hort. Canal./. 1 (ex DC. Prodr. 5: 570. 1836). Coreopsis lanceolata 
var. Succisaefolia DC. Prodr. 5: 570. 1836. Coreopsis lanceolata 
var. angustifolia Torr. & Gray, Fl. N. Amer. 2: 344. 1843. Bidens 
caroliniana Hemsl. in Curtis, Bot. Mag. 27, index. 1906. 

Glabra vel foliorum basibus ciliolata C. lanceolata sensu stricto. 

Pubescens vel hirsuta var. /3 villosa. 

Herba perennis, 2-6 dm. alta, caulibus erectis vel adscendentibus, 
teretibus vel subangulatis, glabris, ramosis, basim versus foliosis, 
superne subnudis ac maxime elongatis (in pedunculos desinentibus). 
Folia opposita, glabra vel basi ciliata, plerumque 5-15 cm. longa, 
superiora sessilia inferiora saepius longe tenuiterque petiolata petiolis 
laminas interdum superantibus, nunc spathulata nunc linearia vel 
lineari-lanceolata nunc lineari-oblanceolata, plerumque simplicia 
raro 1-2 lobis parvis lateralibus divisa. Capitula radiata, pansa ad 
anthesin 3-6 cm. lata et 1-1.4 cm. alta. Involucri bracteae exteriores 
8-10, lanceolatae vel oblongo-ovatae, glabratae vel apice pubescentes, 
4-8 mm. longae; interiores lanceolato-ovatae vel oblongo-ovatae, 
8-12 cm. longae. Flores ligulati circ. 8, flavi, ligula obovati vel 
cuneati, apice 3-lobati lobo mediano plerumque 2-3-dentato, 1.3- 
3 cm. longi. Paleae inferne oblongo-lineares superne filiformes, 
4-6 mm. longae. Florum tubulosorum stylorum rami caudato- 
appendiculati. Achaenia obcompressa, alata, circumambitu (alis 
paululum incurvatis inclusis) orbiculata, 2.3-3 mm. longa, corpore 
atra, ventre gibbis callosis saepe perspicue munita ac saepe tuber- 
culata vel setis nunc superne sensim dilatatis nunc apice capitatis 
setosa, apice dentibus parvis fimbriolatis bidentata. 

Type specimen: No specimen cited. The Linnean concept rested 
upon such clear and unmistakable illustrations as those given by 
Martyn (loc. cit.) and by Dillenius (loc. cit.). 



342 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Distribution: Michigan and Lake Superior southward to Florida, 
Alabama, Louisiana, southeastern Texas, and northern New Mexico; 
widely cultivated for ornament and occasionally escaping; apparently 
becoming established in eastern China. 

Specimens examined: Anon., Florida (Gray; var. angustifolia 
Ton\ & Gray, an official specimen for the treatments in Torr. & 
Gray, loc. cit. and Gray, Syn. Fl. loc. cit.); H. C. Benke 3183, Miller, 
Indiana, May 22, 1922 (Field) ; J. M. Bigelow, Fort Smith, Arkansas 
to the Rio Grande, along 35th parallel of latitude, 1853-1854 (N.Y.); 
Biltmore Herb. 2067a, dry soil near Apalachicola, Florida, May 2, 
1883 (Gray); Dr. Boykin, Georgia (Gray; N.Y.); M. A. Brannon, 
near Crown Point, Indiana, 1889 (Field); Brendel, Illinois (Berl., 
2 sheets) ; Mason Bross, Hyde Park, Chicago, Illinois, June 3, 1878 
(Field) ; S. H. & D. R. Camp, Bay View, Michigan, July 14, 1896 
(Field); H. C. Cheo & W. F. Wilson 11 p.p., near house, Mo Kan 
Shan, Province of Chekiang, China, June 19, 1926 (Calif., 2 sheets); 
Hubert Lyman Clark, Falmouth Road, Woods Hole, Massachusetts, 
August 16, 1899 (Gray); A. H. Curtiss, Pensacola, Florida, 1885 
(N.Y.); idem 1481, dry pine woods, near Aspalaga, Florida, April 
(var. villosae adpropinquans; Berl., 2 sheets; Field, 2 sheets; Gray); 
Arthur W. De Selm 140, west of Kankakee, Illinois, June 1, 1913 
(Field); Royal A. Dixon 528, vicinity of Huntsville, Texas, May 
6-12, 1910 (Field; var. angustifolia Torr. & Gray); idem & Cornelia 
Gage 713, Morgan Park Ridge, Illinois, May 27, 1907 (Field); C. W. 
Duesner, Pine, Indiana, 1908 (Field) ; F. S. Earle & C. F. Baker 243, 
Auburn, Alabama, May 8, 1897 (Field); H. Eggert, rocky hills, 
Jefferson County, Missouri, May 20, 1887 (Gray); idem, eodem 
loco, May 25, 1896 (Gray); J. H. Ehlers 518, dry, sandy soil, near 
Cecil Bay, Emmet County, Michigan, July 14, 1917 (Gray) ; George 
Engelmann, prairies, vicinity of East St. Louis, Illinois, May, 1845 
(Berl.);FrcmA; C. Gates 508, Ravenswood, Chicago, Illinois, June 23, 
1905 (Field); idem 2478, sandy soil, Waukegan, Illinois, June 8, 
1908 (Field) ; idem & M. T. Gates 10547, along sandy road, Cecil Bay, 
Emmet County, Michigan, July 14, 1917 (Field); Jesse M. Green- 
man 2017, sandy soil near Lake Michigan, Beach, Illinois, June 16, 
1907 (Gray); idem 2592, Dune Park, Indiana, June, 1910 (Gray); 
Dr. Hale, Louisiana, 1842 (Gray); Elihu Hall, sandy barrens, 
Athens, Illinois, 1861 (Gray, cum var. villosa commixta, sed alibi 
tantum species ipsa); Roland M. Harper 22, Screeton, Arkansas, 
May 1, 1923 (Gray); herb. Hexamer & Maier, rich woods, June 16, 
1855 (Gray); E. J. Hill, dry grounds, Morgan Park, Chicago, 



REVISION OF THE GENUS COREOPSIS 343 

Illinois, June 17, 1876 (Field); Albert S. Hitchcock, Madison County, 
Florida, June-July, 1898 (Field); Hart. Berol, September, 1832 
(Berl.) ; Mary E. Hutchinson, Randolph, Wisconsin (Field) ; L. N. 
Johnson, sandy ridges in woods, Chicago, Illinois, June 19, 1891 
(Gray); A. B. Langlois, western Louisiana, April, 1880 (Field); 0. E. 
Lansing, Jr. 848, along railroad, Clarke, Indiana, May 31, 1900 
(Field); idem 935, eodem loco, June 20, 1900 (Field); idem 2715, 
swale near railroad, Indiana Harbor, Indiana, June 17, 1908 (Field) ; 
idem 2782, eodem loco et tempore (Gray, var. villosae adpropin- 
quans); /. A. Lapham, Milwaukee, Wisconsin (Gray); Le Conte, 
North America (Berl.); George W. Letterman, top of sandhill near 
Franklin County Road Bridge, Missouri, July, 1883 (Mo.); C. 0. 
Levine 778, Honam Island, Kwang-Tung Province, China, May 12, 
1917 (Gray; Mo.); L. H. Lighthipe, Monticello, Florida, April 5, 
1891 (Field); F. Lindheimer, wet woods west of Houston, Texas, 
early May, 1840 (Berl.) ; Ray N. Lloyd, rich or damp soil, Ravens- 
wood, Chicago, Illinois, June 3, 1887 (Field); Loring, Lake Superior 
(Gray); W. H. Manning, Mackinac Island, Michigan, July 11, 
1915 (Gray); F. E. McDonald, dry, gravelly slopes, Peoria, Illinois, 
June, 1899 (Field); idem, high, dry hills overlooking Horse Shoe 
Bottom, Peoria, June, 1904 (Gray) ; idem, high, gravelly hills, Peoria, 
June, 1915 (Field) and June, 1916 (Field) ; S.B. Mead, Mason County, 
Illinois, June 18, 1845 (Field; var. angustifolia Torr. & Gray); W. S. 
Moffatt 105, sandy thickets, Chicago, Illinois, June, 1895 (Berl.; 
Gray); H. F. Munroe, Hyde Park, Chicago, Illinois, May (Field) 
and June, 1879 (Field); E. J. Palmer 17, cherty barrens, Newton 
County, Missouri, July 1, 1906 (Gray); idem 2386, cherty barrens, 
Reding's Mill, Missouri, July 1, 1909 (Gray); idem 2386 A, eodem 
loco et tempore (Gray); A. S. Pease 17993, damp, sandy beach, 
Bailey's Harbor, Door County, Wisconsin, June 18, 1921 (Gray); 
S. F. Poole, Sharon, Massachusetts, July, 1905 (Gray); Thomas C. 
Porter, banks of Savannah River, near Augusta, Florida, April, 
1847 (Gray) ; P. H. Rolfs 309, Tallahassee, Florida, April 20, 1895 
(Field); B. F. Saurman, rich, grassy plains, Apalachicola, Florida 
(Field; var. angustifolia Torr. & Gray); F. Scammon, Chicago, 
Illinois, June, 1859 (Field) and June 3, 1859 (Field) ; A. K. Schindler 
345a, alt. 1,100 meters, Lu-shan, Kuling Mountains, Province of 
Kiangsi, China, August-September, 1908 (Berl.); J. H. Schuette, 
Door, Wisconsin, July 30, 1887 (Field, 2 sheets) ; idem, shore of Lake 
Michigan north of Sturgeon Bay Canal, Wisconsin, July 14, 1890 
(Field) -EarlE. Sherff5006, cult, in Harvard Univ. Bot. Gard., August 
12, 1929 (Berl.; Brit.; Field); H. C. Skeels 304, Joliet, Illinois, June 



344 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

21, 1904 (Field); Paul C. Standley 4410, alt. about 2,460 meters, 
hillside near Cowles, Pecos River National Forest, New Mexico, 
July 19, 1908 (Gray) ; A. N. Steward, open hillside, Lu-shan, Ruling, 
Province of Kiangsi, China, July 27, 1922 (Calif.); idem 1518, in 
open, Kikungshan, Province of Honan, China, July 24, 1925 (Calif.) ; 
idem 1583, damp meadow, eodem loco, July 27, 1925 (Calif.); W. F. 
Thurrow, Hockley, Texas, 1890 (Field) ; S. M. Tracy 9454, St. Andrews, 
Florida, April 29, 1908 (Gray) ; idem & Lloyd 508, Virginia, August 
15, 1910 (Field; Gray);L. M. Umbach, dry sands, Miller, Indiana, 
June 5, 1897 (Field); idem, sands, Clarke, Indiana, June 4, 1898 
(Field); H. E. Wheeler 82, near Hazen, Grand Prairie, Arkansas, 
May 22, 1924 (Field); H. N. Whitford 64 and 203, waste fields, Cold 
Spring Harbor, New York, August, 1903 (Field) ; Emile F. Williams, 
self-sown and thriving, Concord, Massachusetts, June 20, 1908 
(Gray); Wilbur H. Wright 50, Miller, Indiana, July 2, 1908 (Field). 

Coreopsis lanceolata var. /3 villosa Michx. Fl. Bor. Amer. 2: 
137. 1803. C. crassifolia Dryand. in Ait. Hort. Kew. ed. 1. 3: 252. 
1789; non Sess & Moc. Fl. Mex. ed. 2. 194. 1894. C. oblongifolia 
Nutt. Journ. Acad. Phila. 7: 76. 1834. C. lanceolata var. crassifolia 
Ait. ex Heynhold, Nomencl. 219. 1840. 

E specie planta omnino vel fere usque ad summam pubescente 
differt. 

Type specimen: Collected by Andre Michaux in Carolina (Par.). 

Distribution: South Carolina, Illinois, Missouri, and western 
Arkansas, southwardly to Florida, Alabama, and Louisiana; else- 
where probably introduced. 

Specimens examined: Anon., Tampa Bay, Florida (Gray); H. H. 
Babcock, Chicago, Illinois, June 17, 1874 (Field); J. M. Bigelow, 
Fort Smith, Arkansas, June 23, 1853 (N.Y.); Biltmore Herb. 5693b, 
dry soil, Spartanburg, South Carolina, May 12, 1897 (Gray) ; B. F. 
Bush 84, common in barrens, Eagle Rock, Missouri (Gray) ; William 
M. Canby 55, Columbia, South Carolina, May 9, 1899 (Gray); 
A. W. Chapman, middle Florida (Field); A. H. Curtiss 6390, old 
field near River Junction, Florida, May 2, 1898 (Gray) ; H. Eggert, 
stony hills, Jefferson County, Missouri, June 10, 1891 (Gray; Mo.); 
idem, Hematite, Missouri, May 25, 1896 (Calif.; Field; U.S.); herb. 
George Engelmann, Aiken, South Carolina, April, 1882 (Mo.}; Frank 
C. Gates 942, Chicago, Illinois, June 24, 1905 (Field); idem 2817, 
Beach, Illinois, July 1, 1908 (Field) ;C. A.Geyer, Beardstown, Illinois 
(Mo., 2 sheets); Grossly, Hyde Park, Chicago, Illinois, June 10, 






REVISION OF THE GENUS COREOPSIS 345 

1877 (Field); Dr. Hale, Louisiana (Field; Gray); E. J. Hill 1108, 
top of limestone ledge, Lockport, Illinois, June 14, 1899 (Field); 
idem 2746, eodem loco et tempore (Gray) ; G. G. Kennedy, dry pine 
woods, Ormond, Florida, March 29, 1897 (Gray) ; 0. E. Lansing, Jr. 
24-3, waste ground, Hyde Park, Chicago, Illinois, June 14, 1898 
(Field) ; G. W. Letterman, Allenton, Missouri, August, 1883 (Gray) ; 
idem, Merely, Missouri, July, 1884 (Mo., 2 sheets) ; F. E. McDonald, 
high, dry hills, Peoria, Illinois, May, 1915 (Field); Mrs. J. M. 
Milligan, Jefferson County, Missouri (Field); E. J. Palmer 27056, 
rocky, open ground, along stream near Berry ville, Arkansas, May 7, 
1925 (Gray) ; H. W. Ravenel, San tee Canal, South Carolina (Gray) ; 
F. Scammon, Chicago, Illinois (Field); Earl E. Sherff 5013, cult, in 
Harvard Univ. Bot. Gard., August 12, 1929 (Brit.; Calif.; Field); 
idem 5014., cult., eodem loco et tempore (Field; Kew); idem 5016, 
cult., eodem loco et tempore (Calif.; Field; Kew; N.Y.; U.V.); 
Mary F. Spencer 1114, alt. 300 meters, ditch near San Bernardino, 
California, May 9, 1919 (Gray) ; E. F. Williams, well established, 
Wickford, Rhode Island, June 17, 1908 (Gray) ; R. E. Woodson, Jr. & 
E. S. Anderson 1545, Birmingham, Alabama, April 20, 1927 (Mo.). 

42. Coreopsis corninsularis Sherff, Bot. Gaz. 94: 597. 1933. 

Herba perennis, erecta, gracilior, caulibus tenuibus angulatis 
glabris 2 vel 3 ex unico basi circ. 3-4 dm. altis. Folia opposita, 
plus minusve petiolata petiolis tenuibus usque ad 3 cm. longis basi 
sparsim hispido-ciliatis, petiolo adjecto plerumque 3-8 cm. longa 
et 1-6 mm. lata, saepius integra anguste vel late spathulato-linearia 
crassiuscula faciebus glabra marginibus ciliata apice subacuta vel 
subobtusa, interdum 3-5-partita foliolis lateralibus multo minoribus 
et tantum 1-2 mm. latis. Capitula solitaria longe pedunculata 
pedunculis tenuibus glabris 1-1.8 dm. longis, radiata, pansa ad 
anthesin 3-5 cm. lata et 10-12 mm. alta. Involucri glabri bracteae 
exteriores 8-10, lanceolatae vel oblongo-lineares, 3-5 mm. longae; 
interiores ovato-oblongae, demum circ. 1 cm. longae. Flores ligulati 
circ. 8, flavi, ligula oblongo-oblanceolati, apice saepius 3-lobati lobo 
mediano multo majore apice rursus in 2 dentes inciso, 1.5-2.5 cm. longi. 
Paleae tenuiter lineares, superne filiformes, 3-4 mm. longae. Flores 
tubulosi flavi, styli ramis terminaliter caudato-appendiculatis. 
Achaenia valde obcompressa, dorso convexa, nigra, corpore ipso 
oblongo, 2.5-3.2 mm. longa et (alis exclusis) 1.2-1.5 mm. lata, facie- 
bus glabra vel glabrata, marginibus alata alis expansis membranaceis 
quam corpore saltern 0.5-0.75 angustioribus, apice papposa 2 
fimbriatis squamellis circ. 0.1-0.2 mm. longis. 



346 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: Collected by Samuel Mills Tracy, No. 8542, 
on Horn Island, Mississippi, May 25, 1903 (Mo.). 

Distribution: Known only from type locality on Horn Island, 
Mississippi. 

Specimens examined: Tracy 8542 (type, Mo.; cotype, Field). 

43. Coreopsis debilis Sherff, Bot. Gaz. 89: 366. 1930. 

Herba perennis, caulibus saepius 2-6, e radice lignescenti erectis, 
tenuissimis, angulatis, glabris, 3-6 dm. altis. Folia opposita, 
membranacea, petiolata petiolis tenuibus usque ad 2 cm. longis, 
lamina nunc indivisa linearia vel parce lineari-oblanceolata margine 
integra ciliataque apice vix acuta basim versus sensim angustata 
faciebus saepe aegre hispida petiolo adjecto 2-5 cm. longa et 1-5 mm. 
lata; nunc majora, petiolo saepe 3.5 cm. longo adjecto 7-9 cm. longa, 
aegre pinnata (raro etiam subbipinnata) segmentis lateralibus ple- 
rumque 1-2 jugis, quam terminal! minoribus, linearibus vel subfili- 
formibus. Capitula pauca vel numerosa, tenuiter pedunculata 
pedunculis 1-2 (-3) cm. longis, radiata, pansa ad anthesin circ. 
2-2.5 cm. lata et circ. 7-9 mm. alta. Involucri saepius glabrati 
bracteae exteriores circ. 8, lineares lanceolataeve, apice acutae vel 
obtusae, margine ciliatae, basim versus saepe scariosae, 3-4 mm. 
longae; interiores lanceolatae vel ovato-lanceolatae, 7-8 mm. longae. 
Flores ligulati flavi, ligula cuneato-obovati, apice saepius 4-dentati 
dentibus acutis, circ. 1 cm. longi. Paleae lineares, superne angustis- 
sime elongatae. Disci florum stigmata terminaliter caudata. 
Achaenia obcompressa, tergo papillato-rugoso convexa, corpore 
ipso atro circ. 2 mm. longa et circ. 1 mm. lata, marginibus anguste 
alata, apice 2-squamellata squamellis fimbriolatis; facie ventrali 
glabra vel papillato-scabra, supra infraque valde callosa. 

Type specimen: Collected by John Donnell Smith, No. 602, in 
dry, sandy, old fields along coast of Mississippi Sound, Harrison 
County, Mississippi, September 15, 1885 (Field). 

Distribution: Florida and Georgia westward to Mississippi. 

Specimens examined: A. H. Curtiss, Pensacola, Florida, summer 
of 1885 (N.Y.) ;F. E. Lloyd & S. M. Tracy 527a, Gulfport, Mississippi, 
September 8, 1900 (N.Y.); Charles Mohr, dry, open, sandy hills, 
etc., eodem loco, July 4, 1874 (U.S., 2 sheets); idem, in woods, Cull- 
man, Alabama, June 1, 1882 (Gray) ; idem, low, open places, Cullman, 
June, 1883 (U.S., 3 sheets); idem, dry hillsides, vicinity of Ashland, 
Bibb County, Alabama, June, 1883 (U.S.) ; idem, borders of woods and 
fields, dry openings, Mulberry River Valley, Blount County, Ala- 



REVISION OF THE GENUS COREOPSIS 347 

bama, June 6, 1883 (U.S.) ; idem, sandy soil in open copses, Columbus, 
Mississippi, June 4, 1888 (U.S.); idem, dry bald prairies, Gallion, 
Alabama, May 25, 1893 (U.S.); Charles L. Pollard & William R. 
Maxon 4-96, Lithonia, Georgia, August 12, 1900 (N.Y.) ; John Donnell 
Smith 602 (type, Field; cotype, Gray). 

A species more closely related to Coreopsis lanceolata L. and C. 
grandiflora Hogg. J. Donnell Smith had determined his plant as C. 
lanceolata var. angustifolia Torr. & Gray. That variety, however, 
has a very different habit, the plants being less branched and the 
leaves being basally clustered and almost always simple. Donnell 
Smith had shown Asa Gray his Mississippi material, and Gray had 
called it "a peculiar form" of C. lanceolata L. Mohr's material from 
Pensacola, Florida, likewise had been seen by Gray. Thus, one 
sheet bears a note stating: "Dr. Gray remarks: 'I have nothing like 
this. I suppose it may be an altered, coast form of C. lanceolata. 
But it is very peculiar.' ' (This note is written on the back of a card 
which, by strong transmitted light, is seen to have been Donnell 
Smith's regular herbarium label. This would seem to indicate that 
Mohr and his contemporary had studied their several specimens 
mutually and had regarded them as identical.) This same sheet has 
Mohr's first determination, Coreopsis lanceolata var. angustifolia, 
but this he had subsequently crossed out and replaced with C. 
grandiflora. His other foregoing specimens all had been labeled by 
him as C. grandiflora. This interpretation was retained by him in 
his Plant Life of Alabama (Contr. U. S. Nat. Herb. 6: 805. 1901). 
From C. grandiflora, however, our plants differ in their more slender 
and wiry, less herbaceous stems, in the tendency for some entire 
stems to have minute, simple leaves (smaller than the simple leaves 
formed in C. lanceolata), and in the tiny achenes, which have the 
body proper about 2 mm. long and about 1 mm. wide, as against 
about 2.5 mm. long and about 1.4-1.8 mm. wide in C. grandiflora. 
(A similar or even more pronounced achenial difference separates 
C. debilis also from C. lanceolata. Furthermore, the achenes of C. 
debilis have narrow wings [0.2-0.5 mm. wide] while those of C. 
grandiflora and C. lanceolata have wings 1 mm. or so in width.) 

44. Coreopsis intermedia Sherff, Bot. Gaz. 88: 299. 1929. 

Herba erecta, plus minusve glabra, forsitan perennis, supra parce 
ramosa, 6 dm. alta, caule subtereti vel moderatim angulato, 
sulcato. Folia opposita, simplicia, basalia longe tenuissimeque 
petiolata petiolis usque ad 4.5 cm. longis, laminis oblongo-oblance- 
olatis vel anguste obtuseque obovatis; principalia caulina sessilia, 



348 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

late oblongo-lanceolata, ciliata, crassiuscula, apice subacuta, 5-7 
(-9.5) cm. longa et 1.2-2 (-3.2) cm. lata. Capitula pedunculata 
pedunculis 1.5-2.5 dm. longis, radiata, pansa ad anthesin 3.5-4 
dm. lata et 12 mm. alta. Involucri bracteae exteriores 8-10, 
lanceolatae vel lineari-lanceolatae, tergo glabratae, margine saepe 
diaphana ciliatae, apice acutae cartilagineaeque, 4-7 (rariter -8) 
mm. longae; interiores late lanceolatae, plerumque 12-14 mm. 
longae. Flores ligulati circ. 8, flavidi, unicolores, obovati vel late 
oblanceolati, apice trilobati lobo mediano valde emarginato, circ. 
1.5 cm. longi. Paleae superne elongatae et valde caudato-attenu- 
atae. Disci florum stigmata apice caudata. Achaenia suborbi- 
cularia, valde obcompressa, dorsaliter convexa, brunneo-atra, alata 
alis membranaceis planis vel rarissime parce incurvatis 0.2-0.4 mm. 
latis, apice saepe bidenticulata, faciebus perspicue tuberculata, 
facie ventrali raro callosa, corpore ipso oblongo-obovato vel late 
oblongo-oblanceolato 2-3 mm. longa et 1.3-2 mm. lata. 

Type specimen: Collected by Julian Reverchon, No. 2077 pro 
parte, in sandy woods, Mineola, Texas, June 12, 1900 (Berl.). 

Distribution: Known only from type locality and vicinity, Texas. 

Specimens examined: Reverchon, Pine Island, May 5, 1903 (Mo.); 
idem 2041, sands, Big Sandy, May 27, 1901 (Mo., 2 sheets); idem 
2077 pro parte (type, Berl.; cotype, Mo.); idem 2077 pro parte, 
sands, Mineola, June 10, 1900 (Mo.). 

The stems of the plants examined are uniformly leafy from bottom 
to top as in Coreopsis pubescens and the leaves resemble the glabrous, 
undivided ones occasionally found in that species. The elongate 
peduncles, however, also the pronounced differentiation between 
exterior and interior involucral bracts, are more as in C. lanceolata, 
although not typical for that species. 

45. Coreopsis pubescens Ell. Sketch Bot. S. Carol. & Georgia 
2: 441. 1824. C. auriculata Schkuhr, Handb. ed. 2, pi. 260. 1808 
(non L.). C. auriculata var. 7 Torr. & Gray, Fl. N. Amer. 2: 343. 
1843. C. auriculata var. 8 Torr. & Gray, op. cit. 344 (excl. syn. quod 
C. pubescens var. robusta est). 

Plus minusve pubescens vel hirsuta. . . .C. pubescens sensu stricto. 
Caulis ramique glabri vel glabrati var. robusta. 

Herba perennis, erecta, simplex vel paulum ramosa ramis sub- 
erectis, plus minusve pubescens vel hirsuta, 6-12 dm. alta, caule 
usque ad summam folioso. Folia opposita, polymorpha, saepius 
5-10 cm. longa, inferiora longe (2-5 cm.) tenuiterque petiolata altera 



REVISION OF THE GENUS COREOPSIS 349 

saepe sessilia, laminis nunc integris ovalibus vel oblongo-ovatis vel 
elliptico-lanceolatis nunc (plus minusve irregulariter) 3-5-partitis 
foliolis lateralibus plerumque multo minoribus ac oblonge lineari- 
lanceolatis. Capitula tenuiter pedunculata pedunculis 1-2 dm. 
longis, radiata, pansa ad anthesin 3-5 cm. lata et 8-11 mm. alta. 
Involucri bracteae subaequales plus minusve glabratae, exteriores 
8-10, lineari-lanceolatae, plurivittatae, 7-10 mm. longae; interiores 
ovatae. Flores ligulati circ. 8, flavi, ligula cuneati vel oblongo- 
cuneati, apice 3-lobati lobo mediano rursus 2-4-lobulato, 1-2.3 cm. 
longi. Paleae inferne latiores superne elongato-filiformes, demum 
6-8 mm. longae, achaenia multum superantes. Florum tubulosorum 
stylorum rami apicem versus abrupte angustati et lineari-appendi- 
culati. Achaenia obcompressa, suborbiculata, circ. 2.8-3 mm. longa, 
corpore ipso nigra, alis (0.5 mm. latis) planis integris saepe pur- 
purea, faciebus glabra vel papillata, ventraliter apice basique saepius 
gibbo cartilagineo munita, apice vero squamellis fimbriatis plerumque 
2-dentata. 

Type specimen: None cited, but habitat given as "western dis- 
tricts of Georgia." 

Distribution: Virginia, Illinois, and Oklahoma southward to 
Florida, Alabama, and Louisiana. Sometimes escaped or adventive 
elsewhere. 

Specimens examined (it may well be that these determinations, 
most of which were made many years ago, include a few specimens 
which are properly referable to the var. robusta) : Samuel M. Bain 95 
p.p., fields, Henderson, Tennessee, May, 1893 (Gray); H. C. Beardslee 
& C. A. Kofoid, alt. 900 meters, Great Smoky Mountains, Swain 
County, North Carolina, August, 1891 (Field; Gray); Beyrich, 
grassy banks of rivers, Georgia (Berl., 2 sheets);/ 7 . Boynton, High- 
lands, North Carolina, 1889 (Gray; Mo.); Buckley, Alabama (Gray); 
B. F. Bush 172A, uncommon, in dry ground, McDonald County, 
Missouri, September 1, 1893 (Gray); idem 6041, woods, Webb City, 
Missouri, July 23, 1910 (N.Y.) ; idem 6137, woods, Monteer, Missouri, 
August 8, 1910 (Gray); idem 6531, eodem loco, October 22, 1911 
(Mo.); idem 7872, open woods, eodem loco, October 8, 1916 (Gray); 
Carpenter, Louisiana (Gray); A. W. Chapman, mountains of Georgia 
(Field); A. H. Curtiss 17, rich woods, Jackson County, Florida, 
June 18, 1886 (Gray; foliis sed non achaenis C. auriculatae adpropin- 
quans); H. Eggert, dry woods, Jefferson County, Missouri, July 8, 
1879 (Gray) ; George Engelmann, Mountain Creek bottom, Missouri, 
1833 (Mo.); idem, St. Louis, Missouri, August, 1863 (Berl.); Henry 



350 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

A. Gleason 2749, dry, upland woods, Makanda, Illinois, July 22, 1902 
(Gray); idem 2750, cleared uplands along railroad, eodem loco et 
tempore (Gray) ; P. Graebner, cult, in Hort. Berol. ex sem. ex Amer. 
Boreali missis, August 5, 1910 (Berl.); J. M. Greenman 3771, Shef- 
field Mountain, near Arcadia, Missouri, July 22, 1915 (Mo.); H.E. 
Hasse, Desoto, Missouri, July, 1887 (Field); Elihu Hall, central 
Illinois (Gray); W. Hoffman, Corn Creek, vicinity of Rolla, Mis- 
souri, July 15, 1871 (Berl., 2 sheets); Hort.Berol, 1806-1812 (Berl.); 
Hort. Lips, ex Amer. septentr. (Berl.); Hort. Paris, 1816 (Berl.); 
H. W. Houghton 3864, common, open woods, near Shawneetown, 
Oklahoma, May 28, 1916 (Gray); H. D. House 4023, below 900 
meters alt., sandy soil along A very 's Creek, Pisgah Forest, North 
Carolina, September, 1908 (Gray); John H. Kellogg, Allenton, 
Missouri, August 12 and 13, 1884 (Mo.) ; idem, St. Louis, Missouri, 
July 15, 1903 (Mo.); George W.Letterman, Allenton, Missouri (Mo.); 
Kenneth K. Mackenzie 328, common in woods, Pleasant Grove, 
Missouri, July 21, 1897 (Field); Michaux, North America (Berl.); 
E. J. Palmer 2317 and 2317 A, open woods, Forest Mill, Missouri, 
June 23, 1909 (Gray); idem 2513, barrens, Joplin, Missouri, July 20, 
1909 (Gray) ; idem 29347, rich, moist soil, vicinity of Beaver, Arkan- 
sas, October 24, 1925 (Gray); Thomas C. Porter, Hawk's Nest on 
New River, 9 miles above Kanawha Falls, West Virginia, July 21, 
1880 (Field; N.Y.); Ns. Riehl 65, very moist ground, St. Louis, 
Missouri, July, 1838 (Berl.); F. Rugel, in rocky, wet places of the 
mountains at Broad River, North Carolina, June-July, 1841 (Berl. ; 
N.Y.); idem, in open places and along roads, between Quincy and 
Aspalaga, Florida, May, 1843 (Mo.); Albert Ruth 655, along railroad, 
Valley Head, Alabama, July, 1898 (Mo.); idem 665, Lulu Falls, 
Lookout Mountain, Georgia, July, 1898 (N.Y.); E. M. Shepard, 
Greene County, Missouri, July, 1880 (Gray); Earl E. Sherff 989, 
southeast of Pacific, Missouri, August 7, 1910 (Gray); idem 5005, 
cult., Harvard Univ. Bot. Gard., August 12, 1929 (Berl.; Field); 
C. W. Short 42, Lexington, Kentucky, 1831 (Gray) ; John K. Small, 
alt. 750-1,500 meters, on slopes of Thomas Bald, boundary of North 
Carolina and Georgia, August 19, 1893 (Field); idem, alt. 1,560 
meters, summit of Thomas Bald, August 19, 1893 (Field) ; idem, base 
of Currahu Mountain, Toccoa, Georgia, July, 1896 (N.Y.); Huron 
H. Smith 1089, along railway tracks, Des Arc, Missouri (Field); 
idem 2456, Blue Ridge Mountains, Fannin County, Georgia, July 22, 
1909 (Field) ; idem 2582, eodem loco, August 7, 1909 (Field) ; John 
Donnell Smith, alt. 1,410 meters, north face of cliffs, Wildcat Ridge, 



REVISION OF THE GENUS COREOPSIS 351 

west flank of Whitesides Mountain, 6 miles northeast from High- 
lands, North Carolina, August 21, 1882 (Gray); idem, alt. 1,440 
meters, face of cliffs, eodem loco et tempore (Field; Mo.); idem, 
alluvial banks of river at Cumberland Falls, Whitler County, 
Kentucky, August 23, 1883 (Field) ; idem, thickets, alluvial banks of 
Soquee River, Habersham County, Georgia, September 13-15, 
1883 (Field; Gray); idem, edge of woods, banks of Oconee River, 
above Milledgeville, Georgia, August 16, 1884 (Field; Gray); idem, 
gullies in high, wooded hills west of Cahawba River, Perry County, 
Alabama, September 2, 1885 (Field; Gray); G. W. Stevens 2387, 
shady, grassy thicket in woods near Ottawa, Oklahoma, August 27, 
1913 (Gray); Roland Thaxter, Cullowhee, North Carolina, June- 
July, 1887 (Gray); R. E. Woodson, Jr. 655, Crawford County, 
Missouri, July 10, 1926 (Mo.). 
Passes into the var. robusta. 

Coreopsis pubescens var. /3 robusta A. Gray ex Eames, 
Rhodora 18: 239. 1916. C. testudinea Shuttl. in herb. 
E specie caule ramisque glabris vel glabratis differt. 

Type specimen: Collected by Asa Gray and John Carey, North 
Branch of New River, North Carolina, July, 1841 (Gray). 

Distribution: Virginia and North Carolina westward to 
Kentucky. 

Specimens examined : Biltmore Herb. 307c, slopes of Cedar Cliff 
Mountain, Buncombe County, North Carolina, August 2, 1897 
(Gray; Mo.); John Carey, North Carolina (N.Y.); idem & Asa 
Gray, North Branch of New River, etc. (type, Gray; cotype, N.Y.); 
iidem, Virginia, July, 1841 (Gray) ; E. H. Eames 8825, growing about 
1 meter tall, waste grounds, Bridgeport, Connecticut, July 21, 1914 
(Gray) ; M. L. Fernald & Bayard Long 19265, fallow field northeast of 
East Brewster Station, Brewster, Massachusetts, July 20, 1919 
(Gray); P. A. Rydberg 9445, alt. 1,650-1,800 meters, Craggy Moun- 
tains, Buncombe County, North Carolina, July 21, 1925 (N.Y.; 
Pom.); John Donnell Smith, growing 4 ft. high, upland meadow, 
Richland Gap, Heywood County, North Carolina, August 14, 1882 
(Gray; Mo.) ; idem, alluvial soil and sandy bank of river, Cumberland 
Falls, Whitley County, Kentucky, August 23, 1883 (Gray; Mo.). 

Originally regarded by Asa Gray (Amer. Journ. Sci. 42: 45. 1841) 
as a ''larger form of Coreopsis auriculata, with nearly all the leaves 
undivided." At times very distinct, again scarcely distinguishable 
from the species proper. 



352 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

46. Coreopsis heterolepis Sherff, Bot. Gaz. 89: 365. 1930. 
Herba perennis, glabra, caule angulata, ramosa, dense foliosa, 

3-6 dm. alta. Folia membranacea plus minusve dimorpha: basalia 
dense congregata, petiolata petiolis planis angustissime alatis usque 
ad 6 cm. longis, petiolo adjecto 8-12 cm. longa, indivisa spathulata 
vel oblonga apice obtusa, alia pinnata foliolis nunc linearibus nunc 
oblanceolatis nunc oblongis nunc rhomboideo-subovatis ac etiam 
1-1.7 cm. latis, lateralibus saepius 1-2 jugis ac multo minoribus; 
caulina numerosissima, petiolata petiolis anguste marginatis marginis 
basi rarius setis pluriloculatis sparsim positis obsita circ. 1-2.5 cm. 
longis petiolo adjecto circ. 6-10 cm. longa, plerumque pinnata, foliolis 
anguste linearibus, 2-8 cm. longis et 0.3-2.5 mm. latis, apice sub- 
acutis, lateralibus 1-2 jugis. Capitula tenuissime pedunculata 
pedunculis 1-2 dm. longis, radiata, pansa ad anthesin 3-4 cm. lata 
et 5-7 mm. alta. Involucri glabri vel subglabri bracteae exteriores 
angustissime lineares, saepius patentes, apice acutae, 5-11 mm. 
longae; interiores ovato-lanceolatae nunc breviores nunc longiores. 
Flores ligulati circ. 8, omnino flavi, ligula cuneato-obovati, apice 
obtuse 4-dentati, circ. 1.5-1.8 cm. longi. Paleae superne capillares, 
circ. 4-5 mm. longae. Disci florum stigmata caudato-elongata. 
Achaenia minima plano-convexa, corpore nigro oblongo vel oblongo- 
ovato ipso tantum circ. 1.3-1.7 mm. longa et circ. 1-1.5 mm. lata, 
faciebus levia vel papillato-rugosa, marginibus anguste alata alis 
rubris circ. 0.2-0.4 mm. latis, apice minutissime 2-squamellata 
squamellis fimbriolatis. 

Type specimen: Collected by Ernest Jesse Palmer, No. 6962A, 
on dry, sandy bluffs, Heber Springs, Cleburne County, Arkansas, 
October 30, 1914 (Mo., 4 sheets). 

Distribution: Known only from type locality. 

Specimens examined: Palmer 6962A (4 type sheets, Mo.). 

47. Coreopsis grandiflora Hogg ex Sweet, Brit. Fl. Card. 2: 
pi. 175. 1826; Nutt. Trans. Amer. Phil. Soc. 2, 7: 358. 1841. C. 
Boykiniana Nutt. loc. cit. C. heterophylla Nutt. loc. cit. (nee alior.). 
C. grandiflora var. subintegrifolia Torr. & Gray, Fl. N. Amer. 2: 345. 
1843. 

Caules foliaque pubescentia vel piloso-hispida var. 7 pilosa. 

Caules et foliorum facies plerumque glabrata. 

Foliorum segmenta omnia anguste linearia vel quidem filiformia. 

var. 8 Harveyana. 

Foliorum simplicium lamina vel divisorum segmenta latiora. 



REVISION OF THE GENUS COREOPSIS 353 

Folia caulis basim versus adgregata, pedunculis crassis elongatis 

(saepe 2-3 dm. longis) -. var. /3 longipes. 

Folia subaequaliter distributa, pedunculis tenuioribus saepius 
0.8-1.5 dm. longis C. grandiflora sensu stricto. 

Herba perennis vel raro annua, erecta vel adscendens, plerumque 
non pubescens nisi petiolis ciliatis, 3-6 dm. alta, saepe ramosa, 
plerumque foliosa. Folia opposita, plus minusve petiolata petiolis 
saepius 0.5-4 cm. longis, inferiora simplicia vel irregulariter partita 
lamina (vel segmento terminali) spathulata vel lanceolata, petiolo 
adjecto saepius 5-10 cm. longa; alia plerumque 3-5-partita (raro 
simplicia) segmentis interdum rursus 3-5-partitis segmentis ultimis 
linearibus vel lanceolato-linearibus, plerumque 1.2-5 mm. latis. 
Capitula saepius solitaria, moderate pedunculata pedunculis tenui- 
bus saepius 1-1.5 dm. longis, radiata, pansa ad anthesin 3-6 cm. lata 
et 8-12 mm. alta. Involucri bracteae exteriores 7-10, saepe dupli- 
citer dispositae, lanceolato-subulatae, infra saepe subdilatatae, mar- 
gine ciliatae et plus minusve albidae, 5-9 (raro -18) mm. longae; 
interiores ovatae, nunc paulo longiores nunc paulo breviores. Flores 
ligulati plerumque 8, flavi, ligula cuneato-obovati, apice 3-lobati 
lobo mediano majore rursus 2-partito, 1.3-2.5 cm. longi. Paleae 
lineares, infra latiores supra attenuatae filiformesque, demum circ. 
6-7 mm. longae. Flores tubulosi summa aurantiaci, stylorum ramis 
apice perspicue cuspidati. Achaenia obcompressa et dorso convexa, 
circumambitu (alis inclusis) orbiculata, circ. 2.5 mm. longa, corpore 
nigra, faciebus glabra vel minute papillata, ventraliter apice basique 
saepe verruca callosa magna ornata, marginibus alata alis planis vel 
vix incurvatis quam corpore dimidio angustioribus, apice minute 
biaristata aristis squamellatis fimbriolatisque vel demum calva. 

Type specimen: Sent by a Mr. Hogg from New York to England 
as a new species and represented by Robert Sweet's type plate 
(loc. cit.). 

Distribution: Missouri and Kansas southward to Florida, 
Louisiana, and New Mexico. 

Specimens examined: H. C. Benke 3279, Pensacola, Florida, 
March 9, 1922 (Field) ; George D.Butler 16, prairies, 1.5 miles north 
of Limestone Gap, Oklahoma, June 13, 1877 (Field) ; F. S. & Esther 
S. Earle 73, 4 miles northwest of Auburn, Alabama, July 13, 1899 
(Mo.);EsemE. Hall, Texas, 1877 (Gray); W. H. Haller 982, low 
ground, Wilson County, Kansas, 1895 (Gray) ; ex Hort. Berol. (Gray) ; 
H. W. Houghton 3567, common on hillsides, vicinity of Tishomingo, 
Oklahoma, April 15, 1916 (Gray); Dr. Leavenworth, Louisiana 



354 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

(Gray); Le Conte, North America (Gray); George W. Letterman, 
Gratiot, Missouri, May 30, 1897 (Field) ; idem, along Frisco Railroad 
near St. Louis, Missouri, August, 1899 (Mo.)',Lindheimer, Houston, 
Texas, April, 1842 (Gray); Kenneth K. Mackenzie 100, frequent, 
introduced along railroads, Kansas City, Missouri, June 5, 1898 
(Field); Dr. Pitcher, Arkansas (Gray); J. Reverchon, sandy loams, 
woods, Dallas, Texas, May, 1874 (Mo.); idem, Dallas, May, 1876 
(Gray); idem 513, sandy forests, Dallas, May, 1883 (Mo.); idem 
1478, rich sands, Dallas, May (Berl.; Field); Albert Ruth 482, in 
rich woods near Arlington, Texas, May 24, 1920 (Field, 2 sheets); 
J. H. Schuette 76, cult., Green Bay, Wisconsin, August 28, 1900 (Field) ; 
John K. Small, alt. 300-330 meters, on Little Stone Mountain, De 
Kalb County, Georgia, July 7, 1893 (Field, 2 sheets; Gray); H. E. 
Wheeler 62, near Hazen, Grand Prairie, Arkansas, May 22, 1924 
(Field) ; Charles Wright, Texas (Gray). 

Coreopsis grandiflora var. ft longipes (Hook.) Torr. & Gray, 
Fl. N. Amer. 2: 345. 1843. C. longipes Hook. Bot. Mag. pi. 3586. 
1837. 

Folia caulis basim versus dense vel moderate adgregata, plerum- 
que simplicia vel pinna ta; pedunculis crassioribus elongatis saepe 
2-3 dm. longis. 

Type specimen: Collected by Thomas Drummond in Texas and 
illustrated by Hooker with type plate (loc. cit.). 

Distribution: Texas. 

Specimens examined : Elihu Hall, Texas, 1877 (Field); idem 342, 
wet prairie, Houston, April 18, 1872 (Field) ; Earl E. Sherff 278, cult., 
St. Louis, Missouri, July 7, 1910 (Field). 

Coreopsis grandiflora var. 7 pilosa Sherff, Bot. Gaz. 89: 368. 
1930. 

A specie caulibus foliisque pubescentibus saepe etiam longe pa- 
tenteque piloso-hispidis differt. 

Type specimen: Collected by Steivardson Brown, Nathaniel L. 
Britton, and Peter Bisset, No. 2011, cultivated at Agricultural Station, 
Island of Bermuda, May 22-June 6, 1914 (N.Y.). 

Distribution: Known in a native state only from northern 
Georgia. 

Specimens examined : Brown, Britton, & Bisset 2011 (type, N.Y.); 
John H. Kellogg, cult., Missouri Bot. Card., May 23, 1911 (Mo.); 
John K. Small, alt. 210-300 meters, between Alcovy River and No 



REVISION OF THE GENUS COREOPSIS 355 

Business Creek, Oconee and Gwinnett Counties, Georgia, July 14, 
1893 (Field, 2 sheets; U.S.); idem, alt. 225 meters, on banks of 
Yellow River near McGuire's Mill, Gwinnett County, July 11, 1893 
(Mo.) ; idem, alt. 255 meters, on banks of Yellow River, near Yellow 
River (Store), Gwinnett County, July 20, 1893 (Field). 

The two cultivated specimens examined have a very unique aspect 
because of their marked hispidity. In spontaneous material the hairs 
are shorter and less spreading. 

Coreopsis grandiflora var. 5 Harveyana (A. Gray) Sherff, 
Bot. Gaz. 94: 593. 1933. C. Harveyana A. Gray, Syn. Fl. N. Amer. 1, 
pt. 2: 292. 1884. 

Foliorum segmenta anguste linearia vel quidem filiformia; 
achaeniis paulo minoribus, circ. 2 mm. longis. 

Type specimen: Collected by Francis LeRoy Harvey, No. 22, 
on cliffs near Fort Smith, Arkansas, June (Gray). 

Distribution: Missouri and Arkansas, perhaps also Kansas and 
Oklahoma. 

Specimens examined: Anon,, Arkansas National Forest, Arkan- 
sas, 1909 (U.S.); 5. F. Bush 158, Sheffield, Missouri, June 12, 1894 
(Berl.); Frederick V. Coville 127 Ark., Mountain Park near Little 
Rock, Arkansas, July 17, 1887 (U.S.); H. Eggert, Williamsonville, 
Missouri, June 11, 1893 (Field) ; F. L. Harvey 22 (type, Gray, cum 
C. tinctoria Nutt. pro parte parva commixta); E. J. Palmer 8120, 
wet, open hillsides, Malvern, Arkansas, June 23, 1915 (Mo.); H. S. 
Reynolds, Judsonia, Arkansas, May 31, 1877 (Field); Huron H. 
Smith 1150, railway tracks, Des Arc, Iron County, Missouri, June 8, 
1908 (Field). 

The type sheet of C. Harveyana Gray bears 3 specimens. The 
first at the left is C. tinctoria Nutt., having the oblong, wingless 
achenes, the bicolored ligules, the subtruncate style branch tips, the 
linear-oblong paleae, etc. of that species. The other two specimens 
are very distinct, having orbiculate, winged achenes, caudate-conical 
tips to the style branches, upwardly elongate-attenuate paleae, etc. 
Gray clearly was misled by the admixture of the small C. tinctoria 
specimen, for in his Synoptical Flora he separated C. Harveyana 
partly on the basis of the "rays sometimes brown-purple at base." 
Had the rays of C. Harveyana really possessed the tendency toward 
a bicolored state, a separation of the type as a new species would have 
been justified. Since the rays were really yellow, however, the type 
had nothing left to separate it from C. grandiflora proper except the 



356 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

narrower leaf divisions and the slightly smaller achenes. These 
differences Gray himself elsewhere (e.g. C. Drummondii Torr. & 
Gray, et var. Wrightii ipsi, Syn. Fl. N. Amer. 1, pt. 2: 291. 1884) 
considered as connoting a variety. (A supposedly duplicate sheet of 
the Harvey material at St. Louis [Mo.] bears all C. tinctoria Nutt.!) 
48. Coreopsis auriculata L. Sp. PL 908. 1753. Chrysanthemum 
hirsutum Virginian, auriculato Dulcamaraefolio octopetalon. Chry- 
santh. etc. Plukenet Aim. pi. 83, f. 5. 1691; Pluk. Phytogr. pi. 242, 
f. 4. 1769. Chrysanthemum trifoliatum minus Virgin, foliis obtusio- 
ribus, hirsutis Morison Hist. Univ. Oxon. 3, sect. 6, pi. 3, f. 45 (46). 
1715. Cymbaecarpa auriculata (L.) Hort. Reg. Matrit. ex Elench. PI. 
Hort. Bot. Destremx 1805: 19. 1806. Leachia trifoliata Cass. Diet. 
Sci. Nat. 25: 389. 1822 (ex synon. C. auriculata L.). Chrysomelea 
auriculata (L.) Tausch Hort. Canal. 1823. Coreopsis auriculata var. 
diversifolia Ell. Sketch Bot. S. Car. Ga. 2: 437. 1824. Chysomeka 
auriculata (L.) Tausch ex DC. Prodr. 5: 571. 1836 (sphalm). Core- 
opsis auriculata var. a Ell. ex DC. loc. cit. Coreopsis auriculata 
var. glabrata DC. loc. cit. Anacis auriculata (L.) Schrank, Denk- 
schr. Akad. Muench. Math. Nat. 5: 7. 1817. Coreopsis diversifolia 
DC. loc. cit. (nee alior.). Coreopsis auriculata var. /3 Torr. & Gray, 
Fl. N. Amer. 2: 343. 1843. Coreopsis oculata Hort. in Handlist 
Hardy Rock PI. Harvard Univ. Bot. Gard. 10. 1927 (sphalm). 

Herba perennis, erecta vel adscendens, simplex vel ramosa, 
pubescens vel hirsuta vel supra interdum glabrata, inferne foliosa, 
basi stolonifera. Folia opposita, petiolata petiolis nunc tenuibus 
nunc latis saepe elongatis itaque laminas plurimum excedentibus, 
petiolo adjecto plerumque 4-12 cm. longa; lamina membranacea, 
ovato-orbiculata vel elliptico-ovata, apice obtusa vel subacuta, 
lateribus integra vel 1-2-lobata lobis parvis basalibus. Capitula 
solitaria ramos elongates (saepius 1-2.5 dm. longos) nudos termi- 
nantia, radiata, pansa ad anthesin 3.5-5 cm. lata et circ. 9-10 mm. 
alta. Involucri bracteae subaequales, exteriores circ. 8, pallidae, 
ovato-oblongae vel lanceolato-oblongae, apice rotundatae vel sub- 
acutae, lateribus saepe tenuissime albido-marginatis glabratae vel 
hispido-ciliatae; interiores ovatae. Flores ligulati circ. 8, flavi, ligula 
obovati vel late cuneati, apice 3-lobati lobo mediano emarginato, 
1.5-2.3 cm. longi. Paleae anguste lineares, superne attenuatae, 
demum 6-9 mm. longae. Florum tubulosorum ramorum styli 
terminis caudato-cuspidati. Achaenia oblanceolata vel obovata, 
obcompressa, incurvata, nigra, glabra vel minute papillata, margi- 
nibus anguste alata alis demum cartilagineis involutisque, 2.2-2.9 



REVISION OF THE GENUS COREOPSIS 357 

mm. longa, apice primo minute bisquamellata demum plerumque 
calva. 

Type specimen: Linnaeus' first citation of literature is that of 
"Gron. virg. 105." This refers to a plant collected by John Clayton 
in Virginia and perhaps still extant in London (Brit.). Linnaeus cited 
also the two illustrations by Plukenet (loc. cit.) and the one by 
Morison (loc. cit.); all three are clear and agree unmistakably with 
each other. 

Distribution: Virginia and Kentucky southward to northern 
Florida, Mississippi, and Louisiana. Cited also for Illinois by 
Robinson and Fernald (Gray's New Man. 838. 1908). 

Specimens examined : Biltmore Herb. 176, rich woods, Biltmore, 
North Carolina, May 15, 1896 (Field ; Gray) ; idem 1 76b, eodem loco, 
May 20, 1897 (Field; Gray); ex herb. G. W. Bischoff (Gray); S. B. 
Buckley, Alabama, June, 1841 (Mo.); idem, Virginia (Gray); Wil- 
liam M. Canby 53, Stone Mountain, Georgia, May 3, 1899 (Gray) ; 
idem 54, Chattahoochie, Georgia, May 2, 1899 (Gray); A. W. Chap- 
man, mountains of Georgia (Field); Miss M. A. Coe, Atlanta, 
Georgia, May 6, 1903 (Gray); A. H. Curtiss, Bedford County, 
Virginia, June 1, 1871 (Gray); idem, eodem loco, May 31, 1872 
(Field); idem 1485, rich woods, Jackson County, Florida, June 
(Berl.; Field); F. S. Earle & C. F. Baker, Auburn, Alabama, April 4, 
1897 (Field) ; J. Hale, Alexandria, Louisiana (Gray) ; T. G. Harbison, 
Highlands, North Carolina, May, 1902 (Gray); R. A. Harper 74, 
Talladega County, Alabama, April 16, 1906 (Gray); ex herb. Dr. 
A. C. Hexamer & Dr. F. W. Maier, rich woods, Soluda River, Lex- 
ington District, South Carolina, June 14, 1855 (Gray); ex Hort. 
Berol. (Gray); M. E. Hyams, Statesville, North Carolina, July, 
1880 (Pom.); C. Mohr, open woods throughout the mountains, 
Lawrence County, Alabama, May, 1880 (Field); ex herb, eiusdem, 
Winston County, Alabama, May 1, 1881 (Field) ; P. H. Rolfs 392, 
Pensacola, Florida, April 11, 1894 (Field) ; Albert Ruth, open woods, 
near Knoxville, Tennessee, May, 1895 (Gray; Pom.); idem 67, rich 
woods, Knox County, Tennessee, June 15, 1900 (Gray); idem 652, 
copses, Knoxville, May, 1898 (Mo.) ; C. W. Short, Lexington, Ken- 
tucky, 1835 (Gray); J. K. Small & A. A. Heller, near R. and D. 
Railroad, between Fall Creek and Danville, Virginia, June 3, 1891 
(Field); ex Dr. Steetz, ex Lexington, Kentucky (Berl.); J. T. Stewart, 
northern Mississippi, 1863 (Field); K. A. Taylor, dry, sandy pine 
woods, Columbia, South Carolina, May, 1890 (Field) ; G. R. Vasey, 
North Carolina, 1878 (Field) ;E. H. Wharf, North Carolina (Gray). 



358 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

49. Coreopsis congregata Blake, Journ. Wash. Acad. Sci. 
19:275.1929. 

Herba annua, gracilis, erecta, subsimplex, sparsim pubescens, 
1-3.5 dm. alta, caule sulcato-striato, sparsim piloso praecipue 
superne pilis plerumque adscendentibus, internodiis 2-5 cm. longis. 
Folia opposita sessilia vel petiolata petiolis pilosissimis usque ad 1 
cm. longis, omnino circ. 2-4 cm. longa, pinnatifida vel inferiora 
bipinnatifida, circumambitu triangulata; segmentis ultimis mem- 
branaceis, oblongis vel ovatis vel lanceolato-ovatis, acutis vel obtusis, 
calloso-mucronulatis, utrinque pilosis, marginibus ciliatis, jugo infimo 
paene distincto. Capitula pauca (1-7), nunc solitaria mine corym- 
bosa, tenuissime pedunculata pedunculis sparsim pilosis erectis 
nudis vel minute 1-2-bracteatis 2-5 cm. longis, radiata, pansa ad 
anthesin 1-1.5 cm. lata et 0.5-0.7 cm. alta. Involucri pilosi bracteae 
exteriores circ. 8, tenues, anguste spathulatae vel lineares, obtuse 
calloso-apiculatae, 2.5-3.5 mm. longae et circ. 0.7 mm. latae, 3- 
nervatae nervo mediano multo magis perspicuo; interiores elliptico- 
oblongae, apice obtuso ciliolatae, aureae, 5-7-nervatae, 4.5-4.8 mm. 
longae et circ. 1.8 mm. latae. Flores ligulati 8, aurei, steriles, ligula 
elliptico-oblongi, apice subintegri, tantum circ. 5-nervatae, circ. 
6-7.5 mm. longi et 2.5-2.8 mm. lati, tubo glandulis stipitatis instructo. 
Paleae lineares, acutae vel obtusae, glabrae, flavae, circ. 4-vittatae, 
circ. 3.5 mm. longae. Flores tubulosi 12-20, corollis aureis tubo 
glandulis stipitatis instructis, stylorum ramis termino anguste 
caudato-appendiculatis. Achaenia exteriora obcompressa, obovata 
vel cuneato-rotundata, brunneo-atra, glabra, tergo levia vel superne 
plus minusve muricata, ventris mediane unicostati ac crustaceo- 
mamillati basi callosa, marginibus incrassatis subbrunneis circ. 
0.3 mm. latis longitudinaliter rugulosa infra sinuata supra in lobos 
paucos rotundatos divisa, apice parce depressa ac exaristata, 2-2.3 
mm. longa et 1.3-1.5 mm. lata; interiora angustiora. 

Type specimen: Collected by Ynes Mexia, No. 445, common at 
altitude of 1,200 meters, growing in masses, in damp places in 
openings in oak and pine forests, trail from El Batel to Pica de 
Aguila, Sierra Madre, State of Sinaloa, Mexico, November 14, 1925 
(Gray). 

Distribution: Known only from type locality in State of Sinaloa, 
Mexico. 

Specimens examined: Mexia 445 (type, Gray; cotype, Mo.). 

50. Coreopsis nuecensis Heller, Bot. Expl. S. Texas 106. 1895. 
Coreopsis coronata Hook. Bot. Mag. pi. 3460. 1836 (nee alior.). 



REVISION OF THE GENUS COREOPSIS 359 

Herba annua, sparsim hirsuta vel fere glabra, 2-6 dm. alta, 
caulibus erectis sed debilibus, basim versus ramosis, superne nudis. 
Folia opposita jugis saepe remotis, summa subsessilia, alia petiolata 
petiolis latis angustissimisve hispido-ciliatis (pilis pluriarticulatis) 
usque ad 5 (quidem ad 12) cm. longis, petiolo adjecto 4-10 (-20) 
cm. longa, lamina nunc late ovalia vel oblongo-spathulata integra, 
nunc pinnatim secta segmentis usque ad 5, ultimo multo maximo, 
lateralibus saepius alternis raro lobatis imis saepe petiolulatis. 
Capitula ramos nudos saepe 2-3 dm. longos (pedunculos) terminan- 
tia, radiata, pansa ad anthesin circ. 3.5-4.8 cm. lata et 1 cm. alta. 
Involucri bracteae exteriores circ. 8, oblongo-lanceolatae vel anguste 
deltoideo-ovatae vel subulatae, tergo glabratae, apice angusto 
saepius rotundatae, lateribus saepe diaphanae, marginibus ciliatae 
setis pluriloculatis, 5-8 mm. longae, quam interiores ovatae tergo 
glabrae vel albido-setosae apice subacutae paulo breviores. Flores 
ligulati circ. 8, ligula cuneato-ovati, basim versus maculis magnis 
atro-sanguineis vel brunneo-rubris ornati aliter flavi, apice 3-lobati 
lobo mediano multo majore plerumque in 2 lobulos hos saepe emar- 
ginatos rursus diviso, 1.5-2 cm. longi. Paleae lineares, supra demum 
angustissimae ac coloratae, 3-6 mm. longae. Flores tubulosi nunc 
flavidi nunc superne purpurascentes, stylorum ramis crasso-truncatis 
minute ac abrupte mucronatis. Achaenia obcompressa, circumam- 
bitu alis inclusis obovato-orbiculata circ. 4 mm. longa et circ. 4 mm. 
lata, dorso (convexo brunneo-nigro) spinulis saepe capitatis dense 
ventre (concavo utrinque plerumque perspicue calloso) sparsim 
munita, alata alis membranaceis brunneis irregulariter incisis vel 
lobatis circ. 1 mm. latis, apice 2 squamellis vel spinulis usque ad 0.4 
mm. longis munita. 

Type specimen : Raised from seed collected by Thomas Drummond 
in Texas, about 1835. 

Distribution: Southeastern Texas. 

Specimens examined: Berlandier 1802, from Bejar (Bexar) to 
Austin, Texas, April, 1828 (Gray). A. H. Curtiss 6362, waste ground, 
escaped from cultivation, St. Augustine, Florida, April 13-May 18, 
1898 (Gray; Mo.); Thomas Drummond 199, Texas (Gray); Elihu 
Hall, cult, e sem. texanis, Athens, Illinois, 1864 (U.S.); Hort. 
Cantabr., cult, anno 1848 e sem. texanis a Lindheimero lectis (Gray, 
2 sheets); idem, cult., September, 1856 (Mo.); Marcus E. Jones 
29511, Kingsville, Texas, March 29, 1932 (Pom.); idem 29512, 
Encino, Texas, March 28, 1932 (Pom.};F.Lindheimer, prairies west 
of the Brazos River, Texas, April (Mo.); idem 363, Guadeloupe 



360 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

bottoms at Victoria, Texas, February, 1845 (Mo.); C. Mohr, cult, 
e sem. texanis, Mobile, Alabama, May 10, 1887 (U.S.); H. Ness 
2078, moist forest lands, Brazos County, Texas, April, 1900 (Mo., 
2 sheets) ;#. J. Palmer 9144, sandy, open ground, Inez, Texas, March 
11, 1916 (Mo.); Earl E. Sherff 5017, cult, in Harvard Univ. Bot. 
Gard. (Hort. Cantabr.), August 12, 1929 (Field; Kew);5. C. Tharp 
5635, Woodboro, Texas, March 17, 1929 (U.S.); Tracy 8925 p.p., 
Victoria, Texas, April 27, 1905 (Mo.). 

Heller's name C. nuecensis goes back by direct synonymy to the 
name C. coronata Hook., which it supersedes because of the homonym 
C. coronata L. Heller's cited specimen (A. A. Heller 1548, Corpus 
Christi, Texas, April 11, 1894), however, was very different from 
C. coronata Hook, and was, indeed, at the time an undescribed species. 
It was later named C. similis Boynt. (qu. vide). 

51. Coreopsis cordylocarpa A. Gray, Proc. Amer. Acad. 22: 
428. 1887. 

Perennis, fruticosa, multicaulis, foliosa, 1.5-1.8 meters alta, 
brevissime adpresso-hispida, caulibus subtetragonis. Folia tenuiter 
petiolata petiolis 1-3.5 cm. longis, petiolo adjecto plerumque 0.7-2 
dm. longa, submembranacea, pinnatim 3-5-partita foliolis simplici- 
bus vel rursus 2-3-fidis, segmentis ultimis linearibus vel anguste 
oblongo-lanceolatis saepius 3-11 mm. latis, margine indurato-cras- 
siusculis et apicem acutum subacuminatumve versus acriter (inter- 
dum inciso-) dentatis. Capitula subcymosa pedunculata pedunculis 
tenuibus folia paulo vel multo superantia, radiata, pansa ad anthesin 
3-4 cm. lata et circ. 1 cm. alta. Involucri plus minusve hispidi 
bracteae subulatae, exteriores lineares 4-6 mm. longae, quam 
interiores lanceolatae saepe paulo longiores. Flores ligulati circ. 
10, lutei, ligula oblongi, apice integri vel denticulati, 1.5-2.2 cm. 
longi. Paleae oblongo-lineares, glabrae, apice acutae, demum 5-7 
mm. longae. Florum tubulosorum stylorum rami apice conico- 
angustata. Achaenia obcompresso-clavata, glabra, exalata, brun- 
neo-nigra, obscure longitudinaliterque striata, circ. 8-12 mm. 
longa et 1.5 mm. lata, apice contracto disco parvo calvo terminata. 

Type specimen: Collected by Edward Palmer, No. 172, at Rio 
Blanco, State of Jalisco, Mexico, July, 1886 (U.S.). 

Distribution : Known only from State of Jalisco, Mexico. 

Specimens examined: C. R. Barnes & W. J. G. Land 155, alt. 
1,560 meters, bank of stream, Sierra de San Esteban, State of Jalisco, 
September 28, 1908 (Field, 2 sheets); Palmer 172 (type, U.S.; 



REVISION OF THE GENUS COREOPSIS 361 

cotypes, Gray; N.Y.); C.G. Pringle 2367, river banks, near Guada- 
lajara, State of Jalisco, October 7, 1889 (Field; Gray; N.Y.); idem 
9885, alt. 1,500 meters, banks of Rio Blanco, near Guadalajara, July 
31, 1902 (Berl.; Field; Gray; N.Y.); idem 11506, eodem loco, October 
20, 1903 (Field; Gray). 

52. Coreopsis elgonensis Sherff, Bot. Gaz. 80: 374. 1925. 
Planta erecta, perennis, ramosa, 8-10 dm. alta; ramis angulatis, 

glabris, infra ligneis, internodiis numerosis et saepe tantum 3-10 mm. 
longis. Folia opposita, numerosa, sessilia, tantum circ. 1-1.8 cm. 
longa, ternatim divisa, foliolis membranaceis, margine ciliatis, 
faciebus atro-punctatis et interdum sparsissime hispidis, cuneatis, 
ternatim lobatis vel integris, segmentis ultimis acerrime apiculatis, 
plerumque 2-3 mm. latis. Capitula pauca in corymbis disposita 
pedunculis 2-9 cm. longis, radiata, pansa ad anthesin 2.5-3.5 cm. 
lata et 8-11 mm. alta. Involucri bracteae exteriores 6-11, lineares, 
supra latiores, apice acutae, plerumque glabratae, 5-9 mm. longae; 
interiores lanceolatae, hispidae, paulo breviores. Flores ligulati 
8-12, flavi, ligula anguste elliptico-oblanceolati, apice rotundati 
sed minute plus minusve denticulati, 8-12 mm. longi et 3-4.5 mm. 
lati. Paleae oblongo-lanceolatae, brunneae, pluristriatae, apice 
saepe longo-attenuatae, circ. 6-7.5 mm. longae. Disci florum 
stigmata caudato-attenuata. Achaenia late lineari-oblonga, plana, 
omnino atra, glabra, exaristata, 3.5-4.8 mm. longa et 1.1-1.3 mm. 
lata, marginibus non vere membranaceis. 

Type specimen: Collected by R. A. Dummer, No. 3304, locally 
frequent, at altitude of 3,900 meters, in thicket at edge of cliff, west 
side of crater, Mount Elgon, Uganda, British East Africa, January, 
1918 (Kew). 

Distribution: Known only from type locality. 

Specimens examined: Dummer 3304 (type, Kew). 

Strongly suggestive, in general habit, of such South American 
species as Coreopsis polyactis and C. senaria. The many small, 
sessile, cuneate, ternate or biternate leaves appear at first glance to 
be in whorls rather than in pairs. Correlated with the leaf abun- 
dance is the shortness of the internodes. Thus, for example, one 
branch of the type is seen to have 28 internodes in a length of 2.5 
dm., giving an average length of only about 9 mm. 

53. Coreopsis Chippii M. B. Moss, Kew Bull. 196. 1929. 
Suffrutex, 1 m. altus, caulibus (glabratis) et ramulis (puberulis) 

subteretibus, internodiis 5-12 mm. longis. Folia opposita, tenuiter 



362 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

petiolata petiolis circ. 4-6 mm. longis, petiolo adjecto circ. 1.5-2 cm. 
longa, 1-2-pinnatisecta, segmentis glabris vel glabriusculis, acriter 
apiculatis, crassiusculis, tantum circ. 0.2-0.4 mm. latis. Capitula 
subcorymbosa, tenuiter pedicellata pedicellis valde puberulis et 
circ. 1-2.5 cm. longis, radiata, pansa ad anthesin 2.3 cm. lata et 
5-6.5 mm. alta, demum subglobosa et circ. 1 cm. diametro. Invo- 
lucri hispidi bracteae exteriores 10-20, lineares, obtusae vel etiam 
subtruncatae, 4-5 mm. longae et circ. 0.5 mm. latae, saepe patentes; 
interiores late lanceolatae paulo longiores. Flores ligulati (pro- 
babiliter circ. 8), flavi, ligula anguste obovati, apice 3-denticulati, 
circ. 1 cm. longi. Paleae oblongae, obtusae, glabrae, nitidae, superne 
atriores, 4.5-5.5 mm. longae. Disci florum stigmata truncata, 
minute appendiculata. Achaenia valde obcompressa, dorso convexa, 
anguste obovato-oblonga, nigra, glaberrima, apice truncate exaris- 
tata, ad latera crassiusculo-marginata sed non vere alata, circ. 4 mm. 
longa et 1-1.4 mm. lata. 

Type specimen : Collected by Thomas Ford Chipp, No. 66, grow- 
ing in scrub on mountain top, also in forest in ravines, at altitude of 
3,125 meters, top of Mount Kinetti (Kineti), Imatong Mountains, 
southern Sudan, February 11, 1929 (Kew). 

Distribution: Known only from type locality in southern Sudan. 
("The Imatong Mountains, the highest point of which is Mt. Kineti, 
10,414 feet, are situated between 4 and 4 30' N. and 32 20' and 
33 E., just north of the boundary between the Anglo-Sudan and 
Uganda"; T. F. Chipp, Kew Bull. 177. 1929.) 

Specimens examined: Chipp 66 (type, Kew). 

54. Coreopsis scopulorum Sherff, Bot. Gaz. 88: 302. 1929. 

Frutex +2 dm. altus, glaber, ramosus, ramis foliosissimis, inter- 
nodiis saepe tantum 3-5 mm. longis. Folia opposita, petiolata 
petiolis 6-10 mm. longis, petiolo adjecto tantum 1.2-2 cm. longa, 
plus minusve biternatisecta, segmentis ultimis linearibus, subcarno- 
sis, margine saepe revolutis, apice mucronatis, vix 1 mm. latis. 
Capitula tenuiter pedunculata pedunculis 3-5 cm. longis ac saepe 
ad ramorum terminos 3-5-adgregatis, radiata (radiis in typo defi- 
cientibus sed ovariis circ. 8 sterilibus repraesentatis), demum circ. 
8 mm. alta et 7-10 mm. lata. Involucri bracteae exteriores circ. 
8, anguste lineares, patenti-reflexae, apice rotundatae vel abrupte 
cuspidatae, circ. 3 mm. longae, interioribus lanceolatis atque apicem 
versus reflexis subaequalibus. Paleae perspicuae, late oblongae, 
apice rotundato-obtusae, achaeniorum corpora moderate superantes. 



REVISION OF THE GENUS COREOPSIS 363 

Disci florum stigmata apice perspicue caudata. Achaenia linearia, 
obcompressa, exalata, faciebus marginibusque erecto-setosa, atra, 
singula facie circ. 8-striata, corpore 5-6 mm. longa et 0.7-1 mm. 
lata, apice erecto-setosa ac perspicue biaristata aristis tenuibus 
erecto-hispidis circ. 2.5 mm. longis. 

Type specimen: Collected by Edmund Heller, on cliffs at altitude 
of 2,130 meters, on summit of Mount Garguez, British East Africa, 
August 26, 1911 (U.S.). 

Distribution: Known only from type locality. 

Specimens examined: Heller, on cliffs at alt. 2,130 meters, summit 
of Mount Garguez, British East Africa, August 26, 1911 (type, U.S.). 

Apparently nearest Coreopsis elgonensis (from which it differs in 
its longer and petiolate leaves, its shorter external involucral bracts, its 
aristate achenes, etc.) and C. Chippii (from which it differs in its much 
broader leaf segments, etc.). All three species offer a strong resem- 
blance to certain low, shrubby South American species of Coreopsis. 

55. Coreopsis Jacksonii S. Moore, Journ. Linn. Soc. 35: 
347. 1902. Plate 1. Bidens spathulata Sherff, Bot. Gaz. 76: 149, 
pi. 13. 1923. B. Jacksonii (S. Moore) Sherff, Bot. Gaz. 81: 45. 1926. 

Folia plus minusve glabra C. Jacksonii. 

Folia pro parte maxima dense hispida var. /3 arthrochaeta. 

Herba procumbens vel adscendens, plus minusve glabra, forsitan 
annua, caule subsimplici nonnullis nodis radicante, 3.5-5 dm. 
longo. Folia non manifeste petiolata, 4-8 cm. longa et 1.5-2.5 cm. 
lata, valde membranacea, spathulato-oblanceolata, apice subobtusa, 
inferne sensim angustata, integra vel remotissime minimeque denti- 
culata, ciliata, in sicco plus minusve stramineo-viridia. Capitula 
perpauca (1-3), pedunculata pedunculis pubescentibus usque ad 
6 cm. longis, radiata, pansa ad anthesin circ. 3.5-4.5 cm. lata et 
7-9 mm. alta. Involucri bracteae exteriores circ. 6, membranaceae, 
oblongae vel oblanceolatae, ciliatae, apice orbiculatae, ad anthesin 
circ. 8-10 mm. longae et 2-3 mm. latae; interiores oblongo-ovatae, 
apice subacutae, ad anthesin circ. 3.5-4.5 mm. longae. Flores 
ligulati circ. 6, lutei, ligula elliptico-oblanceolati, apice dentati, 
1.5-1.8 cm. longi et 4.5-6.5 mm. lati. Florum tubulosorum stigmata 
termino anguste elongata. Achaenia subatra vel brunneo-atra, 
crasso-clavata, plus minusve tumida ac quadrangulata, non mani- 
feste striata, circ. 3.2-3.7 mm. longa et 1.1-1.7 mm. crassa, glabrata, 
exalata, exaristata sed apice saepe reliquo basali floris hermaphroditi 
coronata itaque false rostrata. 



364 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: Collected by Frederick J. Jackson, Kikuyu 
region, British East Africa, 1889 (Brit.). 

Distribution: Kikuyu region and northward to Mount Kenya 
(Kenia), British East Africa. 

Specimens examined: Frederick J. Jackson, Kikuyu region, 1889 
(type, Brit.); Edgar A. Mearns 1291, alt. about 3,630 meters, in 
"giant heath" zone, western slopes of Mount Kenya, British East 
Africa, September 21-27, 1909 (type of Bidens spathulata Sherff, 
U.S.); idem 1722, alt. about 3,000 meters, bamboo zone, western 
slopes of Mount Kenya, September 28-October 7, 1909 (U.S.). 

A species strikingly different from all other species of Coreopsis 
and easily distinguished from other African species by its thin, 
spatulate leaves. Its habit of rooting at many of the nodes, 
evidently in response to aqueous conditions of the habitat, is almost 
as distinctive, being common in only a few species of these two genera. 

My examination of the type specimen of Coreopsis Jacksonii S. 
Moore (Brit.) showed that it was merely a tiny, dwarfed specimen 
(cf. Moore, loc. cit., "unfortunately a mere scrap") of the species 
described and illustrated by me under the name of Bidens spathulata. 
(The type of Bidens spathulata, it will be noted, was collected upon 
Mount Kenya, British East Africa, while the type of Coreopsis Jack- 
sonii was collected in the small Kikuyu area immediately to the south- 
west.) Because, however, of its very diminutive stature (being only 
a few centimeters high), it lacked the spatulate type of leaves that 
so strongly characterizes larger plants. The leaves were more 
rotund in outline. For these reasons, Moore's description of Co- 
reopsis Jacksonii had been, of necessity, so misleading that only a 
personal study of his type plant could impart an understanding of 
its true specific status. As to its generic status, its complete lack 
of mature achenes, as also in the case of the type of Bidens spathulata, 
made it necessary to rely upon general habit, which is somewhat 
more that of Bidens than of Coreopsis. The recent finding (1) of the 
var. arthrochaeta of this species and (2) of Mearns 1 722 of the species 
proper makes it possible, however, to refer the species definitely to 
Coreopsis. 

Coreopsis Jacksonii var. $ arthrochaeta Sherff, Bot. Gaz. 
88: 302. 1929. 

A specie foliis pro parte maxima dense hispidis, setis albidis 
elongatis sensim attenuatis multiloculatis differt. 






FIG. 1. Coreopsis Jacksonii (S. L. Moore) Sherff. 

365 



366 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: Collected by R. L. Piemeisel and Leonard W. 
Kephart, No. 166, at altitude of 3,300 meters, vicinity of Camp 
Gusisu, Aberdares Mountains, British East Africa, July 29, 1927 
(U.S.). 

Distribution: Known only from type locality. 

Specimens examined: Piemeisel & Kephart 166 (type, U.S.). 

The type material of the var. arthrochaeta consisted of at least 
six small, entire plants and these offer a strikingly unique appearance 
because of their densely hispid leaves. Since endemism is pronounced 
among the plants of the Kikuyu region, and since elsewhere even 
where endemism is relatively unimportant (in the United States) 
Coreopsis displays many forms that are accepted as varieties, it 
appears worth while to distinguish the Piemeisel & Kephart plants as 
connoting a group of varietal rank. 

Explanation of Fig. 1. Coreopsis Jacksonii S. L. Moore: a, 
flowering specimen, X0.66; b, portion of leaf enlarged to show cilia- 
tion, X2.65; c, exterior involucral bract, X2.65; d, interior involucral 
bract, X2.65; e, ligulate floret, X2.65;/, palea, X2.65; g, disc floret, 
X2.65; all from Mearns 1291, type of Bidens spathulata Sherff, 
Herb. U. S. 

56. Coreopsis Negriana Sherff, Bot. Gaz. 90: 397. 1930. 

Herba annua, erecta, 5 dm. alta, caule gracili, glabro vel ad 
summam subhispido, viridi-stramineo. Folia opposita petiolata 
petiolis usque ad 12 mm. longis saepe tenuissimis petiolo adjecto 
5 cm. longa, 1-2-pinnatisecta, segmentis principalibus lineari- 
oblongis, membranaceis, supra infraque glabratis, marginibus ciliatis, 
acriter serratis dentibus interdum in setas elongatas desinentibus. 
Capitula graciliter pedunculata pedunculis moderate hispidis usque 
ad 14 cm. longis, radiata, pansa ad anthesin 2-2.5 cm. lata et circ. 
6-7 mm. alta. Involucri hispidi bracteae subaequales exteriores circ. 
8, lineari-spathulatae, acriter indurato-apiculatae, 3-4 mm. longae; 
interiores oblongo-ovatae, apicaliter saepe acutae. Flores ligulati 
circ. 8, flavi, ligula lineari-elliptici, apice obtuso obscure denticulati, 
circ. 1 cm. longi et circ. 2-3 mm. lati. Paleae nitido-hyalinae, 
lineari-oblongae vel lineari-lanceolatae, demum conduplicatae ac 
achaenium amplectentes, maturae circ. 6 mm. longae. Disci florum 
stigmata subito capilliformi-caudata. Achaenia lineari-oblonga, 
valde obcompressa, glabra, corpore nigro estriato vel obscurissime 
multistriato tantum 3-4 mm. longa et circ. 1 mm. lata, marginibus 
anguste alata alis brunneis glabrisque, apice exaristata ac glabra. 



REVISION OF THE GENUS COREOPSIS 367 

Type specimen: Collected by Giovanni Negri, No. 915 bis, at 
altitude of 1,500 meters, Arussi Galla, June 30, 1909 (Flor.). 

Distribution: Known only from type locality in Arussi Galla, 
Gallaland. 

Specimens examined: Negri 915 bis (type, Flor.). 

A species close to C. pachyloma 0. & H., from which it differs, 
however, in its smaller capitula, also its smaller achenes which are 
obcompressed, winged, and glabrous. The general habit is deceivr 
ingly like that oiBidens setigera (Coreopsis setigera Schz. Bip.), some 
of the foliar teeth even having the setiform extensions so notable in 
that species. But B. setigera's achenes are distinctly striate, up- 
wardly setose upon the faces and edges, entirely exalate, and (in 
the species proper) at the apex conspicuously slender-aristate. 

57. Coreopsis camporum Hutch. Kew Bull. 381. 1921. 

Herba perennis, erecta, caule 1-1.25 m. alto, basim versus 
dense foliato internodiis brevibus, glabro, simplici vel superne corym- 
boso-ramoso. Folia opposita, petiolata petiolis anguste marginatis 
circ. 1-2.5 cm. longis, petiolo adjecto 0.5-1.5 dm. longa, simpliciter 
vel rarius bipinnata, sicca laete viridia, segmentis linearibus vel 
lineari-lanceolatis, acutis, revolutis, unaquaque facierum glabris, 
margine spinuloso-setulosis et saepe sparsim dentatis (dentibus e 
basibus triangularibus et longe setigeris), usque ad 9.5 cm. longis 
et 7 mm. latis. Capitula pauca vel numerosa, subcorymbosa, 
breviter tenuiterque pedunculata pedunculis superne puberulis, 
radiata, pansa ad anthesin 2-3 cm. lata et 6-9 mm. alta. Involucri 
bracteae exteriores circ. 12, patulae, lineares, acutae, tergo glabratae 
sed marginibus setulosae, 0.8-1.5 cm. longae; interiores adscendentes, 
oblongo-lanceolatae, obtuse acuminatae, circ. 6 mm. longae et 2 
mm. latae, crebre resinoso-striatae. Flores ligulati circ. 10, auran- 
tiaci, fertiles, ligula oblongo-elliptici, apice breviter bilobati, circ. 
1.2-1.5 cm. longi et circ. 4 mm. lati, circ. 9-nervii. Paleae oblongo- 
oblanceolatae, minute mucronatae, circ. 5.5 mm. longae et circ. 
1.5 mm. latae. Achaenia immatura lineari-oblonga, complanata, 
omnino brunnea, alata, ciliato-pubescentia, corpore 4-5 mm. longa, 
biaristata aristis rigidis circ. 2.5 mm. longis. 

Type specimen : Collected by H. V. Lely, No. 383, at altitude of 
1,380 meters on plains between Hepham and Ropp, northern Nigeria, 
July 5, 1921 (Kew). 

Distribution: Northern Nigeria. 



368 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Specimens examined : Lely 383 (type, Kew); idem 449, alt. 1,380 
meters, Ropp, July 19, 1921 (Kew); J. Dent Young 153, alt. 900- 
1,350 meters, waste land, Vom, Bauchi Plateau, 1922 (Kew). 

58. Coreopsis Ellenbeckii 0. Hoffm. Bot. Jahrb. 38: 205. 1906. 
Herba perennis, glaberrima. Folia (caulium juvenilium) petiolata 

petiolis alato-marginatis basi connatis 5-10 mm. longis, petiolo 
adjecto 3-5 cm. longa, bipinnatipartita, rhachi late linearia ac alata 
segmentis linearibus acutis apicaliter callosis 1-2 mm. latis. Capi- 
tula (tantum unicum terminantem ramum juvenilem vidi) tenuiter 
pedunculata pedunculo glabro fere 3.5 cm. longo, radiata, pansa ad 
anthesin (bracteis patentibus inclusis) circ. 4.5 cm. lata et circ. 11 
mm. alta. Involucri bracteae exteriores circ. 5, perspicue foliaceae, 
trifidae, glaberrimae, apicaliter callosae, 1.7-2.4 cm. longae, lobo 
terminali multo majore 1.5 mm. lato et usque ad 1.5 cm. longo; 
interiores oblongae membranaceae brunneae margine angusto 
luteolo cinctae circ. 7-8 mm. longae. Flores ligulati 8, flavi, ligula 
elliptico-oblongi, multistriati, 1.8 cm. longi et 4 mm. lati, apice 
acriter 3-dentati dente mediano acriter bidenticulato. Disci florum 
stigmata terminaliter caudata. Achaenia matura non visa; ovariis 
obcompressis, margine hyalino cinctis faciebus glaberrimis sed 
marginibus dense erecto-ciliatis ciliis superioribus elongato-sericeis, 
biaristatis aristis corolla brevioribus rigide erecto-hispidis. 

Type specimen: Collected by Dr. Elknbeck (Exped. Baron von 
Erlanger), No. 529, at altitude of 2,500 meters in meadows on moun- 
tain slopes, on the Gara Mulata, Harar, State of Harar, eastern 
Abyssinia, March 21, 1900 (Berl.). 

Distribution: Known only from type locality at Harar, eastern 
Abyssinia. 

Specimens examined : Ellenbeck 529 (type, Berl.). 

Founded upon the lone type specimen, which consists of four 
leafy stems 13-20 cm. long, one of them ending in a flowering head. 
These come from the base of an old stem some 6 mm. thick 
which had died away. The juvenile or second-growth nature of the 
parts relied upon for the technical description leave some doubt 
as to the degree to which the characters recorded are truly 
representative. 

59. Coreopsis lineariloba 0. Hoffm. Bot. Jahrb. 30: 430. 1901. 
Herba perennis, ramosa, ramis glabris, erectis, internodiis saepius 

9-13 cm. longis. Folia opposita petiolata petiolis planis ac anguste 
marginatis usque ad 3 cm. longis, petiolo adjecto usque ad 12 cm. 



REVISION OF THE GENUS COREOPSIS 369 

longa, bipinnatipartita segmentis rhachi subsimilibus saepius 1-1.5 
cm. longis et plerumque 1.2-2.5 mm. latis linearibus apice acuto 
mucronatis glabris etiam eciliatis. Capitula mediocria tenuiter 
pedicellata pedicellis glabris cymam laxam bracteis linearibus 
munitam formantia, radiata, pansa ad anthesin circ. 2 cm. lata et 
7-9 mm. alta. Involucri bracteae exteriores 10-12, lineares, apice 
obtusae acutaeve, basi ciliatae ceterum glabrae, 4-6 mm. longae; 
interiores ovatae, acuminatae, apice tomentellae ceterum glabrae, 
saepe paulo breviores. Flores ligulati circ. 8-10, flavi, ligula ovati, 
apice subintegri, circ. 1 cm. longi. Paleae late oblongae, apice 
subacutae. Disci florum stigmata terminaliter caudato-attenuata. 
Achaenia matura ignota; ovariis obcompressis glabris vel saepius 
margine et inter aristas pappi ciliatis, pappi aristis 2 (raro 3) 
subulatis tubum corollae superantibus antrorsum longe pectinato- 
ciliatis. 

Type specimen: Collected by W. Goetze, No. 790, growing 1 
meter high on red laterite at altitude of 1,900 meters on an undulating 
plateau, Liangira, Ubena, southwesternmost German East Africa, 
March 24, 1899 (Berl.). 

Distribution: Known only from type locality in German East 
Africa. 

Specimens examined: Goetze 790 (type, Berl.). 

60. Coreopsis Schimperi 0. Hoffm. Bot. Jahrb. 38: 205. 1906. 
Microlecane abyssinica f. elongata Vatke, Linnaea 39: 497. 1875. 

Herba perennis, glaberrima, e radice fasciculata radiculis 1 dm. 
longis et circ. 2.5 mm. crassis, erecta vel basim versus procumbens 
et nodis radicans, caule gracili subsimplici 6-9 dm. alto. Folia 
opposita petiolata petiolis brevibus basaliter dilatatis connatisque 
tantum circ. 2-7 mm. longis, petiolo adjecto usque ad 5.5 cm. longa, 
quam internodia multo brevioria, circumambitu late deltoideo- 
ovata, 2-3-pinnatipartita segmentis linearibus acutis apice callosis 
margine raro spinuloso-ciliatis circ. 1 (raro 2) mm. latis. Capitula 
in corymbo amplo laxiusculo foliis reductis bracteato disposita 
pedicellis gracilibus spinuloso-hispidis vel glabratis apice paululo 
dilatatis usque ad 4.5 cm. longis, radiata, pansa ad anthesin 2-2.8 cm. 
lata et tantum circ. 5 mm. alta. Involucri bracteae exteriores circ. 
8 (raro usque ad 10), anguste lineares, subherbaceae, glaberrimae, 
subacriter calloso-apiculatae, 3-3.5 mm. longae et circ. 0.4 mm. 
latae; interiores late ellipticae, medio membranaceae purpurascentes 
pilos breves crassos gerentes, margine hyalina glaberrimae, apice 



370 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

in acumen breve obtusum productae, quam exteriores interdum 
paulo longiores. Flores ligulati circ. 8, flavi, ligula oblongo-elliptici, 
apice plerumque 3-lobulati, 1-1.5 cm. longi et 4-7 mm. lati. Paleae 
hyalinae, nitidae, late lineari-oblongae, apice subobtusae, lineis 3 
duplicibus percursae circ. 4-5 mm. longae. Disci florum stigmata 
minima terminaliter caudato-angustata. Achaenia leviter obcom- 
pressa, nitida, crasso-clavata, glaberrima, aegre striata, exalata, 
exaristata, circ. 2.5 mm. longa et fere 1 mm. lata. 

Type specimen: Collected by Wilhelm Schimper, No. 1386, in 
meadows at altitude of 2,460 meters, Gaffat, Abyssinia, October 15, 
1863 (Berl., 2 sheets). 

Distribution : Abyssinia. 

Specimens examined: Rohlfs & Stecker, Godofelassi, Abyssinia, 
January 7, 1881 (Berl.); Schimper 1386 (2 type sheets, Berl.; type 
and cotype of Microlecane abyssinica f. elongata Vatke). 

The type material had been incorrectly ranked by Vatke as a 
forma under Microlecane abyssinica (Schz. Bip.) B. & H. f. (Bidens 
setigera [Schz. Bip.] Sherff). As 0. Hoffmann (loc. cit.) has shown, 
the two are very different. 

61. Coreopsis Prestinariaeformis Vatke, Linnaea 39: 499. 
1875. C. Pristinariaeformis Vatke ex 0. & H. in Oliv. Fl. Trop. Afr. 
3: 387. 1877 (sphalm). C. heterocarpa Chiov., Ann. Bot. Roma 9: 
75. 1911 (pro maxima parte; cf. etiam var. incisa infra). 
Folia simplicia vel pinnatim 3-5-tripartita. 

C. Prestinariaeformis sensu stricto. 
Folia bipinnatisecta var. incisa. 

Herba annua, undique plus minusve pilis crassis flexuosis obsita, 
caule simplex vel quidem e basi valde ramosa, 1-6 dm. alta. Folia 
sessilia vel breviter petiolata, 2.5-7 cm. longa, circumambitu ovato- 
lanceolata vel deltoideo-cordata, sicca subvalde membranacea, 
simplicia vel plurimum subincise 3- vel etiam sub 5-partita, segmentis 
varie ovatis vel rhomboideo-ovatis et margine dentatis vel incisis 
lateralibus multo minoribus. Capitula saepius numerosa vel numero- 
sissima, tenuiter pedicellata pedicellis ante et per anthesin erectis 
postea hamato-recurvis et tune cernua, radiata, pansa ad anthesin 
circ. 2.8-3.3 cm. lata et circ. 1-1.2 cm. alta. Involucri bracteae 
aequilongae, exteriores 10-12, patulae, lineares vel lineari-spathulatae, 
apice obtusae vel acutiusculae, glabrae, circ. 4-6 mm. longae; 
interiores ovato-lanceolatae vel oblongo-ovatae, rufidulae, in dorso 
notatae ibique pilis longis crassis flexuosis flavescentibus onustae. 



REVISION OF THE GENUS COREOPSIS 371 

Flores ligulati 6 vel 7, aurei, ligula varie elliptico-obovati oblongo- 
elliptici vel late spathulato-oblanceolati, apice denticulati, superne 
in centre vel parum subtus macula aurantiaca rotunda elliptica vel 
lunulata praediti, 1.5-2.5 cm. longi et usque ad 8 mm. lati. Paleae 
glaberrimae, supra rufae. Achaenia valde obcompressa apice erecto- 
setosa in 3 vel 4 seriebus exteriora corpore 8 mm. longa et 
2 mm. lata, marginibus latissime alata alis apice truncatis et 
setoso-ciliatis; centralia lineari-oblonga exalata setoso-ciliata corpore 
12 mm. longa et 2.5-3 mm. lata; omnia cum pappo duobus 
aristis linearibus vel demum lanceolato-subulatis acutissimis achaenii 
corpori aequilongis vel quidem tantum 3-4 mm. longis antrorsum 
hispidis formato. 

Type specimen: Collected by Wilhelm Schimper, No. 1173, at 
altitude of 2,460 meters in meadows at Gaffat, Abyssinia, September 
25, 1863 (Berl., 2 sheets). 

Distribution: Abyssinia. 

Specimens examined: Emilio Chiovenda 970, Debarek, District 
of Semien, Province of Amhara, Abyssinia, July 11, 1909 (Flor.); 
idem 1148, Gondar, District of Dembia (Dembea), Province of Am- 
hara, Abyssinia, July 30, 1909 (Flor.); idem 1869, vicinity of Gondar, 
September 8, 1909 (Flor.); idem 1953 et 1954, District of Dembia, 
September 8, 1909 (Flor.); G. W. Grabham, Bahadur Gorges, on 
Lake Tana (L. Tsana), Abyssinia (Kew); Plowden, Abyssinia (Kew); 
Schimper 1173 (type, Berl., 2 sheets; cotype, Kew). 

An important plant in the festal celebrations of some of the 
Abyssinians (cf. Chiovenda, loc. cit.). 

Chiovenda (loc. cit.) cited seven of his own specimens as a basis 
for his Coreopsis heterocarpa: Nos. 1090, 1148, 1869, 1953, 1954, 1955, 
and 970 (in the order here given). His No. 1090 is my var. incisa 
(infra). The others, unless 1955 (which alone I have been unable 
to find at Florence), are C. Prestinariaeformis proper. Chiovenda's 
description as drawn fits primarily C. Prestinariaeformis. 

Coreopsis Prestinariaeformis var. ft incisa Sherff, Bot. Gaz. 
90: 387. 1930. C. heterocarpa Chiov. Ann. Bot. Roma 9: 75. 1911 
(pro minima parte). 

A specie foliis acriter et perspicue bipinnatisectis segmentis 
principalibus oblongo-lanceolatis differt. 

Type specimen: Collected by Dr.Ellenbeck (Expedition of Baron 
von Erlanger], No. 1436, growing 1 meter high at altitude of 2,600 
meters, Djafa, Arussi, Galla, Gallaland, July 21, 1900 (Berl.). 



372 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Distribution: Northern Abyssinia southward into Gallaland. 

Specimens examined: Emilio Chiovenda 1090, very common, 
Gondar, District of Dembia (Dembea), Province of Amhara, Abys- 
sinia, July 26, 1909 (Flor.); Ellenbeck 1436 (type, Berl.). 

62. Coreopsis Barter! 0. & H. in Oliver, Fl. Trop. Afr. 3: 390. 
1877. C. badia Sherff, Bot. Gaz. 76: 90. 1923. 

Herba glabra, adscendens, ramosa ramis teretibus striatis, 3-10 
dm. alta. Folia opposita vel rarius trina, sessilia vel subsessilia, 
membranacea, 1.5-5 cm. longa, nunc cuneato-oblanceolata et apicem 
versus subremote acriterque serrata, nunc irregulariter trilobata 
lobis lineari-oblongis. Capitula ramos singillatim terminantia, 
tenuiter pedunculata pedunculis 3-9 cm. longis, radiata, pansa ad 
anthesin 2-2.6 cm. lata et 7-9 mm. alta. Involucri glabri bracteae 
subaequales, exteriores 7-10, lineari-oblongae, acriter apiculatae, 
demum 5-7 mm. longae et 0.6-0.8 mm. latae; interiores oblongo- 
lanceolatae circ. 2.2-2.6 mm. latae. Flores ligulati circ. 8, lutei, 
ligula lineari-elliptici, apice obscure denticulati, 7-10 mm. longi et 
3-4 mm. lati. Paleae late oblongo-lineares, apice obtusissimae, circ. 
7 mm. longae. Disci florum stigmata parce incrassata, hispida, 
caudata. Achaenia obcompressa, atra, corpore tenuiter vel sublate 
linearia, 5-9 mm. longa et 0.9-1.5 mm. lata, faciebus venuste adpres- 
seque erecto-pilosa, marginibus (saltern achaenia exteriora) late et 
perspicue membranaceo-alata alis erecto-ciliatis, apice erecto-his- 
pido biaristata aristis tenuibus erecto-hispidis 2-4 mm. longis. 
Involucri bracteae interiores, paleae, achaeniorum alae badiae vel 
parce purpureo-badiae. 

Type specimen: Collected by Charles Barter, No. 870, chiefly in 
cultivated ground, Borgu, Upper Guinea (Kew). 

Distribution: Northwestern Africa, from Kamerun west to Togo 
and northward to Borgu. 

Specimens examined: W. B. Baikie, on the Niger River, West 
Tropical Africa (Kew) ; Barter, Borgu, Upper Guinea (type, Kew) ; 
Mission Gironcourt 256, on mountains of gneiss, Togo, 1908-1909 
(type of Coreopsis badia Sherff, Berl.); C. Ledermann 1855, alt. 
1,800 meters, swampy place on shore of stream, Babadju, Kamerun, 
December 15, 1908 (Berl., 2 sheets). 

63. Coreopsis monticola (Hook, f.) 0. & H. in Oliver, Fl. 
Trop. Afr. 3: 390. 1877. Verbesina (Prestinaria) monticola Hook, 
f. Journ. Linn. Soc. 7: 200. 1864. 



REVISION OF THE GENUS COREOPSIS 373 

Caulis demum plerumque glabratus. C. monticola. 

Caulis laxe vel subdense pilosus var. ft pilosa. 

Herba perennis, erecta, rigida, subsimplex, 3-11 dm. alta, caule 
vix subtetragono, superne circ. 2.5-3.5 mm. crasso, demum plerum- 
que glabrato. Folia opposita petiolata petiolis planis submar- 
ginatis 0.6-2 cm. longis, petiolo adjecto circ. 4-5 cm. longa, cir- 
cumambitu ovata, subcoriacea, margine et facie inferiore aegre 
hispida vel mox glabrata, trisecta, foliolis 3-5-sectis, segmentis 
ovatis vel lanceolatis vel anguste oblongis, infra pallidioribus, 
dentibus acriter apiculatis. Capitula corymbosa, pedunculata 
pedunculis tenuibus 2-8 cm. longis, radiata, pansa ad anthesin 
4-5.5 cm. lata et 8-10 mm. alta. Involucri glabri bracteae exteriores 
8 vel 9, lineares, acriter indurato-apiculatae, 8-9 mm. longae, quam 
interiores oblongo-ovatae nitidae rubro-brunneae vix longiores. 
Flores ligulati circ. 8 vel 9, aurantiaci, ligula lineares, circ. 13-15- 
nervii, apice minute sed acriter denticulati, circ. 1.8-2.2 cm. longi et 
4.5-6 mm. lati. Paleae lineari-oblongae, circ. 5-nerviae, apice 
obtusissimae, circ. 9 mm. longae et 2 mm. latae. Disci florum stig- 
mata lanceolato-caudata. Achaenia oblonga, obcompressa, nigra, 
exalata vel subanguste alata alis stramineis, facie ventrali margini- 
busque sursum setosa, corpore 3-5.2 mm. longa et alis inclusis 1.1-2 
mm. lata, apice corona setarum erectarum coronata ac biaristata 
aristis tenuibus stramineis sursum hispidis circ. 2.5-3 mm. longis. 

Type specimen: Collected by Gustav Mann, No. 1922, at altitude 
of 1,800-2,100 meters, Kamerun (Cameroon) Mountains, Kamerun, 
November 6, 1862 (Kew). 

Distribution: Kamerun. 

Specimens examined: W. Kalbreyer 125, alt. 2,100-2,550 meters, 
open plain, dry places, Kamerun Mountain, Kamerun (Berl.); 
Luckhardt 626, Kamerun, December, 1912 (Berl.); Mann 1922 
(type, Kew; cotypes, Berl.; Gray);/ 1 . Mildbraed 3392, "half-shrub" 
growing 7.5-10 dm. high, furrows between lava streams, lower Fuko 
Plateau, Great Kamerun Mountain, Kamerun, June, 1908 (Berl.); 
Mintz 40, alt. 2,400-2,800 meters, Great Kamerun Mountain, 
October 29, 1910 (Berl.) ; Preuss, alt. 2,000-2,800 meters, Kamerun, 
1891 (Berl.); idem 689, Buea, Kamerun, 1891 (Berl., 3 sheets; Gray); 
Gust. Simon 3, on lava fields, eodem loco, December 26, 1909 (Berl.). 






Coreopsis monticola var. ft pilosa Hutch. & Dalz. Fl. West 
Trop. Afr. 2:143. 1931. 



374 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Herba usque ad 3 m. alta, caulibus ramisque laxe vel subdense 
pilosis pilis multiloculatis, foliis membranaceis, petiolo adjecto 6-10 
cm. longis, superne hispidis; involucri bracteis exterioribus oblongo- 
linearibus vel lineari-lanceolatis 6-14 mm. longis et 1.5-3 mm. latis; 
interioribus tergo hispidis. 

Type specimen: Collected by Dr. Dunlap, No. 204, growing up to 
10 feet high in ravine above timber line at altitude of about 2,100 
meters, Kamerun Mountain, Kamerun, January 17, 1926 (Kew). 

Distribution: Kamerun. 

Specimens examined : Dunlap 204 (type, Kew) ; F. W. H. Migeod 
211, grassland, alt. 2,250 meters, Kamerun Mountains, November 
17, 1927 (Kew). 

64. Coreopsis oblonga Sherff, Bot. Gaz. 76: 80, pi. 7. 1923. 
Fig. 2. 

Herba erecta, verisimiliter perennis ac 6-12 dm. alta; caule 
obtuse tetragono, atro, apicem versus pubescenti, alibi glabro. 
Folia petiolata petiolis hispido-ciliatis vel brevibus (usque ad 1.5 
cm. longis) latisque vel longioribus (usque ad 3 cm.) et parce angustis, 
petiolo adjecto 6-7.5 cm. longa, rigida et subcarnosa, pinnata, foliolis 
acriter dentatis dentibus calloso-apiculatis, subprofunde incisis 
sinibus angustis vel foliolis lateralibus infimis etiam tripartitis, 
breviter hispido-ciliatis, setis acribus sparsissime vestitis, terminali 
oblongo-lanceolato, lateralibus oblongo-ovatis. Capitula pauca, 
breviter pedunculata pedunculis subrobustis ac hie illic foliolo- 
bracteatis usque ad 6 cm. longis, radiata, pansa ad anthesin 2.5-3 
cm. lata et 9-11 cm. alta. Involucrum setis validis acribusque his- 
pidum; bracteis exterioribus 6-9, late lineari-spathulatis, apicem 
versus glabratis, apice subobtusis, 8-11 mm. longis, eis capitulorum 
juniorum discum manifeste superantibus; interioribus oblongo- 
oblanceolatis vel oblongo-obovatis, apicem versus constricta, exteri- 
oribus demum subaequalibus. Flores ligulati 7, aurantiaci, ligula 
elliptico-lanceolati, apice late obtusi et integri denticulative, circ. 
15 striis percursi, tubo adjecto 1.5-2.4 cm. longi, tubo circ. 5 mm. 
longo. Paleae lineari-oblongae. Flores tubulosi limbo aurantiaci, 
infra pilosi (ad medium etiam longe hirsuti), supra perpaucis pilis 
interdum conspersi; stylorum ramis terminaliter elongato-attenuatis. 
Achaenia matura ignota. Ovaria subnigra, oblongo-cuneata, alata, 
plana, faciebus glabra, corpore 2-3 mm. longa et 0.7-1 mm. lata, 
apice arrecte setosa et longe biaristata; aristis acribus, supra et ad 
medium glabris, basim versus setulis arrectis 1-3 sparsim munitis, 
2-2.5 mm. longis. 




FIG. 2. Coreopsis oblonga Sherff. 

375 



376 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: Collected by Alex. Carson, No. 106, Lake Tan- 
ganyika, East Equatorial Africa, September, 1893 (Kew). 

Distribution: Known only from type locality in East Equatorial 
Africa. 

Specimens examined: Carson 106 (type, Kew). 

A species approaching Coreopsis pinnatipartita 0. Hoffm., Bidens 
Grantii (Oliv.) Sherff, and B. Holstii (0. Hoffm.) Sherff. The hairy, 
aristate ovaries distinguish it easily from C. pinnatipartita, while 
the definite wings of the ovaries distinguish it from B. Grantii and 
B. Holstii. 

Explanation of Fig. 2. Coreopsis oblonga Sherff: a, main portion 
of flowering specimen, X0.72; 6, lower leaf from a, X0.72; c, exterior 
involucral bract, X4.33; d, interior involucral bract, X4.33; e, 
ligulate floret, X2.16; /, palea, X4.33; g, disc floret with achene 
immature, X4.33; all from type, Herb. Kew. 

65. Coreopsis pinnatipartita 0. Hoffm. Bot. Jahrb. 30: 432. 
1901. 

Fruticosa, perennis, erecta, 1-3 m. alta, ramis erectis, tomentel- 
lis, junioribus foliatis. Folia opposita petiolata petiolis anguste 
marginatis usque ad 3 cm. longis, petiolo adjecto 9-16 cm. longa, 
pinnatipartita, circumambitu ovata, segmentis ovatis vel lanceolatis, 
acutis vel obtusis, irregulariter pinnatilobis vel plus minusve pro- 
funde serratis, coriaceis, supra scaberulis subtus tomentellis. Capi- 
tula in corymbum oligocephalum terminalem congesta, breviter 
subvaldeque pedicellata pedicellis tomentellis, radiata, pansa ad 
anthesin 5-6.5 cm. lata. Involucri bracteae aequilongae, ovatae, 
obtusae, dorso tomentosae, mox reflexae, 8-15 mm. longae, exteriores 
circ. 8-10, plus minusve biseriales. Flores ligulati circ. 10-12, 
flavi, ligula elliptici vel elliptico-oblanceolati, apice plerumque 
acriter denticulati, circ. 3 cm. longi et circ. 6-9 mm. lati. Paleae 
perspicue 3-pluri-lineatae lineis brunneis vel brunneo-purpurascenti- 
bus, apice obtusae, achaenia multo superantes. Disci florum 
stigmata terminaliter attenuato-elongatae. Achaenia lineari-ob- 
longa, valde obcompressa, glaberrima, nigra vel subbrunneo-nigra, 
faciebus multistriata, alata alis margine exteriore stramineis ac 
crassiusculis, 5-6.5 mm. longa et 1.5-1.8 mm. lata, apice exaristato 
calloso-indurata. 

Type specimen: Collected by W. Goetze, No. 1041, at altitude of 
2,300 meters on slopes of Poroto Mountain, southwesternmost 
German East Africa, June 17, 1899 (Berl., 2 sheets). 



REVISION OF THE GENUS COREOPSIS 377 

Distribution: Nyassaland to German East Africa. 

Specimens examined : J. Buchanan 380, Nyassaland, 1891 (Berl.); 
A. Engler 1266, growing 1 meter high at alt. 2,000-2,400 meters, in 
high mountain shrub and eagle-fern formation, "Magamba oberhalb 
Kwai," West Usambara, German East Africa, October 4, 1902 
(Berl.) ; Goetze 1041 (2 type sheets, Berl.); Ad. Stolz 1306, growing 3 
meters high at alt. 1,450 meters, Kyimbila, Nyassaland, May 23, 1912 
(Berl., 2 sheets); A. Whyte, alt. 1,200-1,800 meters, Mount Zomba, 
Nyassaland, December, 1896 (Berl.). 

66. Coreopsis Whytei S. L. Moore, Journ. Linn. Soc. 35: 348. 
1902. 

Herba verisimiliter perennis elataque, caule erecto subtereti vel 
obtuse angulato pubescenti mox glabrato, ad nodos tumido. Folia 
opposita, principalia petiolata petiolis saepius 2-3 cm. longis petiolo 
adjecto 8-10 cm. longa, pinnatifida segmentis ovatis acutis den- 
tato-lobulatis; summa sessilia lanceolata dentato-lobulata integrave; 
omnia supra sparsissime minutissimeque hispida, subtus pubescentia. 
Capitula pedunculata pedunculis pubescentibus circ. 2-4 cm. longis, 
sine dubio radiata, post anthesin fere vel vere 3 cm. lata et 1.3 
cm. alta. Involucri 3-seriati bracteae extimae et intermediae 
anguste obovatae obtusae adpresso-erecte hispidae pilis plurilocu- 
latis 1.2 cm. longae; intimae oblongae obtusae tenuiores et paulo 
breviores. Flores ligulati non observati. Paleae oblongo-lanceolatae, 
obtusissimae, fere 1 cm. longae. Achaenia valde obcompressa, sub- 
cuneate lineari-oblonga, angustissime alata, brunneo-nigra, glabra, 
faciebus striata, apice incrassato exaristata, circ. 6-7 mm. longa et 
circ. 2 mm. lata. 

Type specimen: Collected by Alexander Whyte, No. 35, on Mount 
Milanji (Mount Mlange), southernmost Nyassaland, October, 1891 
(Brit.). 

Distribution: Known only from type locality in southernmost 
Nyassaland. 

Specimens examined: Whyte 35 (type, Brit.). 

67. Coreopsis Lupulina 0. Hoffm. Bot. Jahrb. 30: 432. 1901. 

Fruticosa, glaberrima, perennis, ramosa, 2-3 m. alta, ramis 
angulatis terminaliter foliatis. Folia opposita tenuiter petiolata 
petiolis basaliter connatis usque ad 4 cm. longis, petiolo adjecto 
8-12 cm. longa, principalia bipinnatipartita segmentis oblongis ser- 
ratis vel pinnatifidis apice acutis, superiora simpliciter pinnatipartita. 



378 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Capitula saepius in corymbos densos oligocephalos congesta pedicel- 
lis brevibus crassisque, radiata, pansa ad anthesin circ. 6 cm. lata et 
1.5-1.8 cm. alta. Involucri bracteae demum valde reflexae, exteri- 
ores circ. 7-9 crassiusculae oblongo-ovatae apice obtusae circ. 8 
mm. longae et 4-6 mm. latae, interiores membranaceae ovatae vel 
lanceolato-ovatae obtuse acuminatae apice minute fimbriatae circ. 
12 mm. longae. Flores ligulati circ. 10, flavi, ligula elliptici, apice 
circ. 3-denticulati, circ. 2.5 cm. longi et circ. 7-8 mm. lati. Paleae 
perspicue brunneo-striatae, lineares, obtusae, corollas superantes, 
exteriores apicem versus demum reflexae. Disci florum stigmata 
apicaliter elongato-attenuata. Achaenia valde obcompressa, lineari- 
oblanceolata, omnino atra, marginaliter alata alis quam corpore 
paulo vel etiam dimidio angustioribus, glabra, faciebus multi- 
striata 5.5-8 mm. longa et (alis inclusis) 1.7-2.5 mm. lata, apice 
calloso-marginata et obtuse emarginata. 

Type specimen: Collected by W. Goetze, No. 1069, at altitude of 
2,400 meters on slopes of Beya Mountain, Usafua, south westernmost 
German East Africa, June 27, 1899 (Berl., 2 sheets). 

Distribution: Known only from type locality in German East 
Africa. 

Specimens examined : Goetze 1069 (2 type sheets, Berl.). 

This species and C. pinnatipartita are distinguished from the 
very similar Bidens Holstii (0. Hoffm.) Sherff by their winged 
achenes. 

68. Coreopsis Neumannii Sherff, Bot. Gaz. 76: 85. 1923. 

Herba 1.5 m. alta, caule maximam partem glabro, sulcato, 
ramoso, internodiis folia facile superantibus. Folia subsessilia vel 
petiolata petiolis usque ad 1 cm. longis, petiolo adjecto 2-4 cm. longa, 
plerumque tripartita (inferiora non visa, pinnata ?) foliolis ovatis vel 
interdum lanceolatis, dentatis vel lobulatis, pubescentibus, dentibus 
acerrime apiculatis. Capitula corymbosa, 5-11 in unaquaque corym- 
borum, radiata, pansa ad anthesin 2.2-2.7 cm. lata et circ. 6 mm. 
alta, pedunculis tenuibus plerumque 1-3 cm. longis. Involucrum 
inconspicue hispidum, bracteis exterioribus circ. 8, flavido-viridibus, 
lineari-spathulatis, apice acutis, 3-7 mm. longis, quam interioribus 
lanceolatis paulo brevioribus. Flores ligulati plerumque 8, flavi, 
ligula elliptico-oblongi, paucistriati, apice integri vel denticulati, 
circ. 1.2 cm. longi. Paleae oblongo-lanceolatae, saepius acutae, 
nitidae, achaeniorum aristis superatae. Disci florum stigmata 
caudato-elongata. Achaenia plumbeo-nigra, circumambitu lineari- 



REVISION OF THE GENUS COREOPSIS 379 

oblonga vel oblanceolata, valde obcompressa, alata alis coloris 
similis, glabra, corpore 3-4 mm. longa et (alis inclusis) 1-1.5 mm. 
lata, biaristata aristis brevissime sursumque setosis, circ. 0.5-1 mm. 
longis. 

Type specimen: Collected by Oscar Neumann (Exped. to Sidamo, 
Utadera, and Lake Abbaja), No. 134, at altitude of 2,800 meters, 
mountain meadows, Gardulla (incorrectly printed Gandulla at 
time of my original description in 1923), Gallaland (southern Abys- 
sinia), January 14, 1901 (Berl.). 

Distribution: Known only from type locality in Abyssinia. 

Specimens examined: Neumann IS 4 (type, Berl.). 

The single type specimen has its heads in corymbose clusters, and 
thus simulates in habit Bidens squarrosa HBK. of tropical America. 

69. Coreopsis Feruloides Sherff, Bot. Gaz. 80: 375. 1925. 

Herba erecta, ramosa, forsitan annua, circ. 7-10 dm. alta, 
glabra, viridis vel glaucescens, caule subtetragono. Folia sessilia vel 
subsessilia, 4-8 cm. longa, bipinnata, costa mediana tenui, segmentis 
linearibus vel anguste lanceolatis, crassiusculis, minute ciliatis, 
obtusis vel subacutis, plerumque 2-4 mm. latis. Capitula sub- 
numerosa, corymbosa, radiata, pansa ad anthesin 2.5-3 cm. lata et 
5-7 mm. alta. Involucri subglabrati bracteae exteriores circ. 8, 
lineares, apice subobtusae indurataeque, 3-5 mm. longae, interiores 
lanceolatae circ. 6-7 mm. longae. Flores ligulati circ. 8, flavi, ligula 
lineari-oblongi, apice obtuso saepius emarginati et aegre circ. 3- 
denticulati, 1.5 cm. longi et circ. 4-5 mm. lati. Paleae lineari- 
oblongae, longitudinaliter 1-3-nerviae, apice subabrupte acutae, 
circ. 5.5-7 mm. longae. Disci florum stigmata caudato-attenuata. 
Achaenia matura minuta, non exserta, atra, valde obcompressa, 
lineari-oblonga, leviter alata, faciebus subobscure striata, faciebus 
marginibusque minute erecto-setosa, corpore tantum 3.5-4 mm. 
longa et (alis inclusis) parce 1 mm. lata, apice biaristata; aristis 
tenuissimis, stramineis, setulis minutis erectis numerosis instructis, 
1-1.6 mm. longis. 

Type specimen: Collected by E. Battiscombe, No. 945, at altitude 
of 2,250-2,400 meters, Kinabop Plateau, Western Aberdare Moun- 
tains, British East Africa (Kew). 

Distribution: Known only from type locality. 

Specimens examined : Battiscombe 945 (type, Kew). 

The general habit simulates very closely that of the Mexican 
Bidens serrulata (Poir.) Desf. (B. grandiflora Balb.) and B. Ferulaefolia 



380 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

(Jacq.) DC. The leaves suggest the primary lower lateral portion 
of the more decompound leaves of Ferula glauca L. Aside from the 
habit, the tiny achenes are very distinctive, very few African species 
of Coreopsis having such small ones. The achenial wings are of the 
same color as the body proper, thus easily escaping notice. The 
species is somewhat unwelcome, as it serves further to weaken the 
generic distinctions between Coreopsis and Bidens. 

70. Coreopsis ochracea 0. Hoffm. Bot. Jahrb. 30: 431. 1901; 
Sherff, Bot. Gaz. 80: 375 and pi 19. 1925. C. cosmophylla Sherff, 
op. cit. 76: 90 and pi. 9, ff. h-n. 1923. Bidens ochracea (0. Hoffm.) 
Sherff, op. cit. 158. Fig. 3. 

Herba perennis, e radice crassa caules interdum complures emit- 
tens, elatos, parce ramosos, glaberrimos, infra obtuse supra acrius 
quadrangulatos, 0.8-2 m. altos. Folia pallida petiolata petiolis 
marginatis usque ad 4.5 cm. longis, petiolo adjecto 4-10 vel saepe 
etiam usque ad 18 cm. longa, pinnatim 3-5-partita, segmentis sub- 
crassis, integerrimis vel iterum 3- (rarius 2- vel 4-) partitis, ultimis 
linearibus, plerumque 1.5-5 mm. latis, apice acutis et mucronatis, 
supra glaberrimis vel breviter scabro-pilosis, subtus glabris vel saepe 
ad venam medianam perspicuam sparsim setosis, margine paulo 
revolutis et breviter scabro-ciliolatis. Capitula ramos terminantia, 
longe pedunculata pedunculis ebracteatis vel foliis reductis bracteatis 
et usque ad 3 dm. longis, radiata, pansa ad anthesin 4-8.5 cm. lata 
et 1-1.5 cm. alta. Involucri bracteae exteriores 8-10, basi saepe 
pilosae, margine sparsim spinuloso-ciliatae, ceterum plerumque 
glabrae, lineari-ellipticae vel late lanceolatae, supra basim paulo 
constrictae, latiores saepe ad apicem in acumen breve obtusum 
subito angustatae, 8-13 mm. longae; interiores lanceolatae vel 
ovato-lanceolatae paulo longiores, glabrae vel apice minute pubes- 
centes, paleas aequantes vel paulo superantes. Flores ligulati 
plerumque 8, aurantiaci, ligula lineari-elliptici, saepe minute pul- 
verulento-tomentosi, apice denticulati, 2-3.5 cm. longi. Paleae 
oblongo-lanceolatae, acutae. Disci florum stigmata terminaliter 
acuto-subcaudata. Achaenia plana, atra, alata, marginibus erecto- 
setosa, faciebus glabra, corpore 6-7 mm. longa et 2.5-3.5 mm. lata, 
apice setosa et biaristata aristis tenuibus, calvis vel sparsissime 
erecto-setosis, circ. 2 mm. longis. 

Type specimen: Collected by W. Goetze, No. 731, at altitude of 
about 1,700 meters, in thin bush-growth on red laterite on hilly 
plateaus, Bweni, Uhehe, German East Africa, March 11, 1899 
(Berl.). 




FIG. 3. Coreopsis ochracea O. Hoffm. 

381 



382 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Distribution: Northern Nyassaland northward through German 
East Africa into British East Africa. 

Specimens examined :E. Battiscombe 83, alt. 1,050-1,200 meters, 
Muhoroni, British East Africa (Kew; forma foliis plus minusve 
atypica); Goetze 731 (type, Berl.); Muenzner 159, Msamvia, Lake 
Tanganyika District, German East Africa, February 24, 1909 
(Berl.; type of Coreopsis cosmophylla Sherff); W. H. Nutt, alt. 
about 1,800 meters, between Lake Tanganyika and Lake Rukwa 
(L. Rikwa, L. Leopold), 1896 (Kew); Ad. Stolz 764, growing 2 meters 
tall at alt. 600 meters, Kyimbila, Langenburg, German East Africa, 
June 8, 1911 (Berl.); idem (similiter) 764, alt. 600 meters, eodem 
loco, February 27, 1912 (Del.) ; A. Whyte, northern Nyassaland (Kew). 

Hoffmann had not seen ripe fruits of this species ("Reife Friichte 
fehlen; nach den Fruchtknoten zu erteilen konnten sie gefliigelt sein"; 
Hoffm., loc. cit.). In my own earlier studies it seemed that the 
specimens of this handsome species of the Nyassa region were of two 
kinds, differing generically on the basis of the winged character of the 
achenes. Further specimens have shown, however, that some materi- 
al is merely more tardy than the rest in displaying achenial wings. 
As the specimens mature, their achenes become fairly well winged. 

Explanation of Fig. 3. Coreopsis ochracea 0. Hoffm.: a, 6, 
flowering branch and portion of stem with foliage, X0.58; c, d, 
exterior involucral bracts, X2.33; e, interior involucral bract, X2.33; 
/, ligulate floret, Xl.75; g, palea, X2.33; h, disc corolla, X2.33; i, 
immature achene, X3.5; all from type, Herb. Berl. 

71. Coreopsis leptoglossa Sherff, Bot. Gaz. 76: 88. 1923. 

Herba annua, erecta, gracillima, speciminibus observatis parce 
ramosis, caule tetragono, glabro, 8-10 dm. alto. Folia petiolata, 
petiolis latis hispido-ciliatisque usque ad 7 mm. longis vel angustatis 
glabratisque usque ad 2.5 cm. longis, petiolo adjecto 5-9 cm. longa, 
glabra, bipinnata, segmentis linearibus, plerumque circ. 1 mm. 
rarius 1.5-2 mm. latis, apice acutis, margine non ciliatis sed parce 
revolutis. Capitula tenuiter pedunculata pedunculis 2-10 cm. 
longis, radiata, pansa ad anthesin 4-5 cm. lata et 6-9 mm. alta. 
Involucri bracteae basi dense aliter leviter hispidae setis acribus, 
arcuatis, multi(6-12)-loculatis, exteriores 8-10, lineares, apice 
acutae, margine maximam partem non ciliatae, 5-7 mm. longae et 
0.6-0.9 mm. latae, interioribus lanceolatis plerumque breviores. 
Flores ligulati circ. 8, lutei, leptoglossati, ligula lineari-elliptici, apice 
integri vel obscure denticulati, 2-2.5 cm. longi et 4-6 mm. lati. 



REVISION OF THE GENUS COREOPSIS 383 

Paleae ignotae. Disci florum stigmata angustato-caudata. Achaenia 
matura ignota; ovariis obovatis, valde obcompressis, late alatis, 
supra ad margines et medium erecto-hispidis, biaristatis; aristis 
erecto-hispidis, 0.4-0.7 mm. longis. 

Type specimen: Collected by T. Kassner, No. 2871, on mountain 
slope, Lofuku River, Belgian Congo, May 25, 1908 (Berl.). 

Distribution: Known only from type locality in Belgian Congo. 

Specimens examined : Kassner 2871 (type, Berl.). 

Has flowering heads resembling those of Bidens Steppia (Steetz) 
Sherff, especially as to involucre. 

72. Coreopsis multiflora Sherff, Bull. Jard. Bot. Brux. 13: 
292. 1935. 

Herba erecta, forsitan perennis, probabiliter 1 m. alta, caule 
vel ramis glabratis atque acriter tetragonis. Folia principalia petio- 
lata petiolis marginatis 1.5 cm. longis, petiolo adjecto 1.2 dm. 
longa, pinnatim circ. 7-partita, foliolis lateralibus oppositis (jugis 
remotis), inferioribus circ. 5 cm. longis; omnibus membranaceis, 
circumambitu lanceolatis, cum rhachi confluentes, incise lobulatis 
(lobulis plus minusve dentatis dentibus acutis) utrinque subsparsim 
setosis (setis tenuibus) marginibus ciliatis. Capitula pauca ad 
ramos, pedunculata pedunculis subrobustis inferne glabratis superne 
setis albidis crispis patentibus hispidis 2-6 cm. longis, radiata, ad 
anthesin cernua et pansa 5-6 cm. lata et 1.3 cm. alta. Involucrum 
basi dense hispidum setis albis crispis acribus pluriloculatis, post 
anthesin circ. 2-2.3 cm. latum, bracteis subaequalibus; exterioribus 
linearibus, acutis, crassiusculis, extus glabratis vel sparsim hispidis, 
intus inferne glabris superne densissime hispidis, 8-11 mm. longis; 
interioribus ovato-oblongis, extus subsparsim hispidis. Flores 
ligulati 8-10, aurantiaci, ligula lineares vel lineari-oblanceolati, 
apice minutissime 2-3-denticulati, circ. 2.5 cm. longi et 4-5 mm. lati. 
Flores tubulosi numerosi (125), corollis extrinsecus glabris circ. 5 
mm. longis, stylorum ramis longe tenuiterque appendiculatis. 
Paleae oblongae, apicem ciliatum obtusumque versus atriores, ad 
anthesin saepius aristas aequantes. Achaenia non visa. Ovaria 
plana, ovata, alata, infra medium obsolete supra medium manifeste 
ciliata setis saepe erectis, faciebus glabrata, corpore circ. 3 mm. longa 
et circ. 1.5 mm. lata, apice biaristata aristis tenuibus antrorsum his- 
pidulis 1-1.5 mm. longis. 

Type specimen: Collected by Dom Benoit Thoreau, No. 2. 11, Pla- 
teau of Bianos, Kansenia, Katanga, Belgian Congo, June, 1927 (Bruss.). 



384 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Distribution: Katanga, Belgian Congo. 
Specimens examined: Thoreau 2. 11 (type, Bruss.). 

73. Coreopsis bella Hutch. Kew Bull. 364. 1907. 

Herba erecta, verisimiliter perennis, suffruticosa, 1 m. alta, 
ramis parce pubescentibus. Folia petiolata petiolis 2-3 cm. longis, 
petiolo adjecto 7-15 cm. longa, tenere coriacea, circumambitu trian- 
gulata vel ovato-triangulata, pinnatipartita, supra breviter subtus 
longe pubescentia segmentis ovato-lanceolatis acutis plus minusve 
profunde serratis vel irregulariter pinnatilobis. Capitula tenuiter 
pedunculata pedunculis pubescentibus 2-6 cm. longis, radiata, 
pansa ad anthesin circ. 5.5-6.5 cm. lata et circ. 1.4-1.7 cm. alta. 
Involucri bracteae obtusae, circ. 1.3 cm. longae et circ. 4 mm. latae, 
exteriores oblongae, pilosae; interiores extrinsecus pubescentes. 
Flores ligulati circ. 12, flavi, ligula oblongo-elliptici, circ. 3 cm. longi, 
apice subacuti vel obscure dentati, tubo brevi extrinsecus pilosi. 
Paleae oblongae, glabrae, apice obtusae, apicem versus leviter con- 
strictae, circ. 1 cm. longae et circ. 1.5 mm. latae. Achaenia immatura 
plana, pilosa, corpore circ. 1 cm. longa, apice biaristata aristis 1.5 
cm. longis. 

Type specimen: Raised at Kew in 1907 from seeds collected in 
British East Africa by R. Diespecker (Kew). 

Distribution: British East Africa. 

Specimens examined : Cult, at Kew from seeds collected by Dies- 
pecker (type, Kew). 

Close in general appearance to some African species of Bidens. 
Mature achenes are much to be desired. 

74. Coreopsis Curtisii Sherff, Bot. Gaz. 96: 146. 1934. C. 
Curlissii Sherff, Bull. Jard. Bot. Brux. 13: 289. 1935 (sphalm). 

Herba erecta, verisimiliter perennis, gracilis, forsitan circ. 0.5 
m. alta, subsimplex, caule angulato glaberrimo plus minusve pur- 
pureo-striato. Folia opposita, tenuiter petiolata petiolis glaberrimis 
basi connatis usque ad 2 cm. longis, petiolo adjecto usque ad 8 cm. 
longa, pinnata vel bipinnatisecta, laminae circumambitu triangulato- 
ovata, segmentis primariis circ. 3 vel 5 pallidis membranaceis lanceo- 
latis acriter dentatis secundum venas marginesque nunc sparsissime 
nunc subconferte albido-setulosis. Capitula pauca, 3-adgregata et 
ad rami finem corymbose disposita pedunculis erectis tenuibus glaber- 
rimis 4-8 cm. longis, radiata, pansa ad anthesin circ. 3-3.5 cm. lata 
et circ. 11-13 mm. alta. Involucri pallidi bracteae supra reflexae, 



REVISION OF THE GENUS COREOPSIS 385 

exteriores circ. 8 vel 9, lineari-oblongae, glabrae vel basi extrema 
hispidae, apice acutae, 4 mm. longae, interioribus ovato-oblongis 
apice glanduloso-pubescentibus moderate breviores. Flores ligulati 
circ. 8, aurantiaci, ligula anguste oblongi, apice integri vel sub- 
denticulati, 1.5 cm. longi. Paleae lineari-oblongae, apicem versus 
valde aurantiacae, apice ipso subobtusae. Flores tubulosi extrin- 
secus glabrae vel secundum corollae superne valde aurantiacae 
dentes pubescentes et ad tubi apicem sparsim pilosi; stylorum ramis 
pulcherrime aurantiacis et elongato-appendiculatis. Achaenia ma- 
tura non visa. Ovaria oblanceolata, plana, nitido-brunnea, longi- 
tudinaliter pluristriata, plus minusve alata, marginibus et faciebus 
saltern supra erecte setosa, corpore 4 mm. longa, apice valde 
erecto-setosa et biaristata aristis glaberrimis 1.5 mm. longis. 

Type specimen: Collected by Mrs. Richard C. Curtis from 
material cultivated in 1930 from seeds obtained in Angola, Portu- 
guese West Africa (Field). 

Distribution: Known only from Angola. 

Specimens examined: Mrs. Curtis, cult. 1930 (type, Field; cotype, 
Gray). 

A species offering a superficial resemblance in leaf outline and 
dissection to C. oligoflora. 

75. Coreopsis exilis Sherff, Bull. Jard. Bot. Brux. 13: 288. 1935. 

Herba perennis, erecta, gracillima, simplex, 0.5-1.25 m. alta, 
caule subtetragono glabro infra circ. 3 mm. supra circ. 1.2 mm. 
crasso. Folia petiolata petiolis tenuibus anguste marginatis glabris 
basi vix connatis 1.5-3 cm. longis, petiolo adjecto circ. 7-9 cm. 
longa, pinnatim 3-5-partita, foliolis oblonge lanceolatis (vel superi- 
orum lineari-oblongis), pauciter acridentatis, membranaceis, pallidis, 
ciliatis, utrinque subsparsim adpresso-hispidis, apice acutis. Capi- 
tulum (unicum visum) pedunculatum pedunculo tenui glabrato 
+11 cm. longo, radiatum, pansum ad anthesin circ. 3.5 cm. latum et 
+ 1.5 cm. altum. Involucri bracteae aequilongae, exteriores circ. 
8, adpressae, oblonge lineares, basim versus ciliatae ac moderate 
hispidae supra sparsim adpresso-hispidae vel glabrae, apice acutae, 
longitudinaliter saltern inferne 3-lineata lineis atris, 9-10 mm. 
longae; interiores ovato-oblongae, moderate adpresso-hispidae. 
Flores ligulati circ. 8, aurantiaci, ligula subovati, apice 2-3-denti- 
culati, 1.7 cm. longi. Paleae subacutae, glabrae vel aegre pubes- 
centes, ovariorum aristis superatae. Flores tubulosi extrinsecus 
glabri, supra aurantiaci, corolla circ. 8 mm. longi. Achaenia matura 



386 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

non visa. Ovaria plana, alata, anguste cuneato-obovata, supra 
marginibus faciebusque erecto-setosa, corpore circ. 5.5 mm. longa, 
biaristata aristis acerrimis stramineis infra antrorsum paucisetosis 
supra glabris circ. 4 mm. longis. 

Type specimen: Collected by P. Quarre, No. 419 pro parte, on 
farm of H. Droogmans, Katuba, Katanga, Belgian Congo, May 15, 
1927 (Bruss.; nom. indig. Kibunyubunyu). 

Distribution: Katanga, Belgian Congo. 

Specimens examined: Quarre 419 pro parte (type, Bruss.). 

Similar in habit to C. Curtisii of Angola but differing in exterior 
involucral bracts, which are appressed, moderately hispid below 
the middle, and about 9-10 mm. long (not reflexed, not "glabrous 
except at very base," and not 4 mm. long); in ovarial aristae, which 
are about 4 mm. long and somewhat antrorse-setose below (not 
1.5 mm. long and not extremely glabrous) ; etc. The median leaves 
of the type have the terminal leaflet linear-oblong, about 5 cm. 
long, and few-dentate (1-3 teeth on each side). Root described by 
Quarr as running ("racine tracante"). 

76. Coreopsis oligoflora Klatt, Leopoldina 25: 107. 1889. 
C. oligantha Klatt, Ann. Naturh. Hofmus. Wien 7: 103. 1892. 
Involucri bracteae exteriores lineares, acriter apiculatae, circ. 1 mm. 

latae C. oligoflora sensu stricto. 

Involucri bracteae exteriores late oblongae vel ovatae, apicaliter 
obtusissimae, saepius 2.5-4 mm. latae var. /3 robusta. 

Herba erecta, annua, 5-10 dm. alta, ramosa ramis angulatis, caule 
ramisque sparsim pilis tenuissimis albis elongatis hispidis. Folia 
opposita, pilis tenuissimis argenteo-nitidis elongatis plus minusve 
hispida, breviter petiolata petiolis alato-marginatis basaliter con- 
natis usque ad 1.5 cm. longis, petiolo adjecto 3-6 (-10) cm. longa, 
bipinnatisecta, segmentis majoribus ovatis vel lineari-lanceolatis, 
membranaceis, aegre atropunctulatis, dentatis dentibus acriter 
calloso-apiculatis. Capitula tenuiter pedunculata pedunculis saepius 
7-13 cm. longis, radiata, pansa ad anthesin 4-6 cm. lata et circ. 1 
cm. alta. Involucri hispidi bracteae exteriores circ. 8, lineares, 
nitide acriterque calloso-apiculatae, 6-10 mm. longi et circ. 1 mm. 
latae, interioribus oblongis plerumque aequales. Flores ligulati circ. 
8, lutei vel aurantiaci, ligula oblanceolato-elliptici, apice integri vel 
minutissime denticulati, 2.5-3 cm. longi et usque ad 12 mm. lati. 
Disci florum stigmata terminaliter angustata. Achaenia late 
lineari-oblonga, nigra, plana, apicem versus minute erecto-setosa, 



REVISION OF THE GENUS COREOPSIS 387 

lateraliter alata alls 0.6-0.8 mm. latis, marginibus erecto-setosa, 
corpore circ. 8 mm. longa et cum alis circ. 3-3.5 mm. latis, apice 
erecto-setoso biaristata aristis tenuibus 2 mm. longis antrorsum 
hispidis. 

Type specimen: Collected by Dr. Buchner, No. 32, among bushes, 
Malange, Angola, Portuguese West Africa, March, 1879 (Berl.). 

Distribution: Belgian Congo and south westward across Portu- 
guese West Africa. 

Specimens examined (this list doubtless includes some specimens 
of the more recently described var. robusta) : L. Achten 253B, Luobo- 
Kasai, Belgian Congo (Bruss.) ;F. Allard 65, Kuku Wungu, Belgian 
Congo, 1909 (Bruss.); idem 330, Kikubungu, Belgian Congo, 1910 
(Bruss.); Baum 878, alt. 1,300 meters, between Kulei and Kutsi, 
Mambunda Region, southeastern Angola, Portuguese West Africa, 
April 29, 1900 (Berl.); Bavicchi 14, Esungume Dolo, Belgian Congo, 
1913 (Bruss.); J. Bequaert 113, Bukama, Belgian Congo, May 12, 
1911 (Bruss., 2 sheets); Dr. Buchner 32 (type, Berl.); J. Claessens 67, 
Lubuku, Belgian Congo, September, 1907 (Bruss., 2 sheets); idem 
73, Belgian Congo, August, 1907 (Bruss.); idem 88, Ka-Tembo, 
Belgian Congo, January 18, 1921 (Bruss., 2 sheets}; idem 400, Koloko 
Konbe, Belgian Congo, January, 1910 (Bruss.); F. Flamand, 
Kiamohampa, Belgian Congo, September 20, 1911 (Bruss.); J. 
Gillet 806 and 885, Kisantu, Belgian Congo, 1900 (Bruss.); P. 
Hamerlinck 1502 and 1512, Wombali, Belgian Congo, 1914 (Bruss.); 
Fr. Hens 45, becoming shrubby and growing 1.5 meters high at alt. 
270 meters, marsh of Stanley Pool, Belgian Congo, August 14, 1888 
(Bruss. ; Kew) ; Em. Laurent, between Lusambo and Lomani, Belgian 
Congo, December, 1895 (Bruss.); idem & Marc. Laurent, under- 
growths, Kondue, Belgian Congo, November 25, 1903 (Bruss.); 
Marc. Laurent 512, Bolobo, Belgian Congo, April 15, 1905 (Bruss.); 
E. Lescrauwaet 188, Mokole (Lubi), Belgian Congo, September 12, 
1904 (Bruss.); Alexander von Mechow 33, Pungo Andongo, Angola, 
Portuguese West Africa, January-April, 1879 (Berl., 2 sheets); idem 
459, Malange, Angola, March, 1880 (Berl.); Nelis, Bokala, Belgian 
Congo, May, 1913 (Bruss.); P. Pogge 257, Belgian Congo, March, 
1876 (Berl.); idem 1282, in cultivations, Mukenge tract (campine), 
Belgian Congo, November 6, 1881 (Berl.); idem 1295, eodem loco, 
March 15, 1882 (Berl.); idem 1309, prairie (campine) on the Lualaba 
River, southeastern Belgian Congo, April 24, 1882 (Berl.); A. Sapin, 
N'Dolo, Belgian Congo, May, 1906 (Bruss.); idem, N'Kolokosso, 
Belgian Congo, June 28, 1906 (Bruss.); idem, Sokomini Lake, 



388 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Sankuru (Lubilash) River, Belgian Congo, September, 1906 (Bruss.) ; 
idem, Katola Msek, Belgian Congo, April, 1908 (Bruss., 2 sheets); 
idem, growing on the plains, Munungu, Belgian Congo, April, 1910 
(Bruss.); R. Schlechter 12474, Dolo, Belgian Congo, May- June, 
1899 (Berl.; Bruss.; Kew); Sparano 25, "odor recalling slightly the 
marguerite," Belgian Congo, June 6, 1913 (Bruss.); Hyacinthe 
Vanderyst, Ndembo, Belgian Congo, August 20, 1906 (Bruss.); 
idem, eodem loco, January 7, 1907 (Bruss.); idem, Station at Inkissi 
Falls, Belgian Congo, May, 1907 (Bruss.); idem, Leopoldville, 
Belgian Congo, May 23, 1907 (Bruss.) and May 27, 1910 (Bruss.); 
idem, Kumpako, Belgian Congo, 1908 (Bruss.); idem, N'Boma, 
Belgian Congo, April 28, 1911 (Bruss.); idem B224, Lonzo, Belgian 
Congo, July 26, 1906 (Bruss.); idem 700, Wombali, Belgian Congo, 
May, 1913 (Bruss.); idem 1099, eodem loco, June, 1913 (Bruss.); 
idem 1187, Bandundu, Belgian Congo, June, 1913 (Bruss.); idem 
1341, Wombali, Belgian Congo, June, 1913 (Bruss.); idem 1582, 
Bokala, Belgian Congo, July, 1913 (Bruss.); idem 4304, Bugunu, 
Belgian Congo, May 28, 1914 (Bruss.); idem 4885, Wombali, Belgian 
Congo, 1914 (Bruss.); idem 10097, Kamtsha, Belgian Congo, July, 
1921 (Bruss.); idem 20963, Lulua-gare, Belgian Congo (Bruss.); 

E. Verdick, Lukafu, Katanga, Belgian Congo, March, 1900 (Bruss.) ; 

F. C. Wellman, alt. 1,500 meters, Bailundo District, Benguella 
(Benguela; Abunda) region, eastern Angola (Kew). 

Often confused with Bidens Steppia (Steetz) Sherff, a species 
having more elongate exterior involucral bracts, mostly 11 or 12 
slender-liguled ray florets instead of about 8 rather wide ones, 
leaves larger and more decompound, achenes practically exalate, etc. 

Coreopsis oligiflora var. robusta Sherff, Bot. Gaz. 90: 386. 
1930. 

A specie involucri bracteis exterioribus spathulato-oblongis vel 
oblongis vel ovatis saepius 2.5-4 mm. latis apicaliter obtusissimis 
differt. 

Type specimen: Collected by R. Buttner, No. 408, Bolobo, 
Belgian Congo, November 5, 1885 (Berl.). 

Distribution: Belgian Congo. 

Specimens examined: Joseph Bequaert, Leopoldville, Belgian 
Congo, April 9, 1915 (Bruss.); idem 7205, Belgian Congo, March 
29, 1915 (Bruss.); Buttner 408 (type, Berl.); DeGiorgil42, Kisengwa, 
Belgian Congo, October, 1922 (Bruss.); idem 231, herb of the moist 
plain, Kanda-Kanda, Belgian Congo, November, 1922 (Bruss.); 
D. H. Linder 1 706, moist places at edge of savanna, Bolobo, Belgian 



REVISION OF THE GENUS COREOPSIS 389 

Congo, December 16, 1926 (Berl.); P. Pogge 1292, Mukenge tract 
(campine), Belgian Congo, April 13, 1882 (Berl.); P. Quarre 2173, 
7-9 dm. tall, on plateau, Saeomintra, Belgian Congo, November, 
1930 (Bruss., 2 sheets) ; idem 2826, growing 1 meter tall, Pastorale, 
Division of Lualu, Belgian Congo, November, 1931 (Bruss., 2 sheets) ; 
idem 2921, herb 1.2 meters tall, Lovoi, Kamina, Belgian Congo, 
March, 1932 (Bruss., 2 sheets); Walter Robyns 138, much branched 
at base, 1.5 meters tall, right bank of Guhisi, beyond railroad, 
Kisantu, Belgian Congo, July 3, 1925 (Bruss.); H. Vanderyst 12366, 
between La Labue and La Loange, Belgian Congo, October, 1922 
(Bruss.); idem 21227, Belgian Congo (Bruss.); F. Vermoesen 2403, 
Belgian Congo, September, 1919 (Bruss., 2 sheets). 

Dr. Otto Hoffmann had designated the type varietally under the 
name C. oligantha Klatt in the Berlin Herbarium, but I have not as 
yet found that he ever published the varietal name. 

77. Coreopsis Quarrei Sherff, Bull. Jard. Bot. Brux. 13: 289. 
1935. 

Herba annua, gracilis, erecta, 5 cm. alta, caule moderate 
erecto-ramoso glabrato nunc 1 nunc 2.2 mm. crasso. Folia petiolata 
petiolis usque ad 1.3 cm. longis setoso-ciliatis setis pluriloculatis, 
petiolo adjecto usque ad 5 cm. longa, bipinnatisecta, foliolis mem- 
branaceissimis acute dentatis lobulatisve supra moderate infra 
(interdum sparsius) adpresso-hispidis. Capitula radiata, pansa ad 
anthesin 4 cm. lata et circ. 1 cm. alta, tenuiter pedunculata pedun- 
culis caulem ramosve terminantibus glabris vel summam versus 
aegre setosis usque ad 1.2 dm. altis. Involucri demum hemisphaerico- 
depressi bracteae exteriores circ. 7 vel 8, glabratae vel aegre hispidae, 
lineari-oblongae, supra interdum dilatatae, apice rotundatae vel 
subacutae, 3-4 mm. longae, quam interiores ovato-oblongae sparsim 
setosae dimidio breviores. Flores ligulati 8, aurei, lineari-oblongi, 
apice integri vel bidenticulati, 2.3 cm. longi. Paleae oblongae 
vel lineari-oblongae, apice subcolorato obtusissimae vel subtrun- 
catae, 6-7.5 mm. longae. Flores tubulosi circ. 4 mm. longi, extrin- 
secus praecipue ad tubi summam saepe patenti-hispidi. Achaenia 
plana, brunneo-atra, obovata vel oblonga, alata, corpore 3.5-6 mm. 
longa et alis inclusis 2-2.5 mm. lata, corporis plurisulculati ipsius 
facie ventrali erecte setosa setis e tuberculo ortis, facie dorsali non 
nisi apicem versus tuberculato-setosa, marginibus pectinato-tuber- 
culatis saepius brunneis flavisve numerose erecto-setosa, apice erecte 
setulosa ac minutissime biaristata aristis tenuibus antrorsum his- 
pidis sub 0.5 mm. longis. 



390 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: Collected by P. Quarre, No. 419 pro parte, on 
farm of H. Droogmans, Katuba-Elisabethville, District of Upper 
Katanga, Belgian Congo, May 16, 1927 (Brass.). 

Distribution: District of Upper Katanga, Belgian Congo. 

Specimens examined: Quarre 419 pro parte (type, Brass.). 

Confused by Quarre with C. exilis, from which it differs in being 
an annual, having the exterior involucral bracts 3-4 mm. long and a 
half shorter than the inner ones, the achenial bodies varying from 
3.5 to 6 mm. long, the aristae less than 0.5 mm. long, etc. C. exilis 
is perennial, has involucral bracts subequal and 9-10 mm. long, 
achenial bodies about 5.5 mm. long, aristae about 4 mm. long, etc. 

78. Coreopsis Goffardii Sherff, Bull. Jard. Bot. Brux. 13: 
290. 1935. 

Herba annua, erecta, 2.5-8 dm. alta; caule glabro, subtetragono, 
purpurascenti, suberecte ramoso. Folia petiolata petiolis marginatis 
sparsim hispidis (setis pluriloculatis) 1-3 cm. longis, petiolo adjecto 
usque ad 1 dm. longa, 2-3-pinnatisecta rhachi petiolo marginato 
simile; segmentis ultimis nunc oblongis nunc obovato-oblongis 
nunc plus minusve lanceolatis, utrinque moderate adpresso-his- 
pidula et saepe praecipue supra minutissime glandulosa, membra- 
nacea, dentibus saepius subabrupte minutissimeque apiculatis. 
Capitula ramulos tenues hispidos 1-7 cm. longos terminantia, radiata, 
pansa ad anthesin 3 cm. lata et 11 mm. alta. Involucrum 
hemisphaericum basi dense superne subsparsim adpresso-hispidum, 
bracteis exterioribus circ. 8, lineari-oblongis, superne saepe latioribus, 
tergo longitudinaliter 1-3-atronerviis, apice acutis vel subacutis, 
circ. 4-5 mm. longis, quam interioribus oblongo-ovatis paulum 
brevioribus. Flores ligulati circ. 8, aurantiaci, 1.5 cm. longi, 
ligula anguste oblongi, inferne glabrati superne minutissime adpresso- 
hispiduli, apice subintegri. Paleae oblongae, apice obtusae rotunda- 
taeve ac erecte ciliolatae, ad anthesin quam corollae tubulosae 
breviores. Flores tubulosi extrinsecus glabra vel tubi summa 
(anulo) hispiduli, dentibus saepius aurantiaci, 4-5 mm. longi. 
Ovaria (achaeniis maturis deficientibus) plana, oblongo-obovata, 
faciebus glabratis mediane paucistriata, alata, corpore sub 3 mm. 
longa, marginibus infra obsolete eroso-denticulata summam versus 
erecte setosa, apice cuneato-emarginato erecte setosa ac biaristata 
aristis tenuibus antrorsum hispidulis sub 0.5 mm. longis. 

Type specimen' Collected by P. Quarre, No. 1616, on Mr. 
Goffard's farm, Kipila-Elisabethville, District of Upper Katanga, 
Belgian Congo, May, 1929 (Brass.). 



REVISION OF THE GENUS COREOPSIS 391 

Distribution: Known only from type locality in District of Upper 
Katanga, Belgian Congo. 

Specimens examined: Quarre 1616 (type, Bruss.). 

79. Coreopsis vulgaris Sherff, Bull. Jard. Bot. Brux. 13: 291. 
1935. 

Herba annua, erecta, 4 dm. alta, caule glabrato plus minusve 
tetragono, apicem versus suberecte ramoso. Folia petiolata petiolis 
tenuibus patenti-ciliatis interdum submarginatis usque ad circ. 2 
cm. longis, petiolo adjecto circ. 5-8 cm. longa, 2-3-pinnatifida, 
segmentis primariis 5 ovatis vel ovato-lanceolatis, ultimis mem- 
branaceis utrinque adpresso-hispidis dentatis dentibus acriter 
apiculatis. Capitula pauca (7) , pedunculata pedunculis subrobustis 
sparsissime patenti-hispidis (setis crispis albidisque) 1-4 cm. longis, 
radiata, pansa ad anthesin circ. 3-3.5 cm. lata et 1.2 cm. alta. 
Involucrum magnum depresso-hemisphaericum, 2 cm. latum et 
1 cm. altum, setis acerrimis crispis albidis basi dense alibi sub- 
sparsim conspersum; bracteis exterioribus circ. 8, lineari-oblongis, 
longitudinaliter circ. 3-lineatis lineis atris, apice obtusis vel acutis, 
6-11 mm. longis; interioribus ovato-lanceolatis, apice glanduloso- 
hispidulis, saepius 8-10 mm. longis. Flores ligulati circ. 8, flavi, 
ligula lineari-oblongi, apice integri vel subintegri, circ. 1-2 cm. 
longi. Paleae oblongae, apice atriore subrotundatoque vix apicu- 
latae, aristis superatae. Flores tubulosi extrinsecus glabri. Achaenia 
non visa. Ovaria plana, oblonga vel obovata, faciebus glabrata, 
marginibus (infra breviter supra longe) apiceque erecto-setosa, 
alata, corpore alis inclusis 3-4 mm. longa et 1-1.5 mm. lata, biaristata 
aristis tenuibus antrorsum hispidis circ. 2.5-4 mm. longis et corollae 
tubum dimidio superantibus. 

Type specimen: Collected by A. Becquet, No. 62, at altitude of 
1,000 meters, common on savanna, flowering entire year except in 
dry season, Mukishi-Lomam, District of Middle Katanga, Belgian 
Congo, September, 1927 (Bruss.). 

Distribution : District of Middle Katanga, Belgian Congo. 

Specimens examined : Becquet 62 (type, Bruss.). 

The annual habit, large discs, and comparatively inconspicuous 
rays combine to give this species the appearance of an unattractive 
weed. Referred to Coreopsis (instead oiBidens) with some hesitancy, 
since it is not known that the mature achenes are alate. 

80. Coreopsis Giorgii Sherff, Bull. Jard. Bot. Brux. 13: 
292. 1935. 



392 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Herba annua, erecta, 1 m. alta, caule glabro subtetragono 
tenuiter ramoso. Folia petiolata petiolis tenuibus anguste mar- 
ginatis hispido-ciliatis circ. 1-1.5 cm. longis, petiolo adjecto 7 cm. 
longa, bipinnatifida, segmentis secundariis oblongo-lanceolatis vel 
ovato-lanceolatis membranaceis infra moderate supra subsparsim 
adpresso-hispidis acriter dentatis. Capitula subnumerosa sub- 
corymbose ad ramorum apices disposita, pedicellata pedicellis 
sparsissime setosis 3-6 cm. longis, radiata, pansa ad anthesin circ. 
2.5-3.3 cm. lata et circ. 8-9 mm. alta. Involucrum moderate ad- 
presseque albido-setosum, bracteis exterioribus oblongo-linearibus, 
apice induratis atque acutis subobtusisve 4-6 mm. longis quam 
interioribus ovato-lanceolatis plerumque brevioribus. Flores ligulati 
circ. 8, flavi, ligula lineari-oblongi, apice integri vel minutissime 
denticulati, circ. 1.5-2 cm. longi. Paleae glaberrimae, cuneate 
oblanceolatae, apice atriore rotundatae. Flores tubulosi glabri. 
Achaenia valde obcompressa, nitide brunneo-atra, faciebus pluri- 
striata et (praecipue secundum medium atque apicem versus) erecto- 
setosa, alarum marginibus minutissime numerosissimeque erecto- 
setulosa, corpore alis inclusis exteriora 7 mm. longa et 3.2 mm. 
lata interiora 1 cm. longa et 1.7 mm. lata, omnia apice densissime 
erecto-hispida et biaristata aristis perspicue densissimeque erecto- 
hispidis circ. 1 mm. longis, setulis stramineo-fulvis. 

Type specimen: Collected by De Giorgi, No. 170, Katompe, 
District of Middle Katanga, Belgian Congo, October, 1922 (Bruss.). 

Distribution: District of Middle Katanga, Belgian Congo. 

Specimens examined: De Giorgi 170 (type, Bruss.). 

Related to C. oligoflora and C. Mattfeldii. From C. oligoflora 
it differs at once in having the exterior involucral bracts 4-6 (not 
6-10) mm. long, the ligulate florets about 1-2 (not about 2.5-3) 
cm. long, etc. From C. Mattfeldii it differs in having the exterior 
achenes, wings included, about 3.2 (not 4-5) mm. wide, etc. 

81. Coreopsis Mattfeldii Sherff, Bot. Gaz. 76: 83. 1923. 

Herba erecta, ramosa, verisimiliter circ. 7-10 dm. alta, caule 
subtetragono, glabrato vel disperse piloso, sulcato. Folia petiolata 
petiolis latis 1 cm. longis, petiolis adjectis 4-10 cm. longa, pinna- 
tisecta, utrinque moderatim pilosa, margine parce ciliata, laciniis 
oppositis indivisis vel pinnatisectis, segmentis ultimis linearibus 
(2-6 mm. latis), acutis atque indurato-apiculatis. Capitula corym- 
bosa, radiata, pansa ad anthesin circ. 3.5-4 cm. lata et 9 mm. alta. 
Involucri bracteae dorso hispidae pilis albis multiloculatis atque ad 
basim tumidis, aequilongae (5-8 mm.), exteriores lineari-spathu- 



REVISION OF THE GENUS COREOPSIS 393 

latae, interiores ovato-lanceolatae. Flores ligulati circ. 8, flavi, 
ligula lanceolati vel obovato-lanceolati, striati, apice obscure minu- 
teque denticulati, 1.6-2 cm. longi et 4-9 mm. lati. Paleae hyalinae 
late oblongae, apice subobtusae, circ. 8 mm. longae. Disci florum 
stigmata angustatissimo-caudata. Achaenia valde obcompressa, 
nigrescentia, alata alis 1-1.5 mm. latis, faciebus glabra vel ad costas 
(et supra sparsim) hispida, margine apiceque hispida, biaristata 
aristis tenuibus stramineis sursum hispidis 1-2 mm. longis, exteriora 
obovata 6-8 mm. longa et (alis inclusis) 4.25-5 mm. lata, interiora 
oblanceolata circ. 8 mm. longa et (alis inclusis) 4 mm. lata. 

Type specimen: Collected by Alexander Carson, No. 75, Fwamba, 
Lake Tanganyika, German East Africa, 1894 (Berl.). 

Distribution: Known only from type locality in German East 
Africa. 

Specimens examined: Carson 75 (type, Berl.; cotype, Kew). 

Named very appropriately after Dr. Johannes Mattfeld, who 
most generously assisted in placing at my disposal many of the 
Berlin Herbarium specimens of African Coreopsis and Bidens for 
detailed study. It bears a strong superficial resemblance to Bidens 
Steppia (Steetz) Sherff, but can easily be distinguished by its achenes 
and involucral bracts. The many mature achenes on the type differ 
remarkably from the ones on the type and other specimens (already 
cited in my monograph of the genus Bidens) of Bidens Steppia. For 
B. Steppia they are at most widely oblong-linear in outline, 7.5-9 
mm. long and 2.5-3 mm. wide; there is merely a thickened lateral 
ridge, wings being absent. For Coreopsis Mattfeldii the achenes are 
much thinner and wider, the outer ones being obovate, 6-8 mm. long 
and 4.25-5 mm. wide, the inner ones being oblanceolate, about 8 
mm. long and 4 mm. wide; all are winged, and even on submature 
fruiting heads the rounded upper ends of the wings, adjacent to the 
aristae, are easily visible to the eye. The external bracts about 
equal the inner ones in length in C. Mattfeldii; in Bidens Steppia 
they tend to exceed the inner ones by 2-6 mm. 

The nearest ally in Coreopsis is C. oligiflora Klatt. 

82. Coreopsis injucunda Sherff, Bull. Jard. Bot. Brux. 13: 
293. 1935. 

Herba annua, erecta, ramosa, 4-10 dm. alta, caule ramisque 
subtetragonis glabris. Folia petiolata petiolis tenuibus marginatis 
hispido-ciliatis setis pluriloculatis usque ad 2.5 cm. longis, petiolo 
adjecto 6 cm. longa, bipinnatisecta segmentis linearibus subacriter 



394 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

apiculatis membranaceis utrinque marginibusque sparsim albido- 
hispidis. Capitula moderate numerosa, pedunculata pedunculis 
ramos terminantibus sparsim hispidis tenuibus saepius 1-1.5 dm. 
longis, radiata, pansa ad anthesin circ. 4-4.5 cm. lata et 1.2-1.4 cm. 
alta. Involucri hispidi bracteae exteriores circ. 8, subulato-lance- 
olatae, basi connatae, apice acutae, circ. 4-6 mm. longae, quam 
interiores oblongo-ovatae paulo breviores. Flores ligulati circ. 8, 
flavi, ligula oblonge lineares, apice integri, circ. 2.5 cm. longi. Paleae 
oblongae, apice obtusissimae vel subtruncatae, achaeniorum aristas 
superantes. Florum tubulosorum corollae circ. 5 mm. longae, 
extus glabrae. Achaenia plana, atra, obovata vel oblonga, corpore 
4.5-6.5 mm. longa et alis inclusis circ. 2-2.3 mm. lata, faciebus 
pluristriata dorsali parce ventrali moderate erecto-setulosa, margini- 
bus apiceque numerose erecto-ciliata, biaristata aristis tenuibus 
stramineis antrorsum hispidulis sub 1 mm. longis. 

Type specimen: Collected by P. Quarre, No. 1662, on Mr. 
Goffard's farm, Kipila-Elisabethville, District of Upper Katanga, 
Belgian Congo, May, 1929 (Bruss., 2 sheets). 

Distribution: District of Upper Katanga, Belgian Congo. 

Specimens examined: Quarre 1662 (Bruss., 2 type sheets). 

83. Coreopsis Borianiana Schz. Bip. ex Schweinf. Verh. Zool. 
Bot. Ges. Wien 18: 684. 1868. C. Borianiana var. cannabina Schz. 
Bip. loc. cit. C. guineensis 0. & H. in Oliver, Fl. Trop. Afr. 3: 390. 
1877; etiam in Hutch. & Dalz. Fl. West Trop. Afr. 2: 143, /. 217. 
1931. C. Chevalieri 0. Hoffm. & Muschl. Bull. Soc. France 57, 
Me"m. 8: 118. 1910. C. chrysopterocarpa Chiov. Ann. Bot. Roma 
9: 75. 1911. 
Involucri bracteae exteriores 9-12, uniseriales, 0.7-1.5 cm. longae. 

C. Borianiana sensu stricto. 

Involucri bracteae exteriores 18-22, biseriales, plerumque 1.4-1.8 
cm. longae var. /3 multiplex. 

Herba annua, erecta, plus minusve glabra, 6-1.5 dm. alta, caule 
aegre subtetragona vel etiam fere teres. Folia opposita saepius 
breviter petiolata petiolis latis marginatis plerumque 3-12 mm. 
longis, petiolo adjecto 5-13 (etiam -18) cm. longa, simplicia vel 
2-3-partita, lamina segmentisve remote acriterque denticulatis 
(denticulis saepe subcapilliformibus), linearibus, utrinque elongato- 
angustatis, plerumque 4-10 mm. latis, membranaceis, minutissime 
spinuloso-ciliatis. Capitula non numerosa, tenuiter pedunculata 
pedunculis 5-18 cm. longis ad apicem saepe abrupte dilatatis ac 



REVISION OF THE GENUS COREOPSIS 395 

paucis bracteis bracteae involucri exterioris valde similibus obsitis, 
radiata, pansa ad anthesin 5-7 cm. lata et 1.3-2 cm. alta. Involucri 
bracteae exteriores 9-12, uniseriales, anguste lineares, tergo glabrae 
vel hispidae, ad apicem saepe minute spinuloso-ciliatae, acriter vel 
subacriter (saepe indurate-) apiculatae, 0.7-1.5 cm. longae et 0.6-1 
mm. latae; interiores lanceolato-oblongae vel ovato-oblongae, apice 
plerumque dorso interdum pubescentes, saepius paulo longiores. 
Flores ligulati circ. 8, aurei, ligula obovato-oblongi, 8-14-striati, 
apice obtuso obscure vel manifeste denticulati, 2.5-3 cm. longi et 
8-14 mm. lati. Paleae oblongo-oblanceolatae, demum interdum 
crassiusculae, circ. 10-18 mm. longae et 2.5-4.5 mm. latae. Disci 
florum stigmata caudato-appendiculata. Achaenia valde obcom- 
pressa, alata, circumambitu (alis inclusis) ovato-orbiculata, corpore 
ipso nigra lineari-oblonga tuberculato-setosa 9-15 mm. longa 
et 1.2-2.3 mm. lata, alis badia vel argentea nitida dense setoso- 
ciliata circ. 2.8-3.8 mm. lata, apice setosa et biaristata aristis erectis, 
sursum hispidis, circ. 2.5-4 mm. longis. 

Type specimen: Collected by Boriani, No. 42, in the Fesoglu, 
northeastern Africa, 1839 (Mus. V.). 

Distribution: Across Africa from Eritrea to Se'ne'gal and reaching 
southward to Kamerun and Togo. 

Specimens examined: C. Barter 933, Nigeria, 1857-1859 (Berl.; 
Gray); Boriani 42 (type, Mus. V.; etiam fragmentum typi in herb. 
Schz. Bip. in Par.); Aug. Chevalier 2083, Koulitsoro, Sudan, 1899 
(Berl. ; Bruss.) ; idem 9633, Massenya (Massenia), Bagirmi (Baguirmi), 
Sudan, 1902-1904 (Berl.); idem 9721, Kolke'te and Moito, Dar-el- 
Hadjer, Baguirmi Nord, Territory of Chari, September 6-9, 1903 
(herb. Chevalieri; type of C. Chevalieri 0. Hoffm. & Muschl.); Dr. 
Kersting 14, alt. 350 meters, Trogode, Kerkri, Togo, October 28, 
1897 (Berl.) ; idem 422, alt. 400 meters, Sokode'-Basari, Togo, October, 
1901 (Berl.); Lecard 290, Se'ne'gal (Bruss.); C. Ledermann 5243, 
growing 1-1.5 meters high, alt. 380 meters, between Tschamba and 
Doreba, Karnerun, September 18, 1909 (Berl., 2 sheets); A. Pappi 
7733, alt. 1,000 meters, Baza Cunama-Barentu, Erythrea, November 
28, 1908 (Flor., type of Coreopsis chrysopterocarpa Chiov.); T. Pfund, 
Dar-Fur, 1876 (Berl.) ; idem 336, Takari on the Rahad, Kordofan, 
July, 1875 (Berl., 2 sheets); Dr. Rowland, common in savannas, 
western Lagos, 1893 (Berl., 2 sheets) ;F. Schroeder 74, alt. 350 meters, 
Sokod Farm, Togo, October 19, 1900 (Berl., 2 sheets); G. Schwein- 
furth (Flora of Gallabat No. 439), Matamma, Gallabat (Galabat) 
region, northwestern Abyssinia, middle of October, 1865 (herb. 



396 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Schz. Bip. in Par.); idem (Flora of Gallabat No. 585}, eodem loco 
et tempore (Berl., type material of var. cannabina Schz. Bip.). 

Coreopsis Chevalieri 0. Hoffm. & MuschL, based on Aug. Chevalier 
9721, is seen from its type specimen (which was very kindly obtained 
for my study from the Chevalier Herbarium by Dr. H. Humbert 
of Paris) to belong here. 

Coreopsis Borianiana var. ft multiplex Sherff, Bot. Gaz. 90: 
385. 1930. 

A specie involucri bracteis exterioribus biserialibus 18-22, plerum- 
que 1.4-1.8 cm. longis quam interioribus saepe longioribus differt. 

Type specimen: Collected by "Graf. Zech." under No. 168, 
Kete Kratschy, Togo, October 21, 1898 (Berl.). 

Distribution: Known only from type locality in Togo. 
Specimens examined : Zech. 168 (type, Berl.). 

84. Coreopsis togensis Sherff, Bot. Gaz. 76: 87. 1923. 

Herba perennis, caule glabro, rubro-viridi, sulcato, subtetragono, 
usque ad 2 m. alto. Folia petiolata petiolis 0.5-4 cm. longis, petiolo 
adjecto 10-18 cm. longa et usque ad 20 cm. lata, pinnatim 3- vel 
5-partita, foliolis lanceolatis, apice saepe longe attenuatis, grosse 
vel moderatim serratis dentibus acriter apiculatis infra pallidioribus, 
margine minute ciliatis, aliter glabris, lateralibus usque ad 11 cm. 
longis et 3 cm. latis, terminali usque ad 16 cm. longo et 4 cm. lato. 
Capitula pauca, longe pedunculata pedunculis maximam partem 
glabris usque ad 18 cm. longis, radiata, pansa ad anthesin 7-9 cm. 
lata et circ. 1.5 cm. alta. Involucrum non plerumque pubescens 
nisi basi, bracteis exterioribus 7-10, magnis, foliaceis, lanceolatis, 
apice angustatis, interdum ciliatis, 1-2 (et etiam usque ad 4) cm. 
longis, interiores lanceolatas aequantibus, vel multo superantibus. 
Flores ligulati 7-9, lutei, ligula late elliptico-lineares vel ovato- 
oblanceolati, apice minute vel grosse profundeque (sinu usque ad 
1 cm. alto) dentati, 3-4.5 cm. longi et 0.7-2 cm. lati. Paleae tenuiter 
lineari-oblongae pluribus lineis percursae apice subacutae, omnino 
plus minusve flavidae, 1-1.5 cm. longae. Disci florum stigmata 
angusto-elongata. Achaenia obcompressa, brunneo-atra, oblongo- 
linearia, plerumque alata, margine antrorsum ciliata, supra faciebus 
et apice erecto-hispida, corpore 8-10 mm. longa et (alis inclusis) 
2.5-3 mm. latis, biaristata aristis nudis vel basim versus antrorsum 
1- vel 2-spinulosis, circ. 1 mm. longis. 



REVISION OF THE GENUS COREOPSIS 397 

Type specimen: Collected by R. Buettner, No. 142, on steppe 
behind Ketzabenki, Togo, August 29, 1890 (Berl., 2 sheets). 

Distribution: Togo. 

Specimens examined : Buettner 142 (Berl., 2 type sheets) ; W.Busse 
3416, at alt. of about 700 meters, Hausberg, near Misahoehe, Togo, 
December 7, 1904 (Berl., 2 sheets); Kling 60, Jege, Bismarckburg, 
Togo, 1889 (Berl.); idem 166, eodem loco, October 30, 1889 (Berl.). 

On the type specimens the exterior involucral bracts are much 
larger than in Busse's specimens, being twice the length of the inner 
ones. The type has also larger leaves and ligulate florets, some of 
the ligules reaching 2 cm. in width, but all the other characters are 
too much alike to warrant specific or probably even varietal separa- 
tion of the Busse plants. The achenial measurements are taken from 
Busse's plants, as the achenes of the type are immature. 

85. Coreopsis scabrifolia Sherff, Bot. Gaz. 76: 86. 1923. 

Herba simplex vel parce ramosa, caule quadrangulato, glabro, 
sulcato, perenni e basi lignea, verisimiliter circ. 1 m. alto, non 
robusto. Folia nunc sessilia, nunc petiolata petiolis alatis et elongatis 
usque ad 4 cm. longis, petiolo adjecto 8-15 cm. longa et 1-3 cm. lata, 
lanceolata (raro oblanceolato-linearia), indivisa (raro pauca pinnatim 
lobata), acriter et interclum grosse dentata, utrinque acuminata vel 
tantum acuta, scabra, minute spinuloso-ciliata, infra ad venas 
sparsim piloso-hispida et pallidiora. Capitula pauca (1-4), sub- 
tenuiter pedunculata pedunculis maximam partem glabris 3-12 cm. 
longis, radiata, pansa ad anthesin 5-6 cm. lata et circ. 1 cm. alta. 
Involucri bracteae exteriores 8-11, lineares vel lineari-lanceolatae, 
dorso basi hispidae, 1-1.5 cm. longae, interiores lanceolatas ae- 
quantes vel paulo superantes. Flores ligulati circ. 8, flavi, ligula 
elliptici vel ovato-lanceolati, 2-2.5 cm. longi et 5-12 mm. lati. 
Paleae tenuiter lineari-oblongae, circ. 5-7-purpureo- vel brunneo- 
striatae, circ. 1 cm. longae. Disci florum stigmata incrassata ter- 
minaliter caudata. Achaenia (alis inclusis) brunneo-atra, late linea- 
ria, valde obcompressa, moderatim sed inconspicue alata, antrorsum 
setosa, corpore 9-11 mm. longa et (alis inclusis) 2-3 mm. lata, 
biaristata, aristis nudis vel sursum 1-3-spinulosis, 1.5-2.5 mm. longis. 

Type specimen: Collected by T. Kassner, No. 2776, under trees, 
Kundelungu, Belgian Congo, May 15, 1908 (Berl.). 

Distribution: Southeastern Belgian Congo. 

Specimens examined: S. De Giorgi, vicinity of Elisabethville, 
Belgian Congo, 1923 (Bruss.); Homble 329, bordering stream, 



398 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

eodem loco, May, 1912 (Brass., 2 sheets; nom. indig., musombe'); 
Kassner2776 (type, Berl.: cotypes, Brit., 2 sheets). 

The general habit is deceivingly like that of Bidens Baumii (0. 
Hoffm.) Sherff, and, were it not for the achenial and involucral 
differences, distinction between the two would be at times difficult. 

86. Coreopsis palmata Nutt. Gen. Amer. 2: 180. 1818. Calli- 
opsis palmata Spreng. Syst. Veg. 3: 611. 1826. Coreopsis pauciflora 
Lehm. Ind. Sem. Hort. Hamb. 1833; cf. Linnaea 10: Litt. 76. 1836. 
Coreopsis praecox Fresen. Linnaea 13: Litt. 93. 1839; cf. Ind. Sem. 
Hort. Bot. Francofurt. 1838. 

Herba perennis, erecta, 5-9 dm. alta, caulibus ad basim hori- 
zontalem radicantibus glabris vel ad nodos hirsutis, simplicibus 
vel paulo ramosis, tetragonis, foliosis. Folia opposita, principalia 
(summa plerumque indivisa exclusa) petiolata petiolis planis alatis 
0.4-2.5 cm. longis et 3-6 mm. latis, petiolo adjecto 2-7 cm. longa, 
rigida, crassiuscula, suberecta, pedato- vel palmato-tripartita seg- 
mentis integris vel irregulariter 2-3-lobatis, segmentis (vel lobis) 
ultimis petiolo similibus oblongo-linearibus supra atque infra glabra 
marginibus scabrido-ciliatis apice saepius obtusis vel subrotundatis, 
plerumque 3-6 mm. latis. Capitula plerumque 1-3 (rarius usque 
ad 6), breviter pedunculata pedunculis glabratis 1-4 cm. longis, 
radiata, pansa ad anthesin nunc 2.5-3.5 nunc etiam 4-6 cm. lata, 
0.9-1.5 cm. alta. Involucri bracteae exteriores 8-12, late lineari- 
oblongae, pallidae, obtusae, tergo glabrae, marginibus scabrido- 
ciliatae, 5-9 mm. longae, interdum subbiseriatim dispositae, quam 
interiores oblongo-obovatae irregulariter (et praecipue superne) 
ciliolatae paulo breviores. Flores ligulati circ. 8, flavi, ligula oblongo- 
obovati, apice plus minusve rotundato obsolete dentati, 1.5-2.7 
cm. longi. Paleae angustissime oblongo-lineares (filiformes), supra 
moderate dilatatae, apice acutae, demum 6.5 mm. longae. Florum 
tubulosorum stylorum rami anguste conico-appendiculati. Achaenia 
valde obcompressa, elliptico-oblonga, basim versus attenuata, 
glabra, subnigra, anguste alata, apice alis productis breviter biden- 
tata, omnino 5-6.5 mm. longa et 1.8-2.3 mm. lata. 

Type specimen: No particular specimen cited. Habitat given by 
Nuttall as "on the open plains of the Michigan Territory, Illinois, 
and Lower Louisiana." 

Distribution: Wisconsin and eastern Manitoba southwardly to 
Indiana, Missouri, and Oklahoma. 

Specimens examined: J. A. Allen, Iowa (Field); C. F. Baker, St. 
Croix Falls, northwestern Wisconsin, August 15, 1900 (Gray); 



REVISION OF THE GENUS COREOPSIS 399 

C. R.Barnes, Converse, Missouri, June 30, 1877 (Field); M. S.Bebb, 
prairies, Winnebago County, Illinois, September, 1859 (Gray) ; idem, 
prairies, Marion County, Illinois, 1860 (Field); Robert Bebb 50, 
Clarke, Indiana, 1894 (Field); idem 1048, open hillside, Lauderdale, 
Wisconsin, July 19, 1902 (Field) ; H. C. Benke 3967, Crystal Lake, 
Illinois, June 30, 1923 (Field); 0. W. Blakley 1451, in woods near 
Page, Oklahoma, June 20, 1914 (Gray); E. Bourgeau, Winnipeg 
Valley, Canada, 1859 (Berl.); Fred Brendel, Peoria, Illinois (Field); 
idem, Illinois (Berl.); Buckley, Illinois (Gray); 5. F. Bush 160, Mon- 
tier, Missouri, June 30, 1894 (Berl.; Gray); idem 8249, prairies, 
Purdy, Missouri, August 17, 1905 (Gray); /. W. Clokey 2428, dry 
prairie soil, Niantic, Illinois, June 29, 1915 (Field; Gray) ; George L. 
Clothier & H. N. Whitford 1037, prairies, Brown County, Kansas, 
July 30, 1897 (Gray); Robert Combs & C. R. Ball 517, Ames, Iowa, 
June 25, 1897 (Berl.; Gray); John M. Coulter, field near LaPorte, 
Indiana, July 22, 1878 (Field); idem, Kankakee River, Indiana, 
1882 (Field, 2 sheets) ; R. I. Cratty, Iowa (Field) ; idem, Armstrong, 
Iowa, July 18, 1897 (Gray); idem 16, Emmet County, Iowa, 1879 
(Field); Arthur W. DeSelm 275 and 276, south of Monee, Illinois, 
July 6, 1913 (Field) ; idem 394, highways near Otto, Illinois, July 19, 
1913 (Field);/ 1 . W. Dewart, St. Cloud, Minnesota, September, 1892 
(Mo.) ; H. Eggert, St. Louis, Missouri, June 29, 1875 (Berl., 2 sheets) ; 
idem, dry ground, eodem loco, June 18, 1886 (Gray) ; idem & Sydow, 
prairie and dry hills, eodem loco, July 1, 1874 (Gray) ; W. H. Emig 
146, along roads near Duroc, Missouri, July 2, 1913 (Mo.); ex herb. 
T. J. & M. F. L. Fitzpatrick, common, woods and prairies, Allamakee 
County, Iowa, June 27, 1895 (Field); W. D. Frost, Willmar, Minne- 
sota, July, 1892 (Berl.); Frank C. Gates 598, Ravenswood, Illinois, 
July 17, 1905 (Field); idem 3148, Zion City, Illinois, July 19, 1909 
(Field); H. A. Gleason, in the black oak association, Shirland, 
Illinois, June 29, 1908 (Gray); idem, bunch-grass prairies, Milroy, 
Illinois, July 6, 1908 (Gray) ; idem 326, Forsythe, Illinois, June 10, 
1896 (Gray) ; Elihu Hall, dry, open timber lands, Athens, Illinois, 
1861 (Field, 2 sheets); J. R. Heddle 698, Madison, Wisconsin, 
August 1, 1907 (Field); W. Hoffman, Corn Creek, Ozark Region, 
Missouri, June 21, 1870 (Berl., 2 sheets); J. M. Holtzinger, Lamoille, 
Minnesota, June, 1895 (Field) ; E. W. D. Holway, prairies, Decorah, 
Iowa, July 16, 1888 (Field); Hortus Lipsiae, ex Illinois (Berl.); 
D. Houghton, 200 miles above Falls of St. Anthony, upper Mississippi 
River (Gray); idem, dry prairie of upper Mississippi River above 
Falls of St. Anthony, July, 1832 (N.Y.); 0. E. Lansing, Jr. 307, 



400 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

prairie soil, Hammond, Indiana, June 28, 1898 (Field); idem 2797, 
roadside, Chicago, Illinois, July 19, 1910 (Field; Gray); idem 3113, 
alt. 445 meters, fallow field east of Mansfield, Missouri, June 5-12, 
1911 (Field; Gray); /. A. Lapham, Milwaukee, Wisconsin (Gray); 
Leavenworth, Louisiana (Gray) ; ex Lehmannio ipso sub nom. Coreop- 
side pauciflora (Gray; N.Y.); Ray N. Lloyd, prairies, Ravenswood, 
Illinois, July 11, 1887 (Field) ; W. H. Manning, Lake City, Minne- 
sota, August 13, 1883 (Gray) ; F. E. McDonald, dry prairies, Peoria, 
Illinois, June, 1891 (Field); idem, eodem loco, 1900-1901 (Field); 
idem, eodem loco, June, 1904 (Gray); idem, eodem loco, June, 1915 
(Field); Mead, Illinois, 1850 (Gray); Edgar A. Mearns 144, Fort 
Snelling, Minnesota, July 16, 1888 (Field); R. T. Morgan, vicinity 
of Fountaindale, Illinois, August, 1877 (Berl.);E. J. Palmer 18078, 
dry, open woods, near Ironton, Missouri, June 26, 1920 (Gray); 
Harry N. Patterson, prairies near Oquawka, Illinois, July (Field); 
idem, near Oquawka, June 20, 1872 (Field); A. S. Pease, dry soil 
south of Champaign, Illinois, October 9, 1920 (Gray); idem 12394, 
Champaign, September 11, 1909 (Gray) ; Donald C. Peattie 8, moist 
prairie, East Chicago, Indiana, July 31, 1920 (Field); Ns. Riehl 
41, hills, St. Louis, Missouri, June, 1838 (Berl.); Benjamin L. Robin- 
son, Normal, Illinois, July, 1887 (Gray) ; C. 0. Rosendahl 656, Spring 
Grove, Minnesota, June 30, 1902 (Gray) ; J. H. Sandberg, Hennepin 
County, Minnesota, August, 1889 (Field, 2 sheets); idem, copses, 
eodem loco, July, 1890 (Field) ; Savage & Stull 60, Ironton, Missouri, 
June 16, 1897 (Field) ; J. H. Schuette, St. Anthony Park, Minnesota, 
July 2 and 14, 1888 (Field); idem, eodem loco, July 8, 1888 (Field; 
Gray); idem, St. Anthony, Minnesota, July 16, 1888 (Field); idem, 
Plover, Wisconsin, July 26, 1888 (Gray) ; idem, Green Bay, Wiscon- 
sin, August 13, 1899 (Gray) ; idem, Green Lake, Wisconsin, August 
13, 1899 (Field) ;E. P. Sheldon, Sleepy Eye, Minnesota, July, 1891 
(Field) ; idem, Fort Snelling, Minnesota, June, 1895 (Field) ; Earl E. 
Sherff 284, St. Louis, Missouri, July 7, 1910 (Gray); idem 1642, dry 
sandbarrens west of Kankakee, Illinois, August 3, 1912 (Field); 
idem 1781, dry hill, Elgin, Illinois, August 27, 1912 (Field, 2 sheets); 
H. C. Skeels 375, Gougar's Prairie, Joliet, Illinois, July 14, 1904 
(Field) ; Ernest C. Smith 510, roadsides, Hinsdale, Illinois, September 
2, 1902 (Field); G. W. Stevens 2713, open woods in mountain valley 
near Page, Oklahoma, September 9, 1913 (Gray); J. T. Stewart, 
bluffs around Peoria, Illinois, 1867 (Field) ; E. B. Uline, Fish Lake, 
LaPorte County, Indiana, July, 1891 (Field); idem, Mishawaka, 
Indiana, July 1, 1891 (Field, 2 sheets) ; L. M. Umbach, prairies, Du 
Page, Illinois, July 6, 1898 (Berl.; Field; Gray). 






REVISION OF THE GENUS COREOPSIS 401 

87. Coreopsis verticillata L. Sp. PL 907. 1753. Chrysanthe- 
mum Marianum, etc. Pluk. Mant. pi. 344, / -4- 1769. Coreopsis 
tenuifolia Ehrh. Beitr. 7: 168. 1792; Schkuhr Bot. Handb. edit. 2. 
pi. 260b, fig. ad dextr. 1808. Coreopsis verticillata var. tenuifolia 
Michx. Fl. Bor. Amer. 2: 139. 1803. Bidens verticillata (L.) Baill. 
Hist. PI. 8: 305. 1886. 

Herba perennis, glabra, caulibus e basi plus minusve horizontali 
erectis, 5-9 dm. altis, subangulatis, superne ramosis. Folia opposita, 
sessilia, 3-6 cm. longa, laminis palmatim 3-partitis, segmentis 1-2- 
pinnatis, ultimis lineari-filiformibus, acriter apiculatis, 0.3-1 mm. 
latis. Capitula saepius pauca, saepe corymbosa, tenuissime pedun- 
culata pedunculis glabris vel sparsissime pilosis, radiata, pansa ad 
anthesin 2.5-5 cm. lata et 6-9 mm. alta. Involucri bracteae exteriores 
circ. 8, oblongae vel oblongo-lineares, 4-6 (raro -9) mm. longae, sub- 
glabrae, apice saepius obtusae, quam interiores elliptico-oblongae 
minutissime pubescentes plerumque paulo breviores. Flores ligulati 
circ. 8, flavi, ligula elliptico-oblongi, apice integri vel minute denticu- 
lati, 1.2-2.5 cm. longi. Paleae lineares vel filiformes, superne latiores, 
apice acutae, 4 mm. longae. Flores tubulosi flavi, pubescentes, 
stylorum ramis apice cuspidato-appendiculatis vel acriter conicis. 
Achaenia plana, oblongo-obovata, faciebus glabris subatra, margini- 
bus anguste flavido-alata, apice exaristata vel (alis parce productis) 
vix bidentatis dentibus laceratis, omnino 3-5 mm. longa et 1-1.7 
mm. lata. 

Type specimen: No type cited by Linnaeus, although Virginia 
was given for the habitat. Linnaeus' first citation of literature is 
that of Gronovius, Virg. 105, whence the type would be understood 
to be a plant collected by John Clayton in Virginia (as then bounded) 
and perhaps still extant (Brit.). 

Distribution: Maryland to Florida, Alabama, and Arkansas. 
Reported by some authors as extending to Kentucky and Nebraska. 
Probably not native farther north, although frequently cited for 
Ohio, Michigan, Canada, etc. (cf. Torr. & Gray, Fl. N. Amer. 2: 
342. 1843; Gray, Syn. Fl. N. Amer. 1, pt. 2: 293. 1884). Occasionally 
escapes from gardens. 

Specimens examined: Anon. 576, Arkansas, September, 1835 
(Berl.); anon., cult., June, 1818 (Berl.); W. W. Ashe, Chapel Hill, 
North Carolina, July, 1898 (Field); M. S. Bebb, District of Columbia, 
1863 (Field; Gray); ex Biltmore Herb. 437a, open pine woods, 
Spartanburg, South Carolina, June 17, 1897 (Field; Gray); Walter 
Blair, dry ridges, Prince Edward County, Virginia, June, 1881 



402 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

(Gray) ; S. F. Blake 9542, in colony, sandy field, Washington, District 
of Columbia, July 11, 1926 (Pom.); ex herb. J. W. Blankinship, 
Southern Pines, North Carolina, July 18, 1895 (Gray); Buckley, 
southeastern United States (Gray); Agnes Chase 2382, in woods, 
Riverdale, Maryland, June 22, 1904 (Field); eadem 2474, rocky 
woods, West Chevy Chase, Maryland, July 12, 1904 (Field); A. H. 
Curtiss, Virginia (Field); idem, Bedford County, Virginia, June 10, 
1871 (Gray) ; Earl Jerome Grimes 3768, border of woods beside rail- 
way, vicinity of Williamsburg, Virginia, June 16, 1921 (Gray); 
ex herb. D. Gronovii (Berl. ; cotype?) ; James Hall, Virginia, 1828- 
1834 (Field) ; R. M. Harper 5, alt. 345 meters, rocky bank of Town 
Creek on Sand Mountain, near Chavies, Alabama, November 24, 
1905 (Gray; Mo.); A. A. Heller 933, about Suffolk, Virginia, June 
8-13, 1893 (Field, 3 sheets; Gray); Theodor Holm, open thickets, 
Brookland, District of Columbia, June, 1894 (Gray); Hort. Berol., 
cult., July 25, 1815 (Berl.); idem, cult., September, 1834 (Berl.); 
idem, cult., 1837 (Berl.); H. D. House 150, sandy woods, Takoma 
Park, District of Columbia, August 3, 1904 (Kew) ; G. MacCarthy, 
District of Columbia (Field); E. L. Morris 105, sandy roadsides 
along Potomac River, from Chain Bridge to opposite Cabin John 
Bridge, Virginia, July 1, 1899 (Field); Thomas C. Porter, 40 miles 
south of Lynchburg, Virginia, July 17, 1880 (Field) ;L.F. & Fannie 
R. Randolph 150, dry, sandy soil, sterile hillside field, vicinity of 
Lorton, Virginia, June 14, 1922 (Gray) ; Arthur Schott, near George- 
town, District of Columbia, August, 1857 (Field) ; idem, woods near 
Georgetown, June 22, 1860 (Field); Earl E. Sherff 253, cult., St. 
Louis, Missouri, July 6, 1910 (Field); idem 5004, cult., Harvard 
Univ. Bot. Gard., August 12, 1929 (Field); John K. Small, alt. 
75 meters, on banks of Little Long Creek, Albemarle, North Carolina, 
August 22, 1894 (Field); idem & A. A. Heller 372, between Gold 
Hill and Falls of the Yadkin, Stanley County, North Carolina, 
August 18, 1891 (Field, 3 sheets) ; E, S. Steele, Washington, District 
of Columbia, August, 1889 (Pom., 2 sheets). 

88. Coreopsis pulchra Boynt. in Small, Fl. S. E. United States 
1277, 1340. 1903. 

Herba perennis, erecta, glaberrima, 4-7 dm. alta, perspicue 
foliosa, caulibus moderate angulatis, plus minusve fastigiato-ramosis. 
Folia opposita, sessilia, principalia 2.5-4.5 cm. longa, palmatim 
circ. 3-partita foliolis mox subpalmatimque 2-3-partitis, segmentis 
ultimis anguste oblongeque linearibus, integris, 1-nervatis, mem- 
branaceis, eciliatis, apice obtusis, 2-4 cm. longis et 0.5-1.5 mm. 



REVISION OF THE GENUS COREOPSIS 403 

latis. Capitula numerosa, corymbosa, tenuiter pedunculata pedun- 
culis saepius 4-10 mm. longis, radiata, pansa ad anthesin 3-5 cm. 
lata et circ. 9-11 mm. alta. Involucri bracteae exteriores 8-10, 
lineari-oblongae, superne saepe sensim vel subabrupte latiores, 
apice obtusae, glabrae, 5-8 mm. longae; interiores ovatae, apice 
pubescentes, plerumque tantum 4-6 mm. longae. Flores ligulati 
6-8, intense flavi, ligula elliptico-oblongi vel anguste obovati, 
apice integri vel vix denticulati, 1.6-2.4 cm. longi. Paleae anguste 
lineares, apice saepius dilatatae, 6 mm. longae. Flores tubulosi 
corollis purpureo-brunnei, stylorum ramis apice acriter conicis. 
Achaenia obcompressa, oblongo-obovata, anguste alata, apice corona 
humili lacerataque ornata, circ. 4 mm. longa. 

Type specimen : Biltmore Herb., No. 14728 of the former Biltmore 
Herbarium. Specimen was collected in "northern Alabama," but 
doubtless was destroyed, along with many others, by the flood waters 
at Biltmore. 

Distribution: In region of the Lookout Mountain, northeastern- 
most Alabama, and perhaps in adjacent Georgia. 

Specimens examined: T. G. Harbison, cult., Highlands, North 
Carolina, August 1, 1905 (Gray, 2 sheets) and August, 1906 (Gray) ; 
Charles Mohr, Mentone, Alabama (N.Y., 2 sheets); idem, open, 
rocky woods, Mentone, near the Falls, Lookout Mountain, September 
18, 1892 (U.S.); idem, damp rocks, DeSoto Falls, near Mentone, 
September 12, 1898 (U.S.); Albert Ruth 612, sandy soil, Lookout 
Mountain, DeKalb County, Alabama, July, 1898 (N.Y.); idem 624, 
near DeSoto Falls, Lookout Mountain, July, 1898 (U.S.). 

89. Coreopsis Delphinifolia Lam. Encycl. 2: 108. 1786; Bot. 
Mag. pi. 156, sub nom. C. verticillata. 1791. Ceratocephalus Delphinii 
foliis Vaill. Hist. Acad. roy. sci. 1720: 328. 1722; Ehret PI. et Pap. 
Rar. Depict, pi. 9, f. 1749. C. verticillata var. linearis Michx. Fl. 
Bor. Amer. 2: 138. 1803 (fide Torr. & Gray, Fl. N. Amer. 2: 342. 
1843). C. discolor Link, Enum. Hort. Berol. 2: 353. 1822. 
C. Delphiniifolia Lam. ex Ind. Lond. 2: 294. 1930 (sphalm). 
Foliorum segmenta magna ex parte 2-3.5 mm. lata. 

C. Delphinifolia sensu stricto. 
Foliorum segmenta magna ex parte 0.3-2 mm. lata . var. /3 chlooidea. 

Herba perennis, caulibus erectis, gracilibus, tetragonis, glabris 
vel ad nodos hispidis, ad summam ramosis, 6-15 dm. altis, per totam 
longitudinem foliosis, internodiis folia vix aequantibus. Folia 
opposita, sessilia, suberecta vel parce patentia, 5-9 cm. longa, summa 



404 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

saepe indivisa integraque, alia palmatim 3-partita, foliolo mediano 
plerumque rursus in 3-5 segmenta diviso, foliolis lateralibus simplici- 
bus vel raro 2-partitis, segmentis ultimis linearibus membranaceis 
acutis glabris integris 1-6 (plerumque 2-3.5) mm. latis. Capitula 
corymbosa, tenuissime pedunculata pedunculis glabris 2-7 cm. 
longis, pansa ad anthesin 3-5 cm. lata et circ. 8-10 mm. alta. Invo- 
lucri glabri bracteae exteriores 8-10, oblongo-lineares, obtusae, 4-6 
mm. longae, quam interiores ovatae nunc vix longiores nunc vix 
breviores. Flores ligulati circ. 8, flavi, ligula anguste elliptico- 
oblongi, apice integri vel parce dentati, 1.3-2.5 cm. longi. Paleae 
filiformes, ad summam paulo dilatatae, achaenia superantes. Flores 
tubulosi corolla brunnei, stylorum ramis apice tenuiter caudatis. 
Achaenia obcompressa, circumambitu (alis brunneis 0.5-1 mm. latis 
inclusis) elliptico-oblonga vel obovata, 4.5-6 mm. longa et (alis 
inclusis) 2-4 mm. lata, glabra, corpore subnigra, apice minute 2- 
dentata dentibus fimbriatis. 

Type specimen: The species was founded directly upon Cerato- 
cephalus Delphinifolius Vaill. as illustrated by Ehret (loc. cit.) and 
supported by the material cultivated in the King's Garden at Paris. 
The native habitat was given as Virginia. 

Distribution: Virginia to Georgia and Alabama. 

Specimens examined: Liberty Hyde Bailey, cult., Ithaca, New 
York, July 24, 1935 (Field; sub nom. C. microphylla); ex Biltmore 
Herb., sub num. 2064A, rare in moist soil, Eastman, Georgia, 
June 4, 1900 (N.Y.); A. H. Curtiss 6825, low ground near Millen, 
Georgia, July 1, 1901 (Gray); herb. Drs. A. C. Hexamer & F. W. 
Maier, wet pine woods, Summerville, South Carolina, May 27, 
1855 (Gray); Hort. Berol (Berl.); Hort. Cantabr., 1842 (Gray); 
Hort. Lipsiae (Berl.); Hort. Paris., 1820 (Berl.); Michaux, North 
America (Berl.); Earl E. Sherff 5003, cult, in Harvard Univ. Bot. 
Gard. (Hort. Cantabr.), August 12, 1929 (Field); John K. Small, in 
and about the Altamaha River Swamp, Liberty County, Georgia, 
June 18-21, 1895 (Field); J. T. Stewart, near Charleston, South 
Carolina, June 1, 1865 (Field); K. A. Taylor, pine barren, Summer- 
ville, South Carolina, June 10, 1891 (Field). 

Coreopsis Delphinifolia var. /3 chlooidea Sherff, Bot. Gaz. 
94: 596. 1933. 

E specie foliorum segmentis magna ex parte tantum 0.3-2 
(nonnullis usque ad 3.2) mm. latis differt. 



REVISION OF THE GENUS COREOPSIS 405 

Type specimen: Collected by Samuel Bottsford Buckley, in pine 
woods, mountains of Alabama (Gray). 

Distribution: Georgia and Alabama. 

Specimens examined : Buckley, in pine woods, etc. (type, Gray) ; 
Roland M. Harper 545, dry pine barrens, Sumter County, Georgia, 
August 31, 1900 (Field). 

90. Coreopsis major Walt. Fl. Carol. 214. 1788. C. senifolia 
Michx. Fl. Bor. Amer. 2: 138. 1803. C. maior Walt, ex Ind. Lond. 
2: 295. 1930. 

Folia simplicia var. e Oemleri. 

Folia principalia tripartita. 

Foliola 1-3 (rarius -5) mm. lata var. 8 linearis. 

Foliola plerumque 5-25 mm. lata. 

Foliola plerumque 5-11 mm. lata var. 7 rigida. 

Foliola latiora, plerumque 1-2.5 cm. lata. 

Foliola plus minusve pubescentia C. major sensu stricto. 

Foliola glabra var. stellata. 

Herba perennis, e basi horizontali erecta, gracilis, supra suberecte 
ramosa ramis tenuibus, 5-9 dm. alta, caule ramisque angulatis plus 
minusve pubescentibus. Folia opposita, sessilia, divaricata vel sub- 
erecta, summa simplicia integraque, alia tripartita, segmentis nunc 
subrhomboideo-ovalibus nunc elliptico-lanceolatis vel etiam lanceo- 
latis (rarissime lineari-lanceolatis), membranaceis, integris, plus 
minusve pubescentibus, ciliatis, apice obtusis, terminali 4-7 cm. 
longo et 1.2-2.5 cm. lato, lateralibus plerumque paulo minoribus. 
Capitula subcorymbosa tenuiter pedunculata pedunculis glabris 
vel sparsim hispidis saepius 2-8 cm. longis, radiata, pansa ad anthesin 
2.7-4 ( -5) cm. lata et 7-9 mm. alta. Involucri plus minusve hispidi 
bracteae exteriores circ. 8, lineari-oblongae, superne saepe dilatatae, 
apice obtusae, 4-8 mm. longae, interiores ovato-oblongae vel lance- 
olato-ovatae subaequales. Flores ligulati circ. 8, flavi, ligula oblongo- 
lineares vel clliptico-oblongi, apice integri vel denticulati, 1.5-2.5 
cm. longi. Paleae lineares, supra paulo dilatatae, achaenia multo 
superantia. Florum tubulosorum stylorum rami acriter conici. 
Achaenia valde obcompressa, elliptico-oblonga vel obovata, sub- 
nigra, glabrata, alata alis subbrunneis 0.2-0.7 mm. latis, omnino 
4.5-6 mm. longa et 2-4 mm. lata, apice alis productis breviter biden- 
tata vel plurisetosa. 

Type specimen: No specimen cited. The remnants of Walter's 
collection, however, are in London (Brit.). 



406 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Distribution: South Carolina and Kentucky southwardly to 
Georgia and Mississippi. 

Specimens examined: C. F. Baker, Blount Springs, Alabama, 
May 5, 1898 (Pom.); S. B. Buckley, Alabama, July, 1841 (Mo.); 
ex Cambridge Bot. Gard., Cambridge, Massachusetts, August, 1845 
(Berl.); A. H. Curtiss 6471, dry thickets near Stone Mountain, 
Georgia, June 12, 1899 (Gray); John Davis 1855, Stone Mountain, 
Georgia, June 18, 1921 (Gray; forma foliis glabratis); A. Dedrick, 
dry, sandy soils (Field) ; F. S. Earle 2060, Auburn, Alabama, May 28, 
1899 (Field, 2 sheets); idem 2061, eodem loco, May 28, 1899 (Pom.); 
idem & C. F. Baker, eodem loco, August 17, 1897 (Pom.); H. Eggert, 
Sherwood, Tennessee, June 8, 1897 (Field); ex herb. Hexamer et 
Maier, along fences, Abbeville County, South Carolina, June, 
1855 (Gray); Miss Anna King 116, dry border of woods, Mammoth 
Cave, Kentucky, July 2, 1916 (Field) ; B. F. Leeds, Chattanooga, 
Tennessee, September 29, 1892 (Field) ; Dr. Mohr, Mobile, Alabama 
(Field); Philip A. Munz 1589, dry, open woods, near Montgomery, 
Alabama, June 10, 1917 (Pom.); Josephine Skehan 137, Ocean 
Springs, Mississippi, June, 1892 (Gray; forma var. stellatae adpro- 
pinquans) ; John K. Small, alt. 300-420 meters, on northwest slope of 
Stone Mountain, Georgia, July 3, 1893 (Field) ; John Donnell Smith, 
grassy pine woods, Orangeburg District, South Carolina, August 9, 
1884 (Field). 

Coreopsis major var. /3 stellata (Nutt.) Robins. Rhodora 10: 
68. 1908. C. stellata Nutt. Journ. Acad. Phila. 7: 76. 1834. C. 
senifolia var. stellata (Nutt.) Torr. & Gray, Fl. N. Amer. 2: 342. 1843. 

Glabra, foliolis plerumque angustioribus, saepe longioribus, sae- 
pius rhomboide lineari-lanceolatis, terminali usque ad 1 dm. longo 
(rarissime monstroseque trifido vel tripartito). 

Type specimen: No particular specimen cited. 

Distribution: Virginia to southernmost Ohio and southwardly to 
Florida and Alabama. 

Specimens examined: Anon. (Mt. Desert Nurseries), Bar Harbor, 
Maine, August, 1906 (Gray); Samuel M.Bain 95 p.p., sandy hills, 
Henderson, Tennessee, July, 1893 (Gray); H. C. Beardslee & C. A. 
Kofoid, alt. 900 meters, sandy woods, Swain, North Carolina, 
August, 1891 (Gray); Beyrich, in thickets, Georgia (Berl.); ex 
Biltmore Herb. 175a and 6, light soil in upland woods, North Carolina, 
July 7, 1897 (175a, Field; Gray: 175b, Gray); J. R. Churchill, woods, 
Lookout Mountain, near Chattanooga, Tennessee, June 2, 1911 



REVISION OF THE GENUS COREOPSIS 407 

(Gray) ; John Davis 2097, dry, open woods, Tryon, North Carolina, 
June 29, 1920 (Gray); idem 7818, dry woods, near Anderson, South 
Carolina, August 30, 1917 (Mo. ; forma var. rigidae adpropinquans) ; 
A. H. Curtiss, Bedford County, Virginia, July 1, 1868 (Field); idem, 
eodem loco, June 30, 1871 (Gray, 2 sheets; forma nonnullorum 
foliorum foliolis tripartitis) ; idem, eodem loco, July 3, 1872 (Field) ; 
idem 1487, alt. 600 meters, rocky woods. Chilhowee Mountains, 
Tennessee, June (Berl., 2 sheets; Field); F. S. Earle & C. F. 
Baker, Auburn, Alabama, May 28, 1898 (Field) ; George Engelmann, 
Norristown, eastern Tennessee, August 27, 1876 (Mo.); Juliet 
Fauntleroy 654, near Huddleston, Virginia, August 9, 1914 (Gray); 
Gates, Alabama (Gray); Asa Gray & John Carey, mountains of 
'Virginia and North Carolina, July, 1841 (Gray; var. rigidae adpro- 
pinquans); James Hall, Montgomery County, Alabama, 1828-1834 
(Field) ; T. G. Harbison, Waynesville, North Carolina, 1897 (Gray) ; 
idem, Highlands, North Carolina, 1905 (Gray); Roland M. Harper 
46, alt. 205 meters, dry oak woods, Athens, Georgia, June 25, 1900 
(Berl.; var. rigidae adpropinquans); idem 131, alt. 205 meters, 
eodem loco, July 3, 1900 (Gray); A. Arthur Heller 99, vicinity of 
Salisbury, North Carolina, June 21, 1890 (Field, 2 sheets); idem 
1013, alt. 540 meters, near Hickory, North Carolina, June 23, 1893 
(Field; var. rigidae adpropinquans); Hooker, Kentucky (Berl.); 
Hortus Cantabr., cult., 1842 (Gray); idem, cult., 1873 (Gray); ex 
Hort. Lipsiae (Berl.); Arthur H. Howell 808, Anniston, Alabama, 
June 10, 1913 (U.S.); F. W. Hunnewell, 2nd, dry woods, White 
Sulphur Springs, West Virginia, July 4-6, 1914 (Gray); T. H. 
Kearney, Jr., White Cliff Springs, Tennessee, July 14, 1894 (Field, 
2 sheets); B. F. Leeds, Chattanooga, Tennessee, September 29, 
1892 (Field, 2 sheets) ; E. E. Magee, Hayward County, North Caro- 
lina, September, 1897 (Gray); idem, Waynesville, North Carolina, 
September 9, 1897 (Gray); idem, alt. 750 meters, Highlands, North 
Carolina, September 2, 1902 (Gray) ; idem, eodem loco; below Satulah 
Mountain, August 2, 1902 (Gray);E.L. Morris 1050, alt. 620 meters, 
shaded river bank near Spanishburg, West Virginia, July 23, 1900 
(Field) ; Thomas C. Porter, middle Georgia, 1846 (Gray) ; L. F. & 
Fannie R. Randolph 1066, dry, sterile soil, edge of woods, southeast 
of Granite Falls, North Carolina, July 30, 1922 (Gray, var. rigidae 
adpropinquans); Riddell, Kentucky (Gray);BenjaminL. Robinson 79, 
Asheville, North Carolina, August 2, 1893 (Berl.; Gray);F. Rugel, 
Florida, 1844 (Gray; Mo.); ex herb. Margaret P. Russell, Asheville, 
North Carolina (Gray); Albert Ruth, sandy woods, Knoxville, Ten- 
nessee, June, 1893 (Field); idem, gravelly soil and rocky places, 



408 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Mount Nebo, eastern Tennessee, July, 1893 (Mo., 2 sheets); idem, 
open woodlands, Knoxville, June, 1895 (Berl.) ; idem, woods, Lookout 
Mountain, Georgia, July 20, 1897 (Gray) ; Earl E. Sherff 5007, cult., 
Harvard Univ. Bot. Gard., August 12, 1929 (Berl.; Del.; Field); 
W. L. Sherwood, Waynesville, North Carolina, July, 1897 (Gray); 
idem, Tuckasiegee Valley, North Carolina, September 20, 1897 
(Gray); B. Shimek, Monte Sano, Alabama, July, 1891 (Field); 
C. W. Short, hilly lands, Kentucky (Field) ; Howard Shriver, Wythe- 
ville, Virginia, July, 1877 (Berl.; Field); John K. Small, alt. 900 
meters, on Round Top Mountain, west of Seven Mile Ford, Smyth 
County, Virginia, July 2, 1892 (Field; Gray); idem, alt. 660 meters, 
on Farmer Mountain, on New River, Carroll County, Virginia, July 
12, 1892 (Field); idem, alt. 750 meters, Nick's Creek, base of Pine 
Glade Mountain, Smyth County, Virginia, August 5, 1892 (Field); 
idem & A. Arthur Heller, alt. 1,350 meters, Green's Hill, near Blowing 
Rock Mountain, Watauga County, North Carolina, July 10, 1891 
(Field); iidem, between Cranberry and Linville, North Carolina, 
July 17, 1891 (Field) ; iidem, southern slopes of Grandfather Moun 
tain, North Carolina, July 25, 1891 (Field) ; iidem, alt. 1,050 meters 
Rocky Knob, near Blowing Rock Mountain, Caldwell County 
North Carolina, July 24, 1891 (Field; Gray); iidem 328, Blowing 
Rock Mountain, Watauga County, North Carolina, July 29, 189! 
(Field, 3 sheets); iidem (similiter) 328, eastern slopes of Blowing 
Rock Mountain, Caldwell County, North Carolina, July 31, 189" 
(Field, 3 sheets); John Donnell Smith, Warm Springs, Madison 
County, West Virginia, July 29, 1880 (Field); idem, Caesar's Head 
South Carolina, July, 1881 (Field); Roland Thaxter, Cullowhee 
North Carolina, June-July, 1887 (Gray); S. M. Tracy 9049, Warm 
Springs, Georgia, May 20, 1905 (Berl.; Field; Gray); G. R. Vasey 
northern Alabama, 1878 (Field) ; idem, North Carolina, 1878 (Gray) 
William C. Werner 92, Ironton, Ohio, 1892 (Gray) ; Karl M. Wiegand 
& W. E. Manning 3414, sandy bank by roadside, 4 miles west oj 
Raleigh, North Carolina, June 27, 1927 (Pom.); iidem 3415, edge of 
dry pine woods, 12 miles south of Asheboro, North Carolina, June 
30, 1927 (Pom.}; Charles Wright, northern Georgia (Gray). 

Coreopsis major var. 7 rigida (Nutt.) F. E. Boynton in Small, 
Fl. S. E. United States 1276, 1340. 1903. C. senifolia var. rigida 
Nutt. Gen. Amer. 2: 180. 1818. C. Wrayi Nutt. Journ. Acad. Phila. 
7: 76. 1834. C. senifolia Hook. Bot. Mag. 63: pi 3484- 1836 (non 
Michx.). C. Delphinifolia var. rigida (Nutt.) Torr. & Gray, Fl. N. 



REVISION OF THE GENUS COREOPSIS 409 

Amer. 2: 342. 1843. C. rigida Nutt. ex Torr. & Gray, loc. cit. ; cf. Ind. 
Kew. 1: 617. 1895. 

Folia rigidula, tripartita, plerumque suberecta, foliolis plerumque 
lineari-lanceolatis, faciebus saepissime glabris marginibus scabridis, 
saepius 3-7 cm. longis et 5-11 mm. latis. 

Type specimen: No type cited, but habitat was given as Georgia. 

Distribution: Virginia and Tennessee southwardly to Georgia. 

Specimens examined: Anon., ex Nutallio ipso, loc. ignoto (Gray, 
sub nom. Coreopside Wrayi Nutt.); Harley Harris Bartlett 1638, 
pine barren, vicinity of Thomson, Georgia, July 23, 1909 (Pom.); 
William M. Canby, Florence, South Carolina, July 3, 1878 (Field); 
H. Eggert, dry woods on Yellow River, Gwinne County, Georgia, 
July 27, 1897 (Mo.) ; Roland M. Harper, sunny, rocky hillside, vicin- 
ity of Sawdust, Georgia, June 18, 1927 (Gray; Mo.); Hort. 
Cantabr., cult., 1849 (Gray) ; C. G. Lloyd, near Rockwood, Tennessee, 
August 10, 1880 (Field); C. Mohr, dry pine woods, Bladon (Bladen), 
Alabama, July, 1875 (Field); H. W. Ravenel, Santee Canal, South 
Carolina, July, 1846 (Gray); Albert Ruth, dry, sandy soil, Chil- 
howee Gap, Tennessee, July, 1893 (Mo.) ; idem, sandy soil, Knoxville, 
Tennessee, May, 1896 (Mo.); H. Shriver, Wytheville, Virginia, 1875 
(Field); John K Small & A. Arthur Heller, north of Hickory, North 
Carolina, June 25-26, 1891 (Field); Huron H. Smith 2526, Blue 
Ridge Mountains, Fannin County, Georgia, July 26, 1909 (Field). 

Coreopsis major var. 6 linearis Small, Bull. Torr. Bot. 
Club 22: 48. 1895. 

Parce inconspicueque pubescens, caule 3-6 dm. alta. Folia tri- 
partita segmentis linearibus, 3-11 cm. longis et 1-3 (rarius -5) mm. 
latis utrinque acuminatis. Capitula etiam ligulae achaeniaque paulo 
minora. 

Type specimen: Collected by John Kunkel Small, at altitude of 
300-330 meters, on Little Stone Mountain, DeKalb County, Georgia, 
July 7, 1893 (N.Y., 2 sheets). 

Distribution: South Carolina and Central Georgia. 

Specimens examined: J. K. Small, Little Stone Mountain, etc. 
(type, N.Y., 2 sheets); John Donnell Smith, grassy pine woods, 
Orangeburg District, South Carolina, August 9, 1884 (Field; Gray). 

Coreopsis major var. e Oemleri (Ell.) Britton, Mem. Torr. 
Bot. Club 4: 131. 1894. C. Oemleri Ell. Sketch Bot. S. Carol. & 
Georgia 2: 435. 1824. 

Var. stellatae similis sed foliis simplicibus differt. 



410 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: Collected by Oemler, near the junction of the 
Broad and Saluda rivers, vicinity of Columbia, South Carolina. 

Distribution: North Carolina to Georgia. 

Specimens examined : None. 

Asa Gray (Syn. Fl. N. Amer. 1, pt. 2: 294. 1884) treated this as 
"the abnormal entire-leaved form" of var. stellata. An examination 
of the specimens at Gray Herbarium shows no material of the 
"entire-leaved form," however, and it is probable that Gray was 
guided largely by the inscription on A. H. Curtiss's label for a 
sheet of normal var. stellata at Gray Herbarium. This has the 
observation, "with simple leaves not uncommon." Gray's treat- 
ment was followed essentially by Robinson and Fernald (Gray's 
New Manual, ed. 7. 838. 1908). It may be noted, too, that Boynton 
(in Small, Fl. S. E. United States 1276. 1903) made the var. Oemleri 
to include var. stellata. Boynton's complete omission of the entire- 
leaved plants from his description, however, may well lead to the 
suspicion that he had unintentionally overlooked them. Small 
(Bull. Torr. Bot. Club 22: 48. 1895) describes "a peculiar state" 
of C. major having "leaves undivided, there being two opposite and 
entire leaves at each node in place of the normal three-parted ones." 
Clearly, he referred to plants of var. Oemleri. He states further: 
"It seems to have been first collected by Dr. and Mrs. Britton at 
Black Mountain Station, North Carolina. Later Mr. Heller secured 
it near Salisbury, N.C., and last season (1893) I came upon it at the 
western base of Stone Mountain, Georgia." Apparently the variety 
is somewhat less distinct than the other varieties of C. major (even 
they, however, pass from one into another by various transitional 
forms; cf. A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 294. 1884), but seems 
to bear a relation to its parent species comparable to that of var. 
Smithii to Coreopsis tripteris. 

91. Coreopsis tripteris L. Sp. PI. 908. 1753; Hook. Bot. Mag. 
pi. 3583. 1837. Anacis tripteris (L.) Schrank in Miinch. Math, et 
Nat. 5: 7 (fide DC. Prodr. 5: 568. 1836). Chrysostemma tripteris 
(L.) Less. Syn. Compos. 227. 1832 (fide DC. loc. cit). Coreopsis 
tripteris var. /3 Torr. & Gray, Fl. N. Amer. 2: 341. 1843. Coreopsis 
tripteris vars. Deamii et intercedens Standl. Rhodora 32: 34. 1930. 

Folia divisa. 

Foliorum segmenta subrhomboidea var. 7 subrhomboidea. 

Foliorum segmenta late elliptico-linearia vel anguste oblongo- 

lanceolata C. tripteris sensu stricto. 

Folia simplicia var. /3 Smithii. 



REVISION OF THE GENUS COREOPSIS 411 

Herba perennis, e basi horizontal! erecta, plerumque glabra rarius 
pubescens, plus minusve pallida, 1-3 m. alta, caulibus teretibus 
summam versus ramosis. Folia opposita, principalia petiolata petiolis 
0.5-3 cm. longis, petiolo adjecto 6-12 cm. longa, 3-partita, foliolo 
mediano saepe rursus 3-partita, segmentis late elliptico-linearibus vel 
anguste oblongo-lanceolatis, apice acutis vel subobtusis, marginibus 
aegre revolutis, faciebus glabris vel rarius pubescentibus; folia summa 
minora, saepe simplicia et subsessilia. Capitula (contusa odorata) 
subcorymbosa, tenuiter pedunculata pedunculis saepius 3-8 cm. 
longis, radiata, pansa ad anthesin 3-5 cm. lata et 8-10 mm. alta. 
Involucri saepius glabri bracteae exteriores circ. 8, oblongo-lineares, 
obtusae, circ. 2-3 mm. longae, interiores oblongo-ovatae 4-6 mm. 
longae. Flores ligulati plerumque 7 vel 8, flavi, ligula plus minusve 
elliptico-oblongi, apice subintegri vel dentati, 1.2-2.4 cm. longi. 
Paleae filiformes, apicem versus dilatatae, striis purpureis percursae, 
5-7 mm. longae. Flores tubulosi primo plus minusve flavi demum 
brunnei vel purpurei, stylorum ramis apice breviter caudato-appendi- 
culatis. Achaenia brunneo-nigra, obcompressa, faciebus glabra, 
circumambitu (alis angustis subbrunneis inclusis) cuneato-oblonga 
vel cuneato-obovata, omnino 5-7 mm. longa et 2.8-4.2 mm. lata, 
apice emarginato setis minimis erectis munita. 

Type specimen: No particular type was cited. The habitat was 
given as Virginia and the first cited reference was the Coreopsis foliis 
subternatis integerrimis of the Hort. Upsal. 269. Other citations were 
in the following order: Rudbeckia foliis compositis integris, Roy. 
lugdb. 181; Chrysanthemum virginianum, folio acutiore laevi trifoliatof. 
anagyridis folio, Moris. Hist. 3. p. 21. s. 6. t. 3. f. 44; Raj. Suppl. 215. 

Distribution: Massachusetts, southern Ontario, and Wisconsin, 
southwardly to Georgia, Mississippi, western Louisiana, and eastern 
Kansas. 

Specimens examined: H. C. Beardslee & C. A. Kofoid, alt. 600 
meters, sandy banks, Great Smoky Mountains, Swain County, 
North Carolina, August, 1891 (Field; Gray); M. S. Bebb, Fountain- 
dale, Illinois (Gray); Florence Beckwith 54, Dallas City, Illinois, 
September, 1917 (Field); H.C.Benkel841, Elgin, Illinois, September 
8, 1916 (Field) ; Biltmore Herb. 173a, moist grounds and bordering 
streams, Biltmore, North Carolina, August 1, 1897 (Gray; Pom.); 
0. W. Blakley 3420, open woods near Page, Oklahoma, August 27, 
1914 (Mo.); J. W.Blankinship, Greene County, Missouri, September 
7, 1888 (Gray); Fred W. Brendel, Peoria, Illinois (Field); A. B. 
Burgess 433, railroad bank, Flowerfield, Michigan, September 30, 



412 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

1903 (Field); idem 513, shrubbery at roadside, Portage, Michigan, 
October 5, 1903 (Field); W. W. Calkins 135, prairie, Berwyn, Illinois 
(Field); idem 145, common, Berwyn, Illinois, August 26, 1907 
(Field) ;S.H.& D. R. Camp, dry ground, Jackson County, Michigan, 
August 18, 1897 (Field); iidem, Jackson County, Michigan, August 
18, 1898 (Field); Agnes Chase, open woods, Rogers Park, Illinois, 
August 27, 1896 (Field); Virginius H. Chase, dry prairie north of 
Wady Petra, Illinois, August 12, 1896 (Field); J. R. Churchill, shore 
of Lake Michigan, Indiana, September 4, 1893 (Pom.); John Davis, 
near LaGrange, Missouri, June 9, 1915 (Mo.); C. C. Deam & H. A. 
Gleason, dry upland woods, Bluffton, Indiana, September 1, 1904 
(Gray); Arthur DeSelm 499, highways south of Tucker, Illinois, 
August 7, 1913 (Field); idem 621, highway near Kankakee, Illinois, 
September 7, 1913 (Field) ; W. M. Dick, Lorain County, Ohio, July 
31, 1895 (Gray); S. S. Dickey 96, dry roadside, Warren County, 
Pennsylvania, August 25, 1923 (Gray); Charles K. Dodge, near 
Port Huron, Michigan, August 4, 1895 (Field); C. W. Duesner, 
Miller, Indiana, 1908 (Field) ; F. S. Earle, creek bottoms, Auburn, 
Alabama, September 4, 1899 (Field); H. Eggert, prairie and dry 
hills, St. Louis, Missouri, August 25, 1874 (Gray); idem, woods, 
St. Louis, Missouri, August 23, 1887 (Gray); idem 122, central 
United States, August 30, 1875 (Berl.) ; James Galen 4%89, woods 
and thickets, eastern United States, 1883 (Gray) ; Frank C. Gates 808, 
Edgewater, Illinois, August 23, 1905 (Field); idem 10104, prairie, 
Carthage, Illinois, September 15, 1916 (Field); H. A. Gleason, in 
strip of original prairie, east of Rantoul, Illinois, October 5, 1907 
(Gray); idem 812, prairie soil along roadsides, southwest of Dal ton 
City, Illinois, August 22, 1899 (Gray); idem 1813, dry upland woods, 
Grand Tower, Illinois, August 28, 1900 (Gray); idem 1977, prairie, 
Illinois, September 4, 1900 (Gray); idem 2753, dry upland woods, 
Herod, Illinois, August 23, 1902 (Gray); Grassly, Miller, Indiana, 
August 31, 1880 (Field); J. M. Greenman 544, Tygart Junction, 
West Virginia, September 24, 1904 (Field; Gray) ; idem 3991, Webster 
Grove, Missouri, September, 1920 (Mo.); Dr. Hale, Louisiana 
(Gray); Elihu Hall, barrens, etc., Athens, Illinois, 1861 (Field; 
Pom.); idem, eodem loco, August, 1864-1867 (Field); idem, eodem 
loco, August, 1868 (Field); Roland M. Harper 1954, alt. 50 meters, 
bank of Flint River, near Albany, Georgia, August 27, 1903 (Berl.; 
Field; Gray); A. A. Heller, vicinity of Salisbury, North Carolina, 
August 21, 1890 (Field); idem, near mouth of Tucquan Creek, 
Lancaster, Pennsylvania, September 1, 1890 (Gray); idem & E. 



413 

Gertrude Halbach, near McCall's Ferry, Pennsylvania, September 

9, 1893 (Field); A. A. & E. G. Heller 4129, alt. 90 meters, near 
Texarkana, Arkansas, August 23, 1898 (Field; Gray); Albert S. 
Hitchcock, wet ground, Iowa City, Iowa, 1888 (Calif.); Hort. Bero- 
linensis, cult., September 11, 1876 (Berl.); ex Hort. Goetting. (Berl.); 
Hort. Lipsiae, cult. (Berl.) ; Hort. Bot. de I'Ecole de Medecine de Paris, 
August, 1820 (Berl.); J. W. Huett, Ottawa, Illinois (Gray); T. H. 
Kearney, Jr. 553, Clear Creek, Bell County, Kentucky, September, 
1893 (Field; Gray; Pom.; etiam in Field qua varietati Smithii 
adpropinquans) ; Miss Anna King 534, dry banks or in prairie land, 
Glenwood, Illinois, September 20, 1918 (Field) ; eadem 566, dry soil 
in thickets of waste field, eodem loco, September 20, 1918 (Field) ; 
W. Krebs, Cleveland, Ohio (Berl.); 0. E. Lansing, Jr. 428, open 
woods, Chicago, Illinois, August 14, 1898 (Field); idem 647, prairie 
land, West Pullman, Illinois, July 27, 1899 (Field); idem 1646, 
sandy woods near dunes, near Porter, Indiana, September 16, 1902 
(Field); idem 2832, border of oak woods, Gibson, Indiana, August 
13, 1910 (Field; Gray); idem 3822, sandy roadsides, Starved Rock, 
Illinois, September 7-8, 1914 (Field); idem 3911, open, sandy woods, 
Pine, Indiana, August 22, 1915 (Field) ; Ray N. Lloyd, Ravenswood, 
Illinois, August 31, 1887 (Field) ; John Macoun, thickets, Sandwich, 
Ontario, August 4, 1892 (Gray) ; F. E. McDonald, open, dry woods, 
Peoria, Illinois, August, 1900 (Field) ; idem, eodem loco, September, 
1901 (Field) ; idem, frequent in rich woods, eodem loco, August, 1904 
(Gray); Michaux, North America (Berl.); Charles F. Millspaugh 
3865, railroad ditch, Lakeside, Berrien County, Michigan, September, 
1914 (Field); W. S. Moffatt 492, thickets, Pine, Indiana, August 31, 
1895 (Field; Gray) ; Albert Hanford Moore 2552 and 2553, sandy east 
bank of Tygart's Valley River, near Tygart Junction, West Vir- 
ginia, September 24, 1904 (Gray); E. L. Moseley, Oxford, Ohio, 
August 21, 1895 (Field) ;L. H. Pammel 621, Ames, Iowa, September 

10, 1897 (Gray); H. N. Patterson, dry bluffs, Henderson County, 
Illinois, August, 1871 (Field); Donald C. Peattie 164, sandy fields, 
Tremont, Indiana, August 13, 1920 (Field); idem 2029, Griffith, 
Indiana, August 15, 1925 (Field); Thomas C. Porter, banks and 
islands of Susquehanna River, Lancaster County, Pennsylvania, 
August 22, 1861 (Gray); Miss Carrie A. Reynolds, Stony Island, 
Illinois, August 24, 1909 (Field); Riddell, Ohio (Gray); Robert 
Ridgway 60, Olney, Illinois, September 7, 1917 (Field); Benjamin 
L. Robinson, rich prairie soil, Hendrix, Illinois, August 31, 1904 
(Gray); idem 56, Bloomington, Illinois, September, 1893 (Berl.); 



414 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

F. Rugel, in valley of Blue Ridge Mountains, North Carolina, 
August, 1841 (Berl.); Albert Ruth 69, woods, Lookout Mountain, 
Georgia, July 19, 1897 (Gray) ; H. P. Sartwell, Maumee River, Ohio 
(Field); Earl E. Sherff 1038, near Jefferson Barracks, Missouri, 
August 11, 1910 (Field) ; John K. Small, north of Marysville, Penn- 
sylvania, August 15, 1888 (Field); idem, vicinity of mouth of Tuc- 
quan Creek, Lancaster County, Pennsylvania, September 1, 1890 
(Field); idem, eodem loco, October 11, 1890 (Field); idem, alt. 480 
meters, in canyon at Tallulah Falls, Rabun County, Georgia, 
August 4, 1893 (Field) ; idem, alt. 900 meters, about Estatoah Falls 
on Mud Creek, Rabun County, Georgia, August 12, 1893 (Field); 
idem & A. A. Heller 100, vicinity of Faith Post Office, North Caro- 
lina, August 14, 1891 (Field); Frank Thone 232, Starved Rock, 
Illinois, May-September, 1921 (Field); S. M. Tracy 8670, Taylor- 
ville, Mississippi, August 20, 1903 (Field; Gray); L. M. Umbach, 
woods, Clarke, Indiana, August 28, 1897 (Field) ; idem, eodem loco, 
August 20, 1898 (Field) ; G. R. Vasey, Lookout Mountain, Tennessee, 
1878 (Field) ;L.F. Ward, Potomac flats, Maryland, August 12, 1877 
(Field); Roscoe J. Webb, dry soil, Pippin Lake, Portage County, 
Ohio, September 6, 1909 (Gray) ; E. F. Williams, vacant lots, Boston, 
Massachusetts, September 3, 1903 (Gray); Wilbur H. Wright 221, 
Chicago, Illinois, August 22, 1908 (Field). 

A tall species of attractive appearance and wide distribution 
over the eastern and central United States. Commonly glabrous, 
but at times becoming more or less pubescent. Torrey and Gray 
(Fl. N. Amer. 2: 341. 1843) designated a pubescent form ("leaves 
minutely scabrous-puberulent. . . . Western Louisiana, Dr. Veatch") 
as "var. /3." During the four decades following, however, Asa 
Gray had many opportunities to see the inconstancy of the pubes- 
cence character in C. tripteris. Hence, in his Synoptical Flora of 
North America (1, pt. 2: 294. 1884) he modified the description of 
C. tripteris from "smooth and glabrous" to read "smooth and 
glabrous, or leaves minutely pubescent." Some years ago I myself, 
in a purely tentative way, treated the pubescent extremes as con- 
stituting a variety, which variety was set off with a proper name 
in my manuscript. With the examination of hundreds of herbarium 
and field specimens, however, it became apparent that the pubes- 
cence character was too capricious to admit of utilization, and that 
Gray's reduction of a variety thus segregated, to synonymy, would 
needs be accepted. Recently Standley (loc. cit.), apparently over- 
looking the treatments by Torrey and Gray and by Gray, has 



415 

described the var. Deamii to include forms with leaves pubescent at 
least underneath and involucres hairy; also the var. inter cedens to 
include forms with leaves pubescent at least underneath and 
involucres glabrous. 

Coreopsis tripteris var. Smithii Sherff, Bot. Gaz. 88: 301. 
1929. 

E specie foliis omnibus vel fere omnibus integris, lamina tenuiter 
oblongo-lanceolata differt. 

Type specimen: Collected by John Donnell Smith in low, open 
woods near Montgomery, Alabama, August 26, 1885 (Field). 

Distribution: Kentucky and Arkansas, southwardly to Florida, 
Georgia, and Mississippi. 

Specimens examined: Ex herb. A. W. Chapman, Georgia (Mo.); 
T. H. Kearney, Jr., along Clear Creek, Bell County, Kentucky, 
September, 1893 (Field); George V. Nash 2585, River Junction, 
Gadsden County, Florida, September 5, 1895 (U.S.);#. J. Palmer 
8468, moist, open ground, Malvern, Arkansas, September 4, 1915 
(Mo.); Charles Louis Pollard 1222, Waynesboro, Mississippi, August 
8-9, 1896 (Field) ; John Donnell Smith, low, open woods, etc. (type, 
Field; cotypes, Gray; Mo.). 

According to Smith's additional note on the type sheet, Asa Gray 
had pronounced this plant a new variety of Coreopsis tripteris and, 
in a letter written in November, 1885, had given it a varietal name. 
While Gray would probably have published the name had he not died 
shortly afterward, his proposed name was passed over by me in 
accordance with the recommendations of the Vienna Rules (Recomm. 
XIV. e) and the variety named in honor of the type collector. 

Coreopsis tripteris var. 7 subrhomboidea Sherff, Bot. Gaz. 
88: 303. 1929. 

Folia tripartita foliolis lateralibus lanceolatis terminali rhom- 
boideo-lanceolato 1.7-2.3 cm. lato, petiolo adjecto 6-8 cm. longa. 
Capitula minora, achaeniis tantum 4-4.5 mm. longis. 

Type specimen: Collected by Ernest Jesse Palmer, No. 29421, 
sandy, open woods bordering bog, near Texarkana, Bowie County, 
Texas, October 27, 1925 (Gray). 

Distribution: Known only from type locality in Bowie County, 
Texas. 

Specimens examined: Palmer 29421 (type, Gray). 



416 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

In the more than two hundred collections of the species proper 
studied by me, all the leaflets were variously linear, oblong-linear, 
or narrowly oblong-lanceolate, and the achenes were commonly 
5-6 mm. long. In the Palmer plant the terminal leaflets approach 
very distinctly a rhombus in outline and the achenes are smaller, 
measuring only about 4-4.5 mm. in length. 

92. Coreopsis latifolia Michx. Fl. Bor. Amer. 2: 137. 1803. 
Leiodon latifolium (Michx.) Shuttl. Distrib. Rugelii PI., Herb. Berol., 
etc. 

Herba perennis, erecta, gracilis, glabra vel pubescens, 1-1.5 
m. alta, summam versus erecte ramosa, foliosa usque vel fere usque 
ad summam. Folia opposita, subsessilia vel brevi-petiolata, omnino 
plerumque 1-2.3 dm. longa et 4-10 cm. lata, valde membranacea, 
ovata vel ovalia, basi late cuneata apice graciliter acuminata, mar- 
gine ciliata ac serrata dentibus mucronatis. Capitula pauca vel 
numerosa, paniculato-corymbosa, tenuiter pedicellata pedicellis 
2-10 cm. longis, radiata, pansa ad anthesin 4 cm. lata et 1.2 
cm. alta. Involucri bracteae exteriores 6-9, perspicue herbaceae, 
lineari-oblongae, apice obtusae, patentes, 6-8 mm. longae; interiores 
oblongae, apicem versus abrupte angustatae, paulo longiores. 
Flores ligulati 4 vel 5, flavidi, ligula elliptico-oblongi, apice integri 
vel paululum dentati, circ. 1.5-1.8 cm. longi. Paleae lineares, 
obtusae, 8 mm. longae. Flores tubulosi pauci (circ. 12), flavidi 
vel demum parce brunnei, stylorum ramis ad apicem conicis. Achae- 
nia obcompressa, oblongo-lanceolata, exalata, apice angusto truncata 
ac calva, circ. 7 mm. longa. 

Type specimen: Collected by Andre Michaux at higher altitude 
in the Carolina Mountains (Par.). 

Distribution: North Carolina southward to Georgia. 

Specimens examined: Anon., mountains of North Carolina, 
September, 1843 (Gray); Biltmore Herb. 5374b, slopes of Craggy 
Mountain, Buncombe County, North Carolina, August 12, 1897 
(Gray; N.Y.); Buckley, mountains of Carolina (Gray) and Georgia 
(N.Y.); herb. A. W. Chapman, mountains of Georgia (N.Y.); 
Asa Gray, Hickory Nut Gap, southeastern United States, 1843 
(N.Y.); idem & W. S. Sullivant, mountains of Carolina, 1843 (Gray); 
Harvard Univ. Bot. Gard., cult, anno 1849 e sem. e plantis in monti- 
bus Carolinae lectis (Gray); Hort. Berol., cult. (Berl.); F. Rugel, 
on summits of mountains, Broad River, North Carolina, July, 1841 
(Berl.; N.Y.; appellata Leiodon latifolium Shuttl.). 



417 

93. Coreopsis tinctoria Nutt., Journ. Acad. Phila. 2: 114. 1821. 
Calliopsis bicolor Reichenb. Mag. Aesthet. Bot. pi. 70. 1823. Dip- 
losastera tinctoria (Nutt.) Tausch, Hort. Canalius fasc. 1, icon, et 
descript. 1823. Calliopsis tinctoria (Nutt.) DC. Prodr. 5: 568. 1836. 
Coreopsis tinctoria var. /3 atropurpurea Hook. Bot. Mag. pi. 3511. 
1836. Coreopsis elegans Hort. fide Sieb. & Voss, Vilmor. Blumeng. 
ed. 3. 1. 487. 1894. 

Among the numerous illustrations of this widely cultivated 
species we may cite the following: Bart. Fl. Amer. Sept. pi. 45. 
1822; J. Radius, Schrift. Naturforsch. Gesellsch. Leipz. 1: pi. 4- 
1822; Bot. Mag. pi. 2512. 1824; Bot. Reg. pi. 846. 1824; Sweet, 
Brit. Flow. Gard. 1: pi. 72. 1824. 

Capitula pansa ad anthesin 2-3 cm. lata, achaeniis 1.2-4 mm. longis. 

C tinctoria sensu stricto. 

Capitula pansa ad anthesin 1-1.5 cm. lata, achaeniis 0.9-1.4 mm. 
longis var. ft imminuta. 

Herba annua, erecta, glabra, 6-12 dm. alta, caulibus subtetra- 
gonis vel subteretibus, foliosis, multo ramosis ramis plus minusve 
angulatis. Folia opposita, subsessilia vel breviter petiolata, saepius 
5-10 cm. longa, 1-2-pinnata vel summa indivisa, segmentis (vel 
laminis) anguste linearibus vel lineari-lanceolatis. Capitula nu- 
merosa, subcorymbosa, tenuiter pedunculata pedunculis (pedicellis) 
plerumque 4-10 cm. longis, radiata, pansa ad anthesin 2-3 cm. lata 
et 4-6 mm. alta. Involucri glabri bracteae exteriores circ. 8, plus 
minusve biseriales atque inter se imbricatae, lineari-oblongae vel 
saepius triangulatae, lateribus plerumque scariosae, 2 mm. longae; 
interiores deltoideo-ovatae vel oblongo-ovatae, circ. 5-6 mm. 
longae. Flores ligulati circ. 7 vel 8, inferne brunneo-rubri, alibi 
flavi, ligula obovati, apice plerumque 3-lobati, 0.7-1.5 cm. longi. 
Paleae subfiliformes, superne coloratae atque angustatae, apice 
acutae, demum circ. 4-4.5 mm. longae. Flores tubulosi atro-rubri, 
stylorum ramis apice obtusis. Achaenia lineari-oblonga vel deorsum 
paulo angustata, valde obcompressa, glabra vel glabrata, exalata, 
nigra, apice calva, nunc vix 1.2 nunc etiam 3-4 mm. longa. 

Type specimen: Collected by Thomas Nuttall, in flooded meadows 
on banks of Red River, Arkansas. Should be in London (Brit, or 
Kew). I have seen an authentic, fully labeled cotype (Berl.). 

Distribution: Minnesota, Saskatchewan, and Washington south- 
wardly to Louisiana, Texas, and California; cultivated everywhere 
for ornament and escaping frequently. Apparently becoming 
established at various places in China. 



418 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Specimens examined: Anon. 575, Fort Gibson, Arkansas, June, 
1835 (Berl.); Carrie Barker 31, low ground, Tonka wa, Oklahoma, 
June 27, 1908 (Field) ; J. M. Bates 2958, Red Cloud, Nebraska, July 
20, 1903 (Gray); Robert Bebb 1174, sandy soil, Galveston, Texas, 
May 18, 1903 (Field) ; H. C. Benke 327, Hutchinson, Kansas, October 
5, 1918 (Field); idem 4306, Claflin, Kansas, June 22, 1926 (Field); 
idem 4657, Murphysboro, Illinois, July 6, 1928 (Field); Katharine 
Brandegee, Redstone Park, California, July 25, 1905 (Calif.); N. L. 
Britton, Norfolk, Virginia, June 10, 1892 (Field); B. F. Bush 159, 
Sheffield, Missouri, June 27, 1895 (Berl.); idem 3017, introduced, 
Sugar Creek, Missouri, June 22, 1905 (Gray): George D. Butler 14, 
sulphate flats, fields, and roadsides, north of Limestone Gap, Okla- 
homa, June 18, 1877 (Field); William M. Canby 157, plains of 
Manitoba, Canada, August 7, 1897 (Gray); Harley P. Chandler 
7010, Rio Hondo, Texas, July, 1913 (Berl.; Gray); H. C. Cheo & 
W. F. Wilson 11 p.p., Mo Kan Shan, Province of Chekiang, China, 
June 19, 1926 (Gray); iidem 29, cult., eodem loco, June 28, 1926 
(Calif., 2 sheets; Gray); Mr. & Mrs. J. Clemens 978, Leon Springs, 
Texas, May 29, 1911 (Pom.); Fred Clements 2778, Spencer, Nebraska, 
July 25, 1893 (Gray); Royal A. Dixon 17, Huntsville, Texas, June 
3-12, 1908 (Field); R. R. Dreisbach 1042, escaped, ash dumps, 
Olney, Pennsylvania, July 15, 1922 (Field); Drummond, Texas 
(Gray); George Engelmann, cult., St. Louis, Missouri, September, 
1850, e sem. neo-mexicanis a Fendlero ad Santam Fe. lectis (Mo.); 
Henry Engelmann (Captain Simpson's Expedition), Fort Laramie, 
July, 1858 (Mo.); B. W. Everman, Lake Pend d'Oreille, Idaho, 
August 7, 1893 (Field) ; August Fendler 397 pro parte, prairie hollows 
near Pawnee Fork, New Mexico, September, 1847 (Mo.); Emily 
F. Fletcher, Westford, Massachusetts (Gray); Dr. Hale, Louisiana 
(Gray); Elihu Hall 344 pro parte, wet places, Hempstead, Texas, 
May 28, 1872 (Pom., 3 sheets); idem 345, eodem loco, June 3, 1872 
(Field) ;F.L. Harvey, bluffs, Little Rock, Arkansas (Mo.); Heiland, 
cult., Lychen, Province of Brandenburg, Germany, August 16, 1882 
(U.S.); A. Arthur Heller 11752, along railroad south of Redding, 
California, August 30, 1914 (Field; Gray); Albert S. Hitchcock 279 
and 666, buffalo wallows, Pawnee County, Kansas, August 19, 
1895 (Gray); idem 13943, savanna, Schofield Barracks, Leilihua, 
Island of Oahu, Hawaiian Islands, July 5, 1916 (U.S.); Hooker, 
Arkansas (Berl.); Hort. Paris., 1823 (Berl.); Marcus E. Jones 
29474, north of Corpus Christi, Texas, March 30, 1932 (Pom.); 
W. W. Jones 227, Middle Verde, Arizona, July 12, 1922 (Gray); 
J. B. Leiberg, moist places, Kootenai County, Idaho, June, 1890 



419 

(Field); idem, shores of Lake Pend d'Oreille, Idaho, July, 1892 
(Field) ; F.Lindheimer 102 p.p., prairies, Galveston Island, Texas, 1843 
(Berl.; Gray); idem 548, rocky places, New Braunfels, Texas, May, 
1851 (Berl., 2 sheets); idem 897, Texas, May, 1851 (Berl., forma 
multis achaeniis tuberculatis) ; John Macoun, Winnifred, Assiniboia, 
August 12, 1895 (Gray) ; idem 38, Red Deer Lakes, Canada, July 22, 
1879 (Gray); F. E. McDonald, along roadsides near East Peoria, 
Illinois, July, 1919 (Field); Albert Hanford Moore 1832, escaped, in 
dump by Lake Anthony, Cottage City, Massachusetts, July 16, 
1904 (Gray); Nealley, Chenates, western Texas, 1889 (Field); 
J. C. Nelson 1802, waste ground, Salem, Oregon, July 30, 1917 
(Gray); J. B. Norton 1455, cult., alt. 300-900 meters, Kuliang Hills, 
near Foochow, Province of Fukien, China, July-August, 1919 (U.S.); 
Thomas Nuttall, flooded meadows, etc. (cotype, Berl.);". J. Palmer, 
cherty barrens, Newton County, Missouri, July 15, 1906 (Gray); 
idem 2272, cherty barrens, Joplin, Missouri, June 19, 1909 (Gray) ; 
J. C. Parlin, North Berwick, Maine, July 4, 1892 (Gray); Dr. 
Pitcher, Arkansas (Gray); Birdie T. Powell, Hastings, Nebraska, 
July, 1886 (Field); J. T. Rothrock, cult., Philadelphia, Pennsylvania, 
June 25, 1877 (Field); idem 317, alt. 1,500 meters, Arizona, July, 
1874 (Field; Gray); T. E. Savage, J. E. Cameron, & F. E. Lenocker, 
Spokane, Washington, July, 1898 (Field) ; iidem, The Dalles, Oregon, 
August, 1898 (Field) ; A. K. Schindler 335, alt. 1,100 meters, Lu-shan, 
Kuling Mountains, Kiangsi, China, July-August, 1908 (Berl.); 
J. H. Schuette, cult., Green Bay, Wisconsin, September 14, 1888 and 
August 23, 1897 (Field); idem, cult., eodem loco, July 31, 1901 
(Gray); idem, cult., eodem loco, August 2, 1901 (Field; forma); 
E. P. Sheldon, Pike Island, Minnesota, July, 1895 (Pom.); Sherff 250 
pro parte, cult., St. Louis, Missouri, July 6, 1910 (Field) ; idem 499, 
cult., eodem loco, July 16, 1910 (Field); idem 507, cult., eodem loco 
et tempore (Field; Gray); idem 5000, cult., Chicago, Illinois, July 
3, 1929 (Berl.; Calif.; Field; ligulis supra brunneo-rubris infra plus 
minusve flavis); idem SOOOa, eodem loco et tempore (Berl.; Brit.; 
Calif.; Field; Kew; Mun.);5. Shimek, Oklahoma City, Oklahoma, 
July, 1892 (Field); G. W. Stevens 784, waste place along roadside, 
Cleo, Oklahoma, June 8, 1913 (Gray); idem 1212, waste place, 
Mountain Park, Oklahoma, June 23, 1913 (Gray); A. N. Steward 
1510, Kikungshan, Province of Honan, China, July 23, 1925 (Calif.); 
Tang Chung Chang & Nong Sing Po 3790, campus of Fukien Christian 
University, Foochow, Province of Fukien, China, June 8, 1926 
(Calif.); W. F. Thurrow, Hockley, Texas, 1890 (Field); To Rang 
Peng, Ts'ang Wai Tak, & Ts'ang Un Kin 936 (Canton Christian 



420 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

College No. 12935}, Shiu-chau, Province of Kwangtung, China, 
July 18, 1924 (Calif .) ; S. M. Tracy 7902, Weatherford, Texas, May 
26, 1902 (Berl.; Field; Gray);L. M. Umbach 2752, waste ground, 
Clyde, Illinois, July 14, 1900 (Gray); Van Hermann 2686, cult., 
vicinity of Santiago de las Vegas, Province of Havana, Cuba, May 
16, 1905 (Calif.; Field; Gray); L. F. Ward, Ford County, Kansas, 
October 5, 1897 (Gray); Mark White 218, Summer County, Kansas, 
June 17, 1899 (Mo.); Charles Wright 339 and 341, western Texas, 
May-October, 1849 (Gray); Cyril Zeller, Ellis County, Kansas, 
1908-1911 (Mo.). 

A species often confused with C. basalis var. Wrightii and with 
C. Atkinsoniana. From both it is at once distinguished by its almost 
oblong, entirely wingless achenes. In cultivation, plants occur with 
the rays reddish-brown or brownish-red almost or quite throughout. 
These are the var. atropurpurea of Hooker. At times their rays are 
distinctly yellow on the under surface and brownish-red above. 

Coreopsis bicolor Bosse ex Buchenau (Linnaea 25: 630. 1853, 
nomen) doubtless belongs with C. tinctoria. 

Coreopsis tinctoria var. ft imminuta Sherff, Bot. Gaz. 94: 
594. 1933. 

Capitula pansa ad anthesin 1-1.5 cm. lata, bracteis exterioribus 
1 mm. interioribus circ. 5-6 mm. longis, achaeniis 0.9-1.4 mm. 
longis. 

Type specimen: Collected by John Michael Holzinger, Santa Fe, 
New Mexico, August 13, 1911 (Mo.). 

Distribution: Known only from type locality of Santa Fe, New 
Mexico. 

Specimens examined: J. M. Holzinger, Santa Fe, New Mexico 
(type, Mo.). 

94. Coreopsis basalis (Dietr.) Blake, Proc. Amer. Acad. 51: 
525. 1916. Calliopsis basalis Dietr. in Otto & Dietr. Allgem. Garten- 
zeit. 3: 329. 1835. Coreopsis diversifolia Hook. Bot. Mag. pi. 3474. 
1836 (nee alior.). Calliopsis Drummondii D. Don in Sweet, Brit. 
Fl. Gard. ser. 2. 4: pi. 315. 1838. Coreopsis Drummondii (D. Don) 
Torr. & Gray, Fl. N. Amer. 2: 345. 1843. Coreopsis Drummondii 
var. ft Torr. & Gray, loc. cit. Coreopsis picta Hort. fide Sieb. & Voss, 
Vilmor. Blumeng. ed. 3. 1: 487. 1894. 

Foliorum segmenta linearia vel vix lanceolato-linearia. 

var. ft Wrightii. 



REVISION OF THE GENUS COREOPSIS 421 

Foliorum segmenta nunc lineari-lanceolata nunc elliptico-oblonga 
vel quidem orbiculata C. basalis sensu stricto. 

Herba annua, erecta, ramosa, nunc fere glabra nunc pilis pluri- 
articulatis valde hispida vel tomentosa, 2-4 dm. alta, caulibus 
ramisque angulatis ac sulcatis. Folia opposita, petiolata petiolis 
1-5 cm. longis, petiolo adjecto usque ad 12 cm. longa, principalia 
1-3-pinnata segmentis lineari-lanceolatis vel elliptico-oblongis vel 
quidem orbiculatis, membranaceis, undulato-integris, apice obtusis 
vel raro acutis. Capitula ramos (terminaliter per 5-15 cm. plerum- 
que nudos) terminantia, radiata, pansa ad anthesin 3-4.5 cm. lata 
et 8 mm. alta. Involucri bracteae exteriores 8-10, saepe patentes, 
subulatae vel lineari-lanceolatae, apice plerumque acutae, margini- 
bus plus minusve hispido-ciliatae, tergo longitudinaliter plerumque 
3-vittatae, 5-9 mm. longae, quam interiores ovatae glabrae paulo 
breviores. Flores ligulati circ. 8, magna ex parte flavi sed ad basim 
brunneo-rubri, ligula cuneate ac late obovati, 1.3-2.3 cm. longi, 
apice 3-lobati lobo mediano rursus 2- (raro 3-) lobulato. Paleae 
angustissime lineares, demum contortae et bracteas interiores sub- 
aequantes. Flores tubulosi saltern superne atro-rubri, stylorum 
ramis apice obtuso-conicis. Achaenia obovata, tergo convexo nigra 
ac valde papillata, marginibus incurvatis incrassata cartilagineaque, 
1.4-1.8 (-2) mm. longa, apice calva. 

Type specimen: Cultivated in Berlin Botanical Garden from 
seed said to have come from Missouri. If "Missouri" is correct, 
then the plants there must have been likewise in cultivation, since 
the species is known in the spontaneous state only in Texas. 

Distribution: Native to Texas. Sometimes cultivated for orna- 
ment and hence occasionally found elsewhere as an escape. 

Specimens examined: E. R. Bogusch 994, Gonzales County, 
Texas, June 6, 1926 (Pom.); Mrs. Charles C. Deam 1864, near Lake 
Helen, Florida, April 26, 1906 (Gray); T. Drummond 69 (commun. 
Hooker anno 1835), Rio Brasses (Brazos River), Texas (Gray); 
idem 200 (commun. Hooker anno 1835), Texas (Gray);#. H.Eames 
8839, waste ground, Fairfield, Connecticut, August 6, 1914 (Gray); 
Elihu Hall, Hempstead, Texas, June, 1872 (Field) ; idem 343, sandy 
prairie, eodem loco, May 2, 1872 (Field; Gray; Mo.; N.Y.; Pom., 2 
sheets); Harvard Univ. Bot. Gard., cult., 1844 (Gray); Hort. BeroL, 
July, 1837 (Berl., 2 sheets) ; J. F. Joor, open hills on Galveston Bay, 
Texas, June (Mo.) ; John H. Kellogg, cult, in Missouri Bot. Gard., 
June 30, 1905 (Mo.); F. Lindheimer, Galveston Island, Texas, 
November, 1842 (Mo.); idem, Texas, 1849 (Mo.); idem 54, Mill 



422 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Creek, Texas, April-May, 1844 (Mo.); idem 101, on dunes, Galves- 
ton Island, May, 1843 (Calif.; Field; Gray; Mo., 3 sheets; N.Y.); 
B. Mackensen 76, San Antonio, Texas, May 10, 1911 (Field) ; Manz 
29, Victoria, Texas (Berl.) ; ex herb. C. Mohr, damp meadows near 
coast, eastern shore of Mobile Bay, Alabama, April, 1868 (Field); 
E. J. Palmer 7740, nearly pure sand, Liberty, Texas, May 22, 1915 
(Mo.); J. Reverchon, White Creek, Brazos, Texas, July 4 (Berl.); 
J. Schneck, La Grange, Texas, May, 1894 (Field); J. H. Schuette, 
cult., Green Bay, Wisconsin, September 14, 1888 (Field); Earl E. 
Sherff 498, cult., St. Louis, Missouri, July 16, 1910 (Field); idem 
5015, cult., Harvard Univ. Bot. Gard., August 12, 1929 (Berl.; 
Brit.; Calif.; Field; Kew; Mun.); idem 5018, eodem loco et tempore 
(Field; Del.; Mo.; Mus.V.; U.S.); W. F. Thurrow, Hockley, Texas, 
1890 (Field); S. M. Tracy 8550, Cameron, Louisiana, July 6, 1903 
(Field; Gray; Mo.); idem 8925 p.p., Victoria, Texas, April 27, 1905 
(Berl.; Field; Gray; N.Y.); S. R. Warner, Willis, Texas, June 1 
(Mo.); Charles Wright, Texas (Gray). 

Coreopsis basalis var. ft Wrightii (A. Gray) Blake, Proc. 
Amer. Acad. 51: 526. 1916. C. Drummondii var. Wrightii A. Gray, 
Syn. Fl. N. Amer. 1, pt. 2: 291. 1884; cf. (typo) A. Gray, PL 
Wright. 2: 90. 1853; cf. etiam (descriptione sine typo vel nomine) 
A. Gray, PI. Wright. 1: 109. 1852. 

E specie tantummodo foliorum segmentis linearibus vel vix 
lanceolato-linearibus, capitulis paulo minoribus differt. 

Type specimen: Collected by Charles Wright, rocky hills on the 
San Pedro, western Texas, 1849 (Gray). Gray's name and descrip- 
tion (Syn. Fl. N. Amer. loc. cit.) rest upon the variety given in PI. 
Wright. 2: 90. 1853. The treatment in that work, however, gives 
first the reference to PI. Wright. 1: 109. 1852, concerning a plant 
collected by Wright in western Texas, in 1849; and, second, the 
citation of Wright 1237, collected on prairies of the Leona and Nueces, 
Texas, May, 1851. Fortunately the two collections were identical. 

Distribution: Widely distributed across Texas and reaching 
westwardly into New Mexico. 

Specimens examined: B. F. Biish 696, common in sand, Dallas, 
Texas, May 10, 1900 (Mo.) ; George L. Fisher 5007, Houston, Texas, 
May 13, 1917 (U.S.); 5. H. A.Groth 66, limestone hills near Bracken, 
Texas, July 8, 1903 (Field; Gray); A. A. Heller 1730, alt. 480-600 
meters, Kerrville, Texas, May 7-14, 1894 (Field; Gray; Mo.); 
Hort. Cantabr., cult, anno 1852 e sem. a C, Wrightio lectis (Gray) ; 






REVISION OF THE GENUS COREOPSIS 423 

G. Jermy 657, Iron Creek, Gillespie, Texas (Field; Mo.); Marcus 
E. Jones 28012, west of Uvalde, Texas, April 26, 1931 (Pom.); idem 
28013, east of Carisso Spring, Texas, April 27, 1931 (Pom.); E. J. 
Palmer 12115, rocky hills, San Marcos, Texas, July 1, 1917 (Mo.);' 
J. Reverchon, western Texas, 1882 (Gray) ; Miss Ellen D. Schulz 867, 
Austin, Texas, July, 1920 (U.S.); Charles Wright, Texas (Gray); 
idem 217, New Mexico, 1851 (Gray) ; idem 1237, prairies of the Leona 
and Nueces, western Texas, May, 1851 (Gray; Mo.; N.Y.; U.S.). 

95. Coreopsis rosea Nutt. Gen. N. Amer. PL 2: 179. 1818. 
Calliopsis rosea (Nutt.) Spreng. Syst. Veg. 3: 611. 1826. Conopsis 
rosea Nutt. ex Less. Syn. Gen. Compos. 228. 1832 (sphalm). Coreopsis 
rosea f. leucantha Fern. Rhodora 21: 171. 1919. 

Herba erecta, simplex vel ramosa, glabra, sub terra caulibus 
horizontalibus repens, caule supra terram tereti vel superne vix 
angulato 2-6 dm. alto. Folia opposita axillis saepe foliosis, basi 
ciliata parce connata, 2-5 cm. longa, integra vel irregulariter plus 
minusve 2- vel 3-partita, lamina vel segmentis linearibus, 0.5-3 mm. 
latis, eciliatis, apice parce acutis. Capitula ramulos tenues apicem 
versus nudos (pro pedunculis 3-8 cm. longis) aliter foliosos terminan- 
tia, radiata, pansa ad anthesin 2-2.5 cm. lata et 4-6 mm. alta. 
Involucri glabri bracteae exteriores 6-10, saepe irregulariter positae, 
mox patentes, oblonge lineari-lanceolatae, tantum 1-3 mm. longae; 
interiores oblongo-ovatae, basim versus connatae, 4-6 mm. longae. 
Flores ligulati circ. 8, rosei vel rarius albidi, ligula cuneato-oblongi, 
apice plerumque 3-lobati lobo mediano rursus lobulato, 8-13 mm. 
longi. Paleae lineares, 2-4 mm. longae. Flores tubulosi flavidi, 
stylorum ramis apice abrupte conico-incrassatis. Achaenia obcom- 
pressa, anguste elliptico-oblonga, nigra, exalata, glabra vel papillata, 
apice exaristata sed vix poculata, circ. 2 mm. longa. 

Type specimen: None cited, but "open grassy swamps, from New 
Jersey to Georgia" given as the habitat. 

Distribution: South westernmost Nova Scotia; Massachusetts to 
Delaware and (fide Nutt.) south to Georgia. 

Specimens examined : J. W. Adams 480, Hankens Pond, on Petti- 
coat Branch of Maurice River, Millville, New Jersey, August 15, 
1926 (Gray); anon., Smithfield, Rhode Island, 1871 (Gray); C. F. 
Batchelder, in water at marshy edge of pond, Falmouth, Massa- 
chusetts, August 14, 1906 (Gray) ; herb. William Boott, Winter Pond, 
Winchester, Massachusetts, July 30, 1876 (Gray) ; J. Bernard Brin- 
ton, low grounds, Egg Harbor, New Jersey, September 3, 1888 



424 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

(Field); idem, Egg Harbor, August 18, 1889 (Field); William M. 
Canby, Felton, Delaware, September, 1867 (Gray); idem, swamps, 
Delaware, August, 1877 (Field) ; J. Franklin Collins, Spectacle Pond, 
Lincoln, Rhode Island, September 11, 1900 (Gray); idem, M. L. 
Fernald, & H. H. York 294, wet sand, margin of Sand Pond, War- 
wick, Rhode Island, September 8, 1914 (Berl.; Field); Joseph W. 
Congdon, Lincoln, Rhode Island, August 22, 1871 (Field); herb. 
Carlton C. Curtiss, Martha's Vineyard, August 13, 1892 (Field); 
M. L. Fernald, E. B. Bartram, & B. Long 24687, peaty and cobbly 
beach of St. John (Wilson's) Lake, Yarmouth County, Nova Scotia, 
July 23, 1921 (Gray); iidem 24688, peaty margin of Goven Lake, 
Yarmouth County, Nova Scotia, July 23, 1921 (Gray); M. L. 
Fernald, C. H. Bissell, C. B. Graves, B. Long, & D. H. Linder 22859, 
gravelly margin, northwest side of Tusket (Vaughan) Lake, Nova 
Scotia, August 20, 1920 (Gray}; Fernald &Long 10664, damp, sandy 
shore of Loon Pond, Lakeville, Massachusetts, August 26, 1913 
(Gray); iidem 10669, damp, sandy borders of ponds west of White 
Pond, Chatham, Massachusetts, September 9, 1913 (Gray); iidem 
17591, forming a broad belt, wet, sandy lower beach and inundated 
margin, Buck Pond, Harwich, Massachusetts, August 30, 1918 
(Gray) ; iidem 1 7594, eodem loco et tempore (Gray, 2 sheets) ; iidem 
24689, wet, cobbly beach of Vaughan (Tusket) Lake, Nova Scotia, 
August 13, 1921 (Gray) ; iidem 24690, wet, peaty margin of Vaughan 
Lake at Gavelton, Nova Scotia, August 13, 1921 (Gray); iidem 
24691, peaty and cobbly margin of Gilfilling Lake, Yarmouth 
County, Nova Scotia, August 23, 1921 (Gray); iidem 24692, peaty 
and sandy margin of Salmon (Greenville) Lake, Yarmouth County, 
Nova Scotia, August 25, 1921 (Gray) ; iidem & D. H. Linder 22860, 
cobbly beach of Tusket (Vaughan) Lake, Gavelton, Nova Scotia, 
September 4, 1920 (Gray); iidem 22861, wet, peaty shore, East 
Branch of Tusket River, Gavelton, September 4, 1920 (Gray); 
iidem 22862, sandy and gravelly beach of Butler's (Gavelton) Lake, 
Gavelton, September 4, 1920 (Gray) ; Mrs. E. T. Gibbs, South Pond, 
Plymouth, Massachusetts (Gray); Jesse M. Greenman 437, Nine 
Mile Pcnd, Cape Cod, Massachusetts, September 5, 1898 (Gray); 
H. D. House 9660, Artist's Lake, Middle Island, Suffolk County, 
New York, August 14, 1923 (Gray) ; Edwin Hunt, Sudbury, Massa- 
chusetts, August 12, 1874 (Field); George G. Kennedy, Orleans, 
Massachusetts, July 31, 1891 (Gray); idem, Sharon, Massachusetts, 
September 2, 1904 (Gray); L. A. Kenoyer & F. W. Pennell 3224, 
margin of Mares Pond, Falmouth, Massachusetts, July 29, 1911 



REVISION OF THE GENUS COREOPSIS 425 

(Gray); Henry A. Lang, Egg Harbor, New Jersey, September 10, 
1905 (Gray) ; E. S. Miller, Long Pond, Wading River, New York, 
August 30, 1871 (Gray); idem, Wading River, August 18, 1877 
(Field, 2 sheets) ; ex herb. Thomas Morong, pond borders, Winchester, 
Massachusetts, August 5, 1881 (Field) ; John Murdoch, Jr., Orleans, 
Massachusetts, August 22, 1900 (Field) ; Joseph Murdoch, Rochester, 
Massachusetts, August 24, 1902 (Field); Oakes, in sandy, flooded 
places, Plymouth, Massachusetts (Gray); C. F. Parker, Bristol, 
Pennsylvania, August 11, 1865 (Field; Gray); herb. Margaret P. 
(Mrs. Robert Shaw) Russell, Snow's Pond, Rochester, Massachusetts, 
August 17, 1918 (Gray); F. C. Seymour 1384, in shallow water, 
Seth's Pond, West Tisbury, Martha's Vineyard, Massachusetts, 
September 7, 1917 (Gray); Harold St. John 2939, grassy shore of 
Long Pond, Southampton, New York, August 18-21, 1920 (Gray); 
G. Thurber, Cranston, Rhode Island, August, 1844 (Berl.) ; ex herb. 
E. Tuckermann, Jr., New England (Berl.); Emile F. Williams, Catau- 
met, Cape Cod, Massachusetts, September 15, 1901 (Gray). 

96. Coreopsis Cardaminefolia (DC.) Torr. & Gray, Fl. N. 
Amer. 2: 346. 1843. Calliopsis Cardaminefolia DC. Prodr. 5: 568. 
1836. Coreopsis Cardaminifolia DC. ex Nutt. Trans. Amer. Phil. 
Soc. n. ser. 7: 360. 1841 (sphalm). Coreopsis Cardaminefolia var. 
angustiloba Torr. & Gray, loc. cit. Coreopsis Cardaminefolia var. 
lineariloba A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 291. 1884. 

Herba annua, glabra, erecta, gracilis, suberecte ramosa, 2-5 dm. 
alta, caulibus subteretibus, foliosis. Folia opposita, remota, quam 
internodia breviora, petiolata petiolis plerumque 1-4 dm. longis 
(summa sessilia), inferiora medianaque laminis 1-2-pinnata segmentis 
elliptico-oblongis vel late angusteve linearibus, superiora nunc 
pinnatim paucilobata nunc simplicia. Capitula saepe numerosa, 
tenuiter pedicellata pedicellis glabris 1-5 cm. longis, radiata, pansa 
ad anthesin tantum circ. 1.6-2.4 cm. lata et 6-8 mm. alta. Involucri 
glaberrimi bracteae exteriores 6-9, plus minusve lanceolatae atque 
irregulariter positae, saepe subacutae, marginibus diaphanae, 1-2 
mm. longae; interiores ovatae 5-8 mm. longae. Flores ligulati circ. 
7 vel 8, ligula oblongo-obovati, guttur versus macula brunneo- 
purpurea ornati alibi flavi, apice 3-lobati, 1 cm. longi. Paleae 
filiformes, circ. 3-4 mm. longae. Flores tubulosi minuti, numerosi, 
corolla atro-purpurei, stylorum ramis apice obtusissime conicis. 
Achaenia elliptico-oblonga, valde obcompressa, faciebus glaberrima 
vel minute tuberculata, marginibus anguste vel late alata, circ. 2 mm. 
longa, apice calva vel minute bidentata. 



426 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Type specimen: The species was founded upon four specimens 
("pi. exs. 1668, 1956, 1478, et 2312") collected by Jean Luis Berlandier 
at various locations in southeasternmost Texas and adjacent Mexico 
(Del.). DeCandolle gave three localities: (1) between Bexar (Bejar) 
and Trinity River; (2) at Matamoros, State of Tamaulipas, Mexico; 
(3) at Lake St. Nicholas, Corpus Christi Bay. 

Distribution: Western Louisiana, Kansas, and Arizona south- 
ward to States of Tamaulipas, Coahuila, and Chihuahua, Mexico. 

Specimens examined : Berlandier 566, Texas, May 15, 1829 (Mo.); 
idem 882, about Matamoros, State of Tamaulipas, Mexico, April, 
1831 (Gray); idem 1073, from Matamoros northward to Nueces 
River, Texas, April, 1834 (Gray); idem 1956, Texas, May 15, 1829 
(Gray); idem 2312, about Matamoros, State of Tamaulipas, April, 
1831 (Gray) ; idem 2503, from Matamoros to Nueces River, Texas, 
April, 1834 (Gray; Mo.); idem 2580, Texas (Field); R. S. Cocks, 
Cameron, Louisiana, July, 1903 (Gray); J. F. Collins 8, wastes, 
Providence, Rhode Island, July 30, 1892 (Gray) ; Miss M. B. Croft, 
San Diego, Texas, 1885 (Field); Thomas Drummond 68, Brazos 
River, Texas (Gray); A. Fendler 897 pro parte, Pawneefork, New 
Mexico, August, 1847 (Mo.); idem 441, New Mexico, 1847 (Gray; 
Kew) ; J. Gregg 901 , prairie west of Matamoros, State of Tamaulipas, 
Mexico, June 5, 1847 (Mo.) ; Dr. Hale, Louisiana (Gray) ; Elihu Hall 
344 pro parte, wet places, Hempstead, Texas, May 28, 1872 (Field) ; 
Herbert C. Hanson A13, alt. 2,100 meters, wet places, Flagstaff, 
Arizona, August 22, 1922 (Field); Harvard Univ. Bot. Gard., cult, 
e sem. Texanis anno 1849 (Gray); W. W. Jones 160, Verde Valley, 
Arizona, September 9, 1920 (Gray);Lindheimer 102 p.p., Texas, 1843 
(Gray); D. T. MacDougal 293, alt. 2,100 meters, vicinity of Flag- 
staff, Arizona, July 15, 1898 (Field; Gray); E. W. Nelson 6104, 
near Colonia Garcia, in the Sierra Madre, State of Chihuahua, 
Mexico, August 25, 1899 (Gray); idem 6813, Sabinas, State of 
Coahuila, Mexico, May 21, 1902 (Gray) ; E. W. Olive 72, abundant 
in buffalo wallows, near Cinnamon City, Oklahoma, July, 1893 
(Field); J. H. Oyster 4014, Kansas, September 7, 1884 (Field); 
Edward Palmer 635, States of Coahuila and Nuevo Leon, Mexico, 
February-October, 1880 (Gray) ; idem 655, San Lorenzo de Laguna 
and vicinity, southwest of Parras, State of Coahuila, May 1-10, 
1880 (Field) ; C. G. Pringle 8328, alt. 210 meters, Rio Grande Valley 
near Diaz, State of Coahuila, April 25, 1900 (Field; Gray; Pom.); 
J. Reverchon 83, western Texas, 1882 (Gray); J. T. Rothrock 157, 
alt. 2,010 meters, Zuni Village, New Mexico, July, 1874 (Field; 



REVISION OF THE GENUS COREOPSIS 427 

Gray) ; B. C. Tharp 5627, causeway, Corpus Christi, Texas, March 
16, 1929 (U.S.); J. W. Tourney 594, Flagstaff, Arizona, June 30, 
1892 (Gray); Townsend & Barber 150, alt. 2,250 meters, near Colonia 
Garcia, in the Sierra Madre, State of Chihuahua, Mexico, July 19, 
1899 (Field; Gray); W. F. Thurrow, Hockley, Texas, 1890 (Field); 
C. Wright 34.0, western Texas, October, 1849 (Gray); idem 1236, 
New Mexico, 1851 (Gray) ; M. S. Young, muddy hollow in the plains, 
near Paloduro Canyon, Texas, September 7, 1917 (Mo.). 

97. Coreopsis stenophylla Boynt. Biltmore Bot. Studies 1: 
141. 1902. 

Herba glabra, annua vel vix perennis, 3-4 dm. alta, caulibus 
rigidis sed gracilibus, subfastigiate ramosis, omnino foliosis. Folia 
opposita, rigidula, erecta vel adscendentia, plerumque quam inter^ 
nodia longiora, simplicia vel irregulariter pinnata, lamina vel foliolis 
filiformibus vel anguste linearibus usque ad 2 mm. latis. Capitula 
subnumerosa, tenuissime pedicellata pedicellis 5 cm. longis, radiata, 
pansa ad anthesin 2-3 cm. lata et 6 mm. alta. Involucri bracteae 
exteriores circ. 8, lanceolatae, 1-2 mm. longae; interiores ovatae 
5-6 mm. longae. Flores ligulati plerumque 8, flavi, ligula obovati, 
apice 3-lobati lobis rotundatis, 1-1.4 cm. longi. Flores tubulosi 
atro-rubri, stylorum ramis apice plus minusve truncatis. Paleae 
lineares, acutae. Achaenia plana, elliptico-oblonga, 1.5-2 mm. 
longa, marginibus anguste alata alis integris, apice bidentata dentibus 
brevibus crassis. 

Type specimen: Collected for the Biltmore Herbarium, in pine 
woods near Hammond, Louisiana, September 17, 1900. The type, 
now probably destroyed, was in the Biltmore Herbarium. An 
excellent duplicate specimen is extant (N.Y.). 

Distribution: Louisiana. 

Specimens examined: Ex Biltmore Herb., Hammond, Louisiana, 
September 17, 1900 (N.Y., cotype); R. S. Cocks 3631, dry, open, 
sandy fields, Shreveport, Louisiana, July, 1907 (N.Y.); S. M. Tracy 
3456, Lake Charles, Louisiana, August 9, 1897 (N.Y.); ex herb. 
C. L. Willich & F. W. Weiss, loco ignoto (Berl.). 

98. Coreopsis similis Boynt. in Small, Fl. S. E. United States 
1278, 1340. 1903. 

Herba annua, glabra, erecta, 1-3 dm. alta, caulibus sulcatis 
striatisque, ramosis. Folia opposita, polymorpha, principalia tenuiter 
petiolata petiolis usque ad 7 cm. longis, petiolo adjecto 2-8 (-13) cm. 



428 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

longa, simplicia vel pinnatim partita, lamina vel foliolis oblonge 
linearibus vel elliptico-oblongis (raro oblongo-ovalibus), apice acutis 
vel obtusis, membranaceis, 5-12 mm. latis; summa simplicia vel 
pinnata, lamina vel foliolis plerumque angusto-linearibus. Capitula 
pauca vel numerosa, subcorymbosa, tenuissime pedicellata pedicellis 
5 cm. longis, radiata, pansa ad anthesin 1.5-2.5 cm. lata et 6-8 mm. 
alta. Involucri bracteae exteriores 7-9, lanceolatae vel interdum 
ovatae, glabratae, apice obtusae, 1.5-2 mm. longae; interiores ovatae, 
4-5 mm. longae. Flores ligulati plerumque 8, supra guttur purpureo- 
rubri alibi flavi, ligula obovati, apice 3-lobati lobis rotundatis, 1 cm. 
longi. Flores tubulosi numerosissimi, minuti, corolla atro-purpurei, 
stylorum ramis obtusissimo-apiculati. Achaenia plana, orbiculata, 
faciebus subnigris glabra, marginibus late alata unica ala diametro 
corpus (nunc 2 mm. longum et 0.5 mm. latum, nunc 3 mm. longum 
et 1 mm. latum) aequante, apice biaristata aristis tenuibus glabris 
externe ad alas plus minusve adgregatis tantum circ. 0.5 mm. longis. 

Type specimen: Collected by G. C. Nealley at Brazos Santiago, 
Texas, 1889 (U.S.). The year is erroneously given by Boynton 
as 1899. 

Distribution: Along the Gulf of Mexico coast, Texas. 

Specimens examined: W. L. Bray 48, Virginia Point, Texas, 
April 16, 1899 (U.S.); A. Arthur Heller 1548, sea level to 12 meters 
alt., Corpus Christi, Texas, April 9-12, 1894 (Gray; Mo.; N.Y.); 
G. C. Nealley, Brazos Santiago, etc. (type, U.S.; cotype, Field); 
Robert Runyon 385, Brownsville, Texas, January, 1923 (U.S.). 

99. Coreopsis Atkinsoniana Dougl. in Lindl. Bot. Reg. pi. 
1376. 1830. Calliopsis Atkinsoniana (Dougl. in Lindl.) Hook. Fl. 
Bor. Amer. 1:311. 1833. 

Herba annua vel plus minusve perennis, erecta, glabra, 6-12 
dm. alta, ramosa ramis arcuato-adscendentibus vel suberectis, 
caulibus angulatis, internodiis saepius elongatis. Folia radicalia 
l-3-(saepius 2-) pinnatisecta segmentis lineari-oblongis vel anguste 
elliptico-lanceolatis terminali multo majore petiolo 2-8 cm. longo, 
1-2 dm. longa; caulina plerumque 1-2-pinnatisecta segmentis 
anguste linearibus vel lineari-subspathulatis. Capitula subnume- 
rosa, tenuiter pedunculata pedunculis glabris plerumque 5-15 cm. 
longis, radiata, pansa ad anthesin 3-5 cm. lata et 7-10 mm. alta. 
Involucri bracteae exteriores 8-12, saepe irregulariter positae, 
lineari-oblongae, glabrae, margine scariosae, apice obtusae, 1-3 mm. 
longae; interiores ovatae, saepissime glaberrimae, 6-8 mm. longae. 



REVISION OF THE GENUS COREOPSIS 429 

Flores ligulati plerumque 8, guttur versus brunneo-purpurei alibi 
flavi, ligula cuneate obovati, apice 3-lobati lobo mediano rursus 
2-3-lobulato, 1.2-1.8 cm. longi. Paleae tenuiter oblongo-lineares, 
marginibus scariosae alibi aurantiaco-rubrae, 5 mm. longae. Flores 
tubulosi corolla purpureo-rubri, stylorum ramis apice abrupte sub- 
truncatis. Achaenia valde obcompressa, nigra, oblonga vel oblongo- 
oblanceolata, faciebus glabra vel minutissime sparsimque papillatal 
marginibus angustissime alata, 2.4-2.8 mm. longa, apice calva ve, 
vix bidentata. 

Type specimen: Collected by David Douglas on Mewries (or 
Menzies, cf. DC. Prodr. 5: 568. 1836) Island in the Columbia River 
in 1825. 

Distribution: Saskatchewan and British Columbia southwardly 
to Oregon, Arizona, and South Dakota. 

Specimens examined: L. R. Abrams 9465, Hood River, Hood 
River County, Oregon, July 25-28, 1922 (Pom.); Anon., cult, ex 
sem. ex herb. DeCand., anno 1839 (Gray) ; E. Bourgeau, rare, near 
lake on the prairie, Saskatchewan, September 17, 1857 (Kew); 
Townsend S. Brandegee, the Dalles, Oregon, 1882 (Calif.); Douglas, 
North America (Gray); idem, Fort Vancouver, British Columbia 
(Gray) ; Drake & Dickson, Bradfords Island, Columbia River, June, 
1889 (Field) ; A. D. E. Elmer 601, sand-gravel shores of Palmer Lake, 
northwest of Loomiston, Washington, August, 1897 (Berl.; Pom.); 
Griffiths, Highmore, South Dakota, July, 1897 (Berl.);Elihu Hall, 
Oregon (Field) ; idem, cult, from Dakota seed, 1877 (Gray) ; W. G. W. 
Harford & George W. Dunn, Cascades, northwestern United States, 
May 28, 1869 (Berl.); A. Arthur Heller, Coeur d'Alene River, near 
Harrison, Idaho, July 12, 1892 (Field); idem, about Lake Pend 
d'Oreille, near Lakeview, Idaho, August 1-10, 1892 (Pom.); Thomas 
J. Howell, Oregon, 1881 (Field) ; idem, Klickitat, Washington, June, 
1881 (Field); idem, banks of Columbia River, July, 1881 (Berl.); 
idem 224, Multnomah County, Oregon, July, 1877 (Gray); Marcus 
E. Jones, Sand Point, Idaho, September 18, 1908 (Pom.); Frank 0. 
Kreager 315, marshy shore of Lake Calispell, Stevens County, 
Washington, July 28, 1902 (Gray); John B. Leiberg 1542, alt. 640 
meters, region of Coeur d'Alene Mountains, Idaho, August 17, 
1895 (Field; Gray; Pom.); Lemmon 4159, meadow near San Fran- 
cisco Mountains, Arizona, September, 1884 (Gray); Dr. Lyall, 
Cascade Mountains to Fort Colville, lat. about 49 N., 1860 (Gray) ; 
idem, from Fort Colville to Rocky Mountains (Oregon Boundary), 
1861 (Berl.); Daniel T. MacDougal 617, Lakeview, Idaho, August, 



430 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

1892 (Field) ; Ernest MacKay 25, very abundant in wet sand, head 
of Grand Coulee, Douglas County, Washington, July 12, 1902 
(Gray; Pom.); J- M. Macoun, between Kettle and Columbia Rivers, 
Waneta, British Columbia, July 28, 1902 (Berl.; Field; Gray; Pom.); 
J. C. Nelson 1980, sandy shore of Columbia River, Hayden Island, 
Oregon, September 8, 1917 (Gray) ; Frank W. Peirson 3900, near 
Biggs, Oregon, July 16, 1923 (Pom.) ;C.G. Pringle, banks of Columbia 
River, Washington, September 10, 1881 (Field); Margaret P. (Mrs. 
Robert Shaw) Russell, Hood River, Oregon, August 15 (Gray); 
J. H. Sandberg, D. T. MacDougal, & A. A. Heller 617, on shores 
near Farmington Landing, Lake Coeur d'Alene, Idaho, July 11, 
1892 (Berl.; Field; Gray; Pom.); E. P. Sheldon 11081, Bonneville, 
Oregon, August 10, 1902 (Gray; Pom.); Sherff 250 pro parte, cult., 
St. Louis, Missouri, July 6, 1910 (Gray); Rev. Mr. Spalding, Clear 
Water, Oregon (Gray) ; W. N. Suksdorf, bottom lands of the Columbia 
River, Washington, August 26, 1882 (Berl.); G. R. Vasey 551, 
Washington, 1889 (Gray); Sereno Watson 217, Bed of Columbia 
River, Old Fort Colville, Washington, October 3, 1880 (Gray); 
idem (similiter) 217, mouth of the Chelan, Columbia River, Okan- 
ogen County, Washington, October 12, 1880 (Gray). 

100. Coreopsis Leavenworthii Torr. & Gray, Fl. N. Amer. 
2: 346. 1843. Coreopsis angustata Greene, Pittonia 4: 239. 1901. 

Folia simplicia var. d Lewtonii. 

Folia principalia 1-2-pinnata. 
Foliorum caulinorum foliola terminalia nunc obovata nunc lanceo- 

lato-oblonga, 0.8-1.5 cm. lata var. /3 Garberi. 

Foliorum caulinorum foliola terminalia (vel eorum segmenta 
ultima) circ. 1-3 mm. lata. 

Folia basalia pinnatim circ. 9-partita var. 7 Curtissii. 

Folia basalia integra C. Leavenworthii sensu stricto. 

Herba annua, erecta, glabra, 5-15 dm. alta, caulibus teretibus 
subsimplicibus vel suberecte ramosissimis, omnino vel fere usque ad 
summam foliosis. Folia opposita, ima integra elongate lineari- 
spathulata, quam internodia longiora, petiolis 1 dm. longis, 
laminis 5 cm. longis et saepius 3-5 mm. latis; principalia internodiis 
longiora, saepius 2-5-partita segmentis anguste linearibus plerumque 
1-3 mm. latis et petiolo elongate plus minusve similibus; summa 
integra, internodiis breviora. Capitula pauca vel numerosa, pa- 
niculato-corymbosa, tenuiter pedicellata, radiata, pansa ad anthesin 
2-3 cm. lata et circ. 5-6 mm. alta. Involucri bracteae exteriores 



REVISION OF THE GENUS COREOPSIS 431 

circ. 6-10, saepe irregulariter positae, lineari-lanceolatae vel ovato- 
lanceolatae, saepe coloratae, apice obtusae, margine plus minusve 
diaphanae, tergo plerumque scabridae, tantum 1-2 mm. longae; 
interiores ovatae, supra pubescentes, 5-7 mm. longae. Flores 
ligulati circ. 8, flavi, ligula obovati, apice 3-lobati lobis triangulato- 
rotundatis mediano majore saepe emarginato, 0.8-1.5 cm. longi. 
Paleae lineares vel subfiliformes, acutae, demum circ. 2-4 mm. 
longae. Flores tubulosi numerosi, minuti (circ. 2.2 mm. longi), 
stylorum ramis apice lato-conicis. Achaenia obcompressa, glabra, 
corpore nigro 2-3 mm. longo et circ. 0.5-0.7 mm. lato lineari-oblonga, 
marginibus membranaceo-alata ala subbrunnea corporis latitudinem 
aequante vel superante, apice biaristata aristis erectis tenuibus 
antrorsum hispidulis circ. 0.3-0.6 mm. longis. 

Type specimen: Collected by Melines C. Leavenworth, in Florida. 
The first cited specimen came from Tampa Bay (Gray). 

Distribution: Florida. 

Specimens examined: John H. Barnhart 2715, moist ground near 
Ehren, April 26-May 16, 1899 (Field) ;F. S. Blanton 6972, wet ditch, 
roadside, south of Lakeland, March 28, 1930 (Field); Mrs. E. G. 
Britton 132, prairie west of Cutter, March 23, 1904 (Field) ; Buckley, 
Florida (Gray); ex herb. William Canby, shores of Indian River, 
February, 1889 (Gray) ; A. H. Curtiss 5375, shore of Lake Worth, 
May 8, 1895 (Gray; Pom.); Mrs. Charles C. Deam 1709, border of 
Lake Worth, near West Palm Beach, February 22, 1904 (Gray); 
A. Fredholm 5^73, prairie, Orange County, August 8, 1902 (Gray) ; 
idem 5778, low prairie, Okeechobee region, Brevard County, April 
13, 1903 (Gray); idem 6094, swamp, Osceola County, October 12, 
1903 (Gray); A. P. Garber, Tampa Bay, 1876 (Gray); idem, Keys of 
Florida, 1876 (Gray); idem, Mellonville, March, 1876 (Field); idem, 
Lake Monroe, March, 1876 (Field); idem, Tampa, May, 1876 (Field); 
idem, Rosewood, June, 1876 (Field); W. Garvens, vicinity of Palm 
Beach, February 15-March 15, 1908 (Field); Albert S. Hitchcock, 
New River, December 26, 1895-January 11, 1896 (Field); idem, 
Palm Beach, eodem tempore (Field); idem, Levy County, June- 
July, 1898 (Field); William Kellogg, Owanita, about March 18, 
1907 (Gray) ; Dr. Leavenworth, Tampa Bay (type, Gray) ; George V. 
Nash 832, hummock land, vicinity of Eustis, May 16-31, 1894 (Berl. ; 
Field; Gray); idem 1257, eodem loco, July 1-15, 1894 (Berl.; Gray); 
idem 1876, Orange Bend, May 28-June 15, 1895 (Pom.); Edward 
Palmer 286, Indian River, 1874 (Gray);Fcmme R. Randolph 136, 
moist sand among sand dunes near Lake Worth, Kelsey City, 



432 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

January 2, 1921 (Gray; foliis omnibus simplicibus) ; F. Rugel 102, 
Florida (Field); idem 142, Florida, 1845 (Mo.); John K. Small & 
J. J. Carter, Bull Key, opposite Lemon City, November 6, 1903 
(Field); iidem 1219, Fort Lauderdale, November, 1903 (Field); 
John Donnell Smith, grassy pine barrens, Sumter County, March 
21, 1883 (Field); Miss Jeanette P. Standley 195, in pine land, vicinity 
of Fort Myers, May 20, 1916 (Field; Gray); F. C. Straub 97, Port 
Orange, April 9, 1895 (Gray); idem 159, eodem loco, May 18, 1895 
(Gray); S. M. Tracy 6920, Palma Sola, May 16, 1900 (Mo.); idem 
6921, eodem loco et tempore (Field, 2 sheets; N.Y.; U.S.; cotypes of 
Coreopsis angustata Greene); idem 7704, Braidentown, June 15, 
1901 (Berl.; Field; Gray); H. J. Webber, Sanibel Island, January 29, 
1896 (Field); idem 414, margin of Lake Worth, West Palm Beach, 
June 7, 1896 (Field). 

Coreopsis Leavenworthii var. /3 Garberi A. Gray, Syn. Fl. 
N. Amer. 1, pt. 2: 291. 1884. 

Folia caulina omnia pinnatim 5-7-partita foliolis brevioribus 
latioribusque, terminali subobtuso nunc obovato nunc lanceolato- 
oblongo, 0.8-1.5 cm. lato. 

Type specimen: Collected by Abraham Pascal Garber, Tampa, 
Florida, May, 1876 (Gray). 

Distribution: Known only from type locality of Tampa, Florida. 

Specimens examined: A. M. Ferguson, pine barrens, Tampa, 
June-July, 1898 (Mo.); A. P. Garber, Tampa, May, 1876 (type, 
Gray). 

Coreopsis Leavenworthii var. 7 Curtissii Sherff, Bot. Gaz. 
94: 592. 1933. 

Folia 1-2-pinnata; basalia pinnatim circ. 9-partita, foliolis 
oblongo-linearibus ad basim sensim angustatis plerumque 2-4 cm. 
longis et 2-5 mm. latis, lateralibus (praecipue inferioribus) utrinque 
saepe rursus 1-lobatis lobo parvo linearique; caulina quam internodia 
nunc multo longiora nunc multo breviora, plerumque 2-pinnata 
foliolis lateralibus saepius 3 vel 4 jugis, segmentis ultimis omnibus 
0.5-2.5 cm. longis et circ. 1-3 mm. latis. 

Type specimen: Collected by Allen Hiram Curtiss, No. 6734, in 
dry, pastured ground along the Suwannee River, near Branford, 
Florida, October 23, 1900 (Gray, 2 sheets). 

Distribution: Known only from type locality in Florida. 

Specimens examined: Curtiss 6734 (Gray, 2 type sheets). 



REVISION OF THE GENUS COREOPSIS 433 

Coreopsis Leavenworthii var. 8 Lewtonii (Small) Sherff, 
Bot. Gaz. 94: 592. 1933. C. Lewtonii Small, Bull. Torr. Bot. Club 
25: 146. 1898. 

E specie foliis simplicibus plerumque quam internodiis breviori- 
bus radice saepe perenni differt. 

Type specimen: Collected by Frederick Lewis Lewton, Forest 
City, Florida, 1894 (N.Y.). 

Distribution: Florida. 

Specimens examined : Liberty H. & Ethel Zoe Bailey 13047, sands 
at Sarasota, September 28, 1929 (Field); A. H'Curtiss 1480, swamps 
and shores, Halifax River, June (Berl.; Field, 2 sheets; Gray); A. A. 
Eaton 469, Little River, Bade County, December 5, 1903 (Field, 

2 sheets); Albert S. Hitchcock 139, around flatwood ponds, Myers, 
July-August, 1900 (Field) ; Frederick L. Lewton, Forest City, 1894 
(type, N.Y.); idem, Cedar Hammock, Sumter County, August 3, 
1894 (N.Y.); George V. Nash 2255, Sanford, July 24, 1895 (Berl.; 
Field; Gray); F. Rugel 133, Florida, 1843 (Gray; Mo., 2 sheets); 
J. K. Small & J. J. Carter 3025, the Everglades, intersecting Long 
Key, January 18-26, 1909 (N.Y.); John Donnell Smith, pine-barren 
ponds, Pellicier's Creek, St. John's County, March 4, 1882 (Field, 

3 sheets; Gray); S. M. Tracy 7256, Long Key, May 8, 1901 (Berl.; 
Field; Gray); idem 7355, Perico Island, December 6, 1901 
(Gray; Mo.). 

101. Coreopsis saxicola Alex. Torreya 32: 161. 1932. 

Herba perennis, erecta, glabra, suberecte ramosissima, saltern 
superne foliosissima, 7-10 dm. alta, caule subtereti ramis tetragonis. 
Folia opposita, principalia petiolata petiolis eciliatis inferne dilatatis 
basi connatis saepius 1-4.5 cm. longis, pinnata vel bipinnata seg- 
mentis oblongo-linearibus sparsim minuteque ciliatis 2-9 (rarius 
-13) mm. latis (raro multa simplicia spathulataque). Capitula 
corymbose disposita, tenuiter pedunculata pedunculis 1 dm. 
longis, radiata, pansa ad anthesin 2.5-4 cm. lata et circ. 1-1.2 cm. 
alta. Involucri bracteae exteriores plerumque 7 vel 8, lanceolatae, 
apice acutae vel subacutae ac saepe hispido-ciliatae alibi glabrae, 
marginibus albidae, 7-9 mm. longae; interiores oblongo-ovatae 
paulo longiores. Flores ligulati circ. 8, subaurantiaci, ligula oblongi 
vel cuneato-obovati, apice tenuiter lobati lobis plus minusve lobula- 
tis, 1.5 cm. longi. Paleae lineares, superne attenuatae atque 
elongatae, demum circ. 6-8 mm. longae. Florum tubulosorum styli 
ramorum terminis breviter conici. Achaenia obcompressa, valde 



434 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

incurva, circumambitu orbiculata vel obovato-orbiculata, atra, 
faciebus glabra vel papillata, circ. 2 mm. longa et (alls brunneis 
irregulariter pectinato-laceratis inclusis) circ. 2-2.5 mm. lata, apice 
demum calva. 

Type specimen: Collected by Francis W. Pennell, No. 4029, on 
thin soil over granite, Stone Mountain, DeKalb County, Georgia, 
August 2, 1912 (N.Y.). 

Distribution: DeKalb County, northern Georgia, westward into 
Tallapoosa County, Alabama. 

Specimens examined: F. S. Earle 2147, Double Bridges, Talla- 
poosa County, Alabama, August 9, 1899 (N.Y.); H. Eggert, Stone 
Mountain and vicinity, DeKalb County, Georgia, July 23, 1897 
(Mo., 2 sheets; N.Y.);F. W. Pennell 4029 (type, N.Y.); JoJm Kunkel 
Small, alt. 300-450 meters, on and about Stone Mountain, Georgia, 
August 1-6, 1895 (Field; N.Y.); Karl M. Wiegand & W. E. Manning 
S421, sandy soil, northwest base of Stone Mountain, Georgia, 
August 17, 1927 (Pom.; forma foliis plerumque simplicia); iidem 
8422, turfy rocky slope, base of Stone Mountain, Georgia, August 17, 
1927 (Pom.). 

An anomalous species, combining the foliar habit, involucre, and 
paleae of Eucoreopsis (especially C. grandiflora') with the lacerate- 
winged achenes of species like C. Linifolia. The tips of the style 
branches of the disc florets are somewhat intermediate, being less 
elongate-cuspidate than in the former but more definitely conical 
than in the latter. 

102. Coreopsis paludosa M. E. Jones, Contr. West. Bot. 12: 
46. 1908. 

Herba annua, erecta, glabra, subsucosa, simplex vel paniculato- 
ramosa, 6 dm. alta, internodiis elongatis usque ad 1.7 dm. longis. 
Folia alterna, nunc simplicia (pro ramis vel plantis parvis), anguste 
spathulata, 3.5-7 cm. longa et 2-5 mm. lata, lamina sensim in 
petiolum subaequalem decurrente, integra, apice subobtusa; nunc 
1-2-pinnata (pro plantis majoribus), majora (radicem versus posita) 
1.5-2.25 dm. longa petiolata petiolo 8 cm. longo foliolis lineari- 
spathulatis apice subobtusis (lateralibus circ. 3 jugis, 2-4 cm. longis 
et 1.5-3.5 mm. latis terminali circ. 6-8 cm. longo et 7-8 mm. lato) 
saepius (ad marginem ad folii terminum spectantem) 1-lobatis lobo 
8 mm. longo; altera minora, 1-pinnata, foliolis lateralibus 2 jugis. 
Capitula pauca (15), tenuiter pedicellata pedicellis usque ad 8 cm. 
longis, radiata, pansa ad anthesin 3.5 cm. lata et circ. 7-8 mm. 



REVISION OF THE GENUS COREOPSIS 435 

alta. Involucri glabri bracteae exteriores 4-6, oblongae, apice 
obtusae, plurinervatae, margine plus minusve hyalinae, adpressae 
vel rarius patentes, circ. 2 mm. longae; interiores ovatae, superne 
subelongatae, demum 6-8 mm. longae. Flores ligulati circ. 8, ligula 
rhomboideo-obovati, ad ligulae basim purpureo-maculati aliter flavi, 
supra medium 3-lobati lobis lateralibus parvis terminal! multo 
majore, 1.5-2 cm. longi. Paleae anguste lineares, glaberrimae, 4-5 
mm. longae. Florum tubulosorum stylorum rami termino truncati. 
Achaenia obcompressa, nigra, anguste obovata, basi flavo-callosa, 
faciebus glaberrima, marginibus pectinatim cartilagineo-dentata 
dentibus brunneis quam 0.1 achaenii latitudinis non longioribus, 
apice calva, 2.4-3.2 mm. longa et circ. 1.2-1.4 mm. lata. 

Type specimen: Collected by Marcus E. Jones at altitude of 
2,100 meters, growing in water in creeks, etc., Marsh Lake, Sierra 
Madre, State of Chihuahua, Mexico, September, 1903 (Pom.). 

Distribution: Known only from type locality in State of Chi- 
huahua, Mexico. 

Specimens examined: Marcus E. Jones, alt. 2,100 meters, Meadow 
Valley, Sierra Madre, State of Chihuahua, September 17, 1903 
(Pom.; an associate type). 

103. Coreopsis Linifolia Nutt. Journ. Acad. Phila. 7: 75. 
1834. Coreopsis callosa Bertol. Misc. Bot. 7: 42. 1848. 

Herba perennis, glabra, pallida, erecta, 5-7 dm. alta, caulibus 
gracilibus, supra ramosis, plus minusve tetragonis. Folia opposita 
vel inferiora alterna, integra, inferiora laminis spathulata vel oblanceo- 
lata quam internodia breviora (vel basalia interdum duplo longiora) 
4-9 mm. lata apice rotundata basi petiolata petiolis saepius 1-4 cm. 
longis; alia sessilia, linearia, apice saepe truncata vel obtusissima, 
multo minora. Capitula pauca, corymboso-paniculata, tenuiter 
pedunculata pedunculis saepius 3-7 cm. longis, radiata, pansa ad 
anthesin 2-3 cm. lata et circ. 6-7 mm. alta. Involucri bracteae 
exteriores circ. 6-8, glabratae, saepe irregulariter positae, ovatae, 
longitudinaliter pluristriatae, lateribus scariosae, apice subacutae 
vel rotundatae, 1-3 mm. longae; interiores oblongo-ovatae, saepius 
glabrae, 5-8 mm. longae. Flores ligulati circ. 7 vel 8, flavi, ligula 
obovati, apice 3-lobati lobo mediano emarginato, 1-1.5 cm. longi. 
Paleae lineares, apice acutae, circ. 5-6 mm. longae. Flores tubulosi 
corolla atro-purpurei, stylorum ramis apice obtuse conicis. Achaenia 
plana, oblonga vel ovalia, subnigra, faciebus glabra, marginibus 
ornata ala irregulariter lacerata circ. 0.2-0.3 corporis latitudinis 



436 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

lata, corpore circ. 3 mm. longa, apice tenuiter biaristata aristis 
sparsim antrorsumque hispidis 1 mm. longis. 

Type specimen: Collected in Alabama (Brit.?). 

Distribution: North Carolina to Florida, thence westwardly to 
Alabama and Mississippi. 

Specimens examined (formerly confused by me in herbaria with 
C. longifolia Small; for that reason there are cited here only the 
few specimens which have been recently reexamined) : A. W. Chap- 
man, wet barrens, Florida (Field); ex herb, eiusdem 174b, swamps 
in the pine barrens, Apalachicola, Florida, September 22, 1879 
(Gray; Mo.) ; A. B. Langlois, Spring Hill, southern Alabama, October 
1, 1880 (Field) ; J. Skehan, Ocean Springs, Mississippi, September 16, 
1895 (Mo.) ; John Donnell Smith, open tidal marshes on Biloxi Bayou, 
Harrison County, Mississippi, September 16, 1885 (Field; Gray); 
S. M. Tracy 4778, Koshtaw, Mississippi, October 25, 1898 (Field; 
Mo.); idem 8565, Gateswood, Alabama, October 31, 1903 (Field; 
Gray; Mo.). 

The type of Coreopsis callosa Bertol. has not been seen by me. 
Bertoloni's long description, however, fits C. Linifolia. His concluding 
observation ("Haec species appropinquaret Coreopsidem angusti- 
foliam Ait., si haberet omnia folia alterna, et lanceolato-linearia") 
shows beyond doubt that he had our C. Linifolia before him. More- 
over, Bertoloni's type had been collected by Dr. Gates in Alabama. 
A Gates specimen still extant (N.Y. ; not cited above) was undoubt- 
edly a duplicate. I found this latter to be typical C. Linifolia. 

104. Coreopsis longifolia Small, Bull. Torr. Bot. Club 22: 47. 
1895. C. angustifolia Dryand. in Ait. Hort. Kew. ed. 1. 3: 253. 
1789. (Non L. Sp. PI. 908. 1753.) 

Herba perennis, glabra, pallida, 7-12 dm. alta, erecta, caulibus 
simplicibus vel summam versus parce ramosis, teretibus. Folia 
alterna, integra, basalia (petiolis elongatis adjectis saepius 1-2 dm. 
longa) quam internodia multo longiora laminis linearia vel lineari- 
oblonga vel lineari-lanceolata 2-11 mm. lata apice plus minusve 
acuta marginibus saepe subscariosa; superiora sessilia, ad bracteas 
lineares plus minusve reducta. Capitula pauca paniculato-corym- 
bosa, tenuiter pedunculata pedunculis usque ad 2 dm. longis, radiata, 
pansa ad anthesin 3-5.5 cm. lata et 1 cm. alta. Involucri bracteae 
glabrae exteriores circ. 7-9, ovatae vel oblongo-lanceolatae, lateribus 
plus minusve diaphanae, apice saepe rotundo-dilatatae, 2-5 mm. 
longae; interiores oblongo-ovatae 6-12 mm. longae. Flores ligulati 
circ. 8, flavi, ligula cuneate obovati, apice 3-lobati lobo mediano 



REVISION OF THE GENUS COREOPSIS 437 

rotundato vel emarginato, 1.5-2.5 cm. longi. Paleae oblonge lineares, 
acutae, demum usque ad 1 cm. longae. Flores tubulosi corolla atro- 
purpurei, stylorum ramis apice obtusissime conicis. Achaenia plana, 
corpora ipso circ. 4 mm. longo atra et circumambitu (alis exclusis) 
oblonga vel late linearia, faciebus glabra margine pectinato-alata 
dentibus brunneis quam corporis diametro 0.3-0.6 brevioribus, 
apice breviter biaristata aristis antrorsum setulosis saepe caducis. 

Type specimen: Collected by Allen Hiram Curtiss, No. 4489, in 
dry, grassy pine woods about Jacksonville, Florida, October 16, 1893 
(N.Y.). 

Distribution: North Carolina, Georgia, and Florida. 

Specimens examined (perhaps several additional specimens 
determined by me some years ago for various herbaria as C. Linifolia 
Nutt. will be found to belong here): Curtis, North Carolina (N.Y.); 
A. H. Curtiss, Florida, November (Field) ; idem, near Jacksonville, 
Florida, September (Gray); idem 1470, margin of swamp near Jack- 
sonville, Florida, October (Pom.); idem 14.77 pro parte, moist pine 
barrens near Jacksonville, November (Field, 2 sheets; N.Y.); 
idem 4489 (N.Y., 2 type sheets) ; idem 5266, moist pine barrens near 
Jacksonville, October 15, 1894 (Field; Gray; N.Y., 2 sheets; U.S.); 
idem 5348 pro parte, thickets bordering swamps near Jacksonville, 
October 22, 1894 (N.Y.); idem 5568, moist, grassy pine barrens near 
Jacksonville, October 7, 1895 (Field; Gray; Mo., 2 sheets; Pom.); 
idem 6278, sides of ditch near Jacksonville, October 31, 1898 (Field; 
Gray); Mary E.Francis 155, low grounds, Alva, Florida, November 
25, 1917 (U.S.); A. P. Garber, Levy County, Florida, November, 
1877 (Field; Gray; U.S.); herb. Otto Katzenstein, southeastern 
United States, September 23, 1897 (Gray); L. H. Lighthipe 557 1 
South Jacksonville, Florida, October 30, 1896 (N.Y.); F. Rugel 480, 
Florida, 1845 (Field; U.S.); J. H. Simpson, Manatee, Florida, 1889 
(U.S.) ; J. K. Small & J. J. Carter 1028, pine lands, Fort Lauderdale, 
Florida, November 19-25, 1903 (Field); J. K. Small &G. V. Nask, 
the Everglades, west of Miami, Florida, November 1-9, 1901 (N.Y.); 
Mm Jeanette P. Standley 423, low pine land, vicinity of Fort Myers, 
Florida, October 25, 1916 (Pom.); S. M. Tracy 7142, Braidentown, 
Florida, October 16, 1900 (Field; Gray; Mo.; N.Y.; U.S.). 

Passes into C. gladiata. 

105. Coreopsis falcata Boynt. Biltmore Bot. Studies 1: 141. 
1902. 

Herba perennis, pallida, glabra, 8-12 dm. alta, caulibus teretibus, 
summam versus corymbose ramosis. Folia alterna, inferiora petio- 



438 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

lata petiolis 1-1.5 dm. longis lamina (aequaliter 1-1.5 dm. longa; 
0.6-3 cm. lata) elliptico-oblonga vel linearia vel lineari-oblanceolata 
terminaliter nunc obtusa nunc longe attenuata marginibus Integra 
vel saepe 1-2-lobata lobis angustis adscendentibusque; superiora 
sessilia, multo minora, angustiora, lateraliter saepe 1-plus-lobata. 
Capitula pauca vel vix subnumerosa, tenuiter pedunculata pedunculis 
1 dm. longa, radiata, pansa ad anthesin 3-5 cm. lata et 6-9 mm. 
alta. Involucri glabri bracteae exteriores circ. 8-10, lanceolatae, 
4-7 mm. longae; interiores ovatae vel oblongo-lanceolatae, dimidio 
longiores. Flores ligulati circ. 8, aurantiaco-flavi, ligula cuneato- 
obovati, apice 3-lobati lobo mediano majore rotundato vel emar- 
ginato, 1.3-2.3 cm. longi. Paleae lineares, acutae. Achaenia plana, 
oblonga, corpore circ. 4.5 mm. longa, marginibus alata alis latis 
pectinatis dentibus corporis diametrum fere aequantibus, apice 
biaristata aristis usque ad 1 mm. longis. 

Type specimen: Collected for the Biltmore Herbarium, in shallow 
water near Pembroke, North Carolina, June, 1901. The type proper 
was preserved in the Biltmore Herbarium, where it was doubtless 
destroyed later on by flood waters. A sheet of fine duplicate material 
is still extant, however (N.Y.). 

Distribution: North Carolina to Georgia. 

Specimens examined: Anon., Georgia (Gray); anon., Macon, 
Georgia (N.Y.) (these first two specimens are doubtless of the same 
collection; the first had been studied by Asa Gray and was regarded 
by him as C. gladiata in his Synoptical Flora of North America; the 
second had come from John Torrey's private herbarium and was 
under the name C. gladiata; this latter specimen was probably the 
basis for Torrey and Gray's words "sometimes 1-2-ternately-parted" 
in describing the lower leaves of C. gladiata [Fl. N. Amer. 2: 347. 
1843]); ex Biltmore Herb., shallow water near Pembroke, North 
Carolina, etc. (cotype, N.Y.); ex eodem 14?58b, shallow water or wet 
places in pine lands, Selma, North Carolina, May 31, 1902 (N.Y.); 
Cuthbert, in pool, Georgia, October 10, 1898 (N.Y.); Karl M. 
Wiegand & W. E. Manning 3410, dry pine woods, 7 miles north of 
Washington, North Carolina, June 21, 1927 (Pom.). 

106. Coreopsis gladiata Walt. Fl. Carol. 215. 1788. C. dicho- 
toma Michx. Fl. Bor. Amer. 2: 137. 1803. 

Herba perennis, glabra, pallida, erecta, 6-8 dm. alta, caulibus 
teretibus vel vix angulatis, supra ramosa et subnudata . Folia plerum- 
que alterna, integra, inferiora petiolata petiolis elongatis quam inter- 



REVISION OF THE GENUS COREOPSIS 439 

nodia plerumque multo longiora laminis elliptico-oblonga vel 
oblanceolata marginibus scariosa plerumque 1-2.5 cm. lata apice 
rotundata atque interdum aegre mucronata; superiora sessilia, multo 
minora, linearia vel spathulato-linearia. Capitula pauca, sub- 
corymbosa, tenuiter pedunculata pedunculis glabris 1 dm. longis, 
radiata, pansa ad anthesin 3-4 (-6) cm. lata et 8-11 mm. alta. 
Involucri glabri bracteae exteriores 4-10, saepe biseriatim dispositae, 
ovatae vel lanceolatae, plerumque obtusae, lateraliter scariosae, 
2-4 (-6) mm. longae; interiores oblongo-ovatae 8-12 mm. longae. 
Flores ligulati circ. 8, flavi, ligula cuneato-obovati, apice 3-lobati 
lobo mediano rotundato vel saepius emarginato, 1-2.5 cm. longi. 
Paleae lineares, apice acutae, demum 8 mm. longae. Flores 
tubulosi corolla atro-purpurei, stylorum ramis apice late breviterque 
conicis. Achaenia plana, elliptico-oblonga, faciebus nigris glabra, 
marginibus anguste pectinato-alata, corpore circ. 3.5 mm. longa, 
apice biaristata aristis tenuibus antrorsum hispidis. 

Type specimen: No specimen cited. The plant in mind was 
native to Carolina. My notes, contain no mention of an authentic 
(author's) specimen among the Walter specimens in London (Brit.). 

Distribution: North Carolina to southern Florida and west- 
wardly to southern Mississippi. 

Specimens examined: C. Billington 131, low places near salt water, 
Lynn Haven, Bay County, Florida, October 15, 1921 (U.S.); 
Biltmore Herb, (ex herb. A. W. Chapman) 20661), low pine barrens, 
Apalachicola, Florida, September 22, 1879 (Mo.; U.S.); J. Brown- 
field, Summerville, South Carolina, October, 1892 (Mo.); William 
M. Canby, vicinity of Wilmington, North Carolina, October, 1867 
(Field; Mo.); ex herb. A. W. Chapman, southern Florida (Field; 
U.S.); A. H. Curtiss, Chattahooche, Florida, 1875 (U.S.); idem 
1477 pro parte, moist pine barrens near Jacksonville, Florida, 
November (Gray; Mo.); Roland M. Harper 1666 p.p., wet barrens 
north of Moultrie, Georgia, September 24, 1902 (Berl.; Mo.; U.S.); 
Albert S. Hitchcock 910, New River, Florida, December 26, 1895- 
January 11, 1896 (Field) ; Gerald MacCarthy, eastern North Carolina, 
1883 (Field) ; Michaux, North America (Berl., sub nom. Coreopside 
dichotoma Michx.); Charles Mohr, margins of pine-barren swamps, 
Poplarville, Mississippi, October 18, 1894 (U.S.); John H. Redfield 
5630, near foot of Blue Ridge, Caesar's Head, South Carolina, 
September 3, 1876 (Mo.); F. Rugel, Florida, 1842-1849 (Mo.; 
foliorum apicibus C. longifoliae adpropinquans) ; John K. Small & 
J. J. Carter 694, in the Everglades, between Cocoanut Grove and 



440 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Cutler, Florida, October 31-November 4, 1903 (N.Y.); Udem 954, 
in pine lands, Fort Lauderdale, Florida, November 19-25, 1903 
(Field; N.Y.); iidem 2731, pine lands between Perrine and Larkin, 
Florida, November 16, 1906 (N.Y.); iidem & G. K. Small 3401, in 
the Everglades, Cutler to Homestead, Florida, February 15, 1911 
(N.Y.); J. T. Stewart, in swamp, near Charleston, South Carolina, 
1865 (Field); ex herb. Katherine A. Taylor, Summerville, South 
Carolina (Pom.). 

Passes into C. longifolia and apparently also into C. Linifolia. 

107. Coreopsis Helianthoides Beadle, Bot. Gaz. 25: 448. 1898 
(non C. Helianthoides Forst. Prodr. 91, num. 543. 1786; nomen 
subnudum). 

Herba perennis, erecta, pallida, 5-12 dm. alta, caulibus glabris, 
plus minusve teretibus, summam versus erecto-ramosis. Folia 
alterna, moderate adpresso-hispida setis pluriloculatis, marginibus 
scariosae, apice obtusae vel acutae, inferiora longe petiolata petiolis 
glabratis conduplicatis saepius 0.5-1.5 dm. longis, lamina (saepe 
rhomboide vel oblonge) ovata vel ovato-lanceolata 6-13 cm. longa 
et 2-6 cm. lata; superiora abrupte minora, sessilia, elliptico-oblonga 
vel linearia. Capitula pauca, corymbosa, tenuiter pedunculata 
pedunculis 1 dm. longis, radiata, pansa ad anthesin 3-4.5 cm. lata 
et 7-9 mm. alta. Involucri glabri bracteae exteriores 6-11, saepe 
subimbricatae, ovatae vel deltoideo-lanceolatae, tergo longitudinaliter 
pluristriatae, basi plus minusve subauriculatae, lateribus subdia- 
phanae, apice obtusae, plerumque 1-5 mm. longae; interiores ovatae 
vel interdum oblongo-lanceolatae, 8-12 mm. longae. Flores ligulati 
plerumque 8, flavi, ligula cuneato-obovati, apice grosse 3-lobati lobo 
mediano saepe emarginato, 2 cm. longi. Paleae tenuiter oblongo- 
lineares, 4-5 mm. longae. Flores tubulosi corolla atropurpurea, 
stylorum ramis apice obtuse conicis. Achaenia oblonge oblanceolata 
vel obovata, valde obcompressa, subnigra, faciebus glaberrima, 
marginibus subregulariter pectinato-alata dentibus brevibus longi- 
tudine circ. 0.1-0.2 corporis latitudinis aequantibus, circ. 3 mm. 
longa, apice brevissime biaristata. 

Type specimen: Collected by Alvin Wentworth Chapman, Aspa- 
laga, Florida, October, 1897. Herbarium not cited but presumably 
was the Biltmore Herbarium. The Chapman specimen No. 2066c 
in Herb. N. Y. had been distributed from the Biltmore Herbarium. 

Distribution: North Carolina southward to Florida and Alabama. 



REVISION OF THE GENUS COREOPSIS 441 

Specimens examined: Ex Biltmore Herb. 2139, Thomasville, 
Georgia, September 21, 1901 (N.Y.); William M. Canby, near 
Wilmington, North Carolina, October, 1867 (Field; Mo.; N.Y.); 
A. W. Chapman, Aspalaga, Florida, 1897 (U.S.); ex. herb, eiusdem 
2066b, low pine barrens, Apalachicola, Florida, September 22, 1879 
(Gray); (exBiltmore Herb.) ex eodem herb. 2066c, eodem loco, October, 

1897 (N.Y.; undoubtedly cotype material); A. H. Curtiss .1477 
pro parte, pine barrens, near Jacksonville, Florida, November 
(Berl.; Field; Mo.; U.S.); idem 4-487, near Jacksonville, Florida, 
October 21, 1893 (N.Y.; U.S.); idem 5348 pro parte, thickets 
bordering swamps, eodem loco, October 22, 1894 (Field; N.Y., 2 
sheets; U.S., 2 sheets); idem 10223, Chattahooche, Florida, October, 
1875 (Mo.); A. Cuthbert, peat bog, Augusta, Georgia, September 25, 

1898 (N.Y.); Roland M. Harper 653, alt. 90 meters, sandy bog 
just southeast of Americus, Georgia, September 15, 1900 (N.Y.); 
idem 1666 p.p., wet barrens north of Moultrie, Georgia, September 
24, 1902 (Gray; N.Y.) ; Charles Mohr, below lake, pine barren swamp, 
Yellow Pine, Alabama, August 8, 1895 (U.S.); J. B. Norton, swamp 
near mouth of Kilgore Branch, east of Hartsville, South Carolina, 
November 5, 1921 (U.S.); H. Ravenel, damp pine land, Santee 
Canal, South Carolina, September (Gray) ; F. Rugel 570 pro parte, 
Florida, 1845 (Field) ; John Donnell Smith, swampy woods, Columbia, 
South Carolina, September, 1883 (Field, 2 sheets); idem 2028, 
swampy woods 4 miles northeast of Columbia, September 25, 1883 
(Field; Gray; U.S., 2 sheets); Taylor, Thomasville, Georgia, Septem- 
ber 30, 1903 (Gray); Otto Vesterland, Marion County, Florida, 
November, 1889 (U.S.). 

108. Coreopsis integrifolia Poir. Encycl, Suppl. 2: 353. 1811. 

Herba perennis, erecta, gracilis, 6-9 dm. alta, caule glabro, 
tereti vel subangulato, subsimplici, fere omnino folioso. Folia 
opposita, petiolata petiolis hispido-ciliatis usque ad 3 cm. longis (vel 
summa sessilia); laminis diverse ovalia vel ovata vel suboblonga, 
crassiuscula, utrinque acuta vel subobtusa, marginibus venisque 
saltern primo hispida aliter glabra, 1-4 cm. longa, majora 1.5-2 cm. 
lata. Capitula pauca, tenuiter pedunculata pedunculis glabratis 
1 dm. longis, radiata, pansa ad anthesin 2.5-4 cm. lata et 9 mm. 
alta. Involucri bracteae exteriores 6, lineari-oblongae, ciliatae, 
obtusae, demum patentes vel reflexae, 2-4 (-6 ?) mm. longae; interi- 
ores ovatae 4-8 mm. longae. Flores ligulati plerumque 8, flavi, 
ligula cuneato-obovati, apice 3-lobati, 1-1.5 cm. longi. Paleae 
tenuiter lineari-oblongae. Florum tubulosorum stylorum rami 



442 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

apice obtusissime brevi-conici. Achaenia non visa. Ovaria plana, 
subalata, biaristata aristis nudis. 

Type specimen: Collected byBosc in Carolina (Par.?). 

Distribution: South Carolina and Georgia, also (fide F. E. 
Boynt. in Small Fl. S. E. United States 1279. 1903) in Florida. 

Specimens examined : Biltmore Herb. 14759a, low, flat land along 
St. Mary's River, Folkston, Georgia, August 27, 1904 (N.Y.); 
A. W. Chapman, banks of Spring-brook, Decatur County, Georgia 
(N.Y.); Ravenel, damp, rich soils, Santee Canal, South Carolina, 
September (Gray). In addition, I have seen the fragment (Gray) 
obtained by Asa Gray at the DeCandolle Herbarium, doubtless 
from a cotype specimen. 

A species very rare in herbaria and imperfectly known as to 
fruiting characters. Boynton (loc. cit.) classes it, in his key, with 
the species having achenial margins fimbriate or dissected-winged. 
His specific description (p. 1279) omits, however, all reference to 
achenes. 

109. Coreopsis nudata Nutt. Gen. N. Amer. PL 2: 180. 1818; 
Hook. f. Bot. Mag. pi. 6419. 1929. Calliopsis nudata (Nutt.) Spreng. 
Syst. Veg. 3: 611. 1826. Conopsis nudata Nutt. ex Less. Syn. Gen. 
Compos. 228. 1832 (sphalm). 

Herba perennis, glaberrima, 6-12 dm. alta, caulibus e radice fere 
tuberosa erectis, teretibus, flexuosis, summam versus corymbose 
pauciramosis, internodiis elongatis. Folia alterna, simplicia, anguste 
linearia, plus minusve teretia et juncoidea, basalia elongatissima 
saepe 3-4 dm. longa, mediana superioraque 1 dm. longa, summa 
subulata 1 cm. longa. Capitula pauca, radiata, longe tenuiterque 
pedunculata, pansa ad anthesin 4-6 cm. lata et circ. 7-8 mm. alta. 
Involucri bracteae exteriores 6-9, plus minusve irregulariter positae, 
lanceolatae, medio longitudinaliter vittatae, marginibus diaphanae, 
basi subauriculatae, tergo saepe subscabridae, 2-4 mm. longae; 
interiores ovatae, saepe subscabridae, 7-10 mm. longae. Flores 
ligulati circ. 8, rosei, obovato-cuneati, apice 3-lobati lobis lateralibus 
rotundatis mediano plerumque multo majore truncate vel emargi- 
nato, 1.5-2.2 cm. longi. Paleae late lineari-oblongae, apice subacutae, 
circ. 5-6 mm. longae. Flores tubulosi corolla flavidi, stylorum ramis 
obtuse truncato-conicis. Achaenia subplana oblonga, faciebus nigra, 
marginibus irregulariter pectinato-dentata dentibus saepe 2-4- 
congregatis subbrunneis plus minusve planis apice obtusissimis 
quam corporis diametro saltern dimidio brevioribus, corpore circ. 



REVISION OF THE GENUS COREOPSIS 443 

2.5-3.1 mm. longa et (dentibus lateralibus inclusis) 1-1.4 mm. lata, 
apice biaristata (aristis linearibus stramineis antrorsum hispidis 
0.6-1 mm. longis) atque inter aristas poculo minuto central! coronata. 

Type specimen: Collected by William Baldwin, near St. Mary's, 
"west Florida." 

Distribution: Southern Georgia and northernmost Florida. 

Specimens examined: Biltmore Herb. 2070b, bordering pond in 
pine barrens, near Apalachicola, Florida, April 14, 1894 (Gray); 
A. W. Chapman, Apalachicola, Florida (Gray; N.Y., 2 sheets); 
herb, eiusdem 2070b, bordering ponds in pine barrens, near Apa- 
lachicola, Florida, April 14, 1894 (Field; N.Y.; cf. Biltmore Herb. 
2070b); A. H. Curtiss 1484, pine barren swamps, near Jacksonville, 
Florida, May (Berl.; Field, 3 sheets; Gray) ; idem 4488, near Jackson- 
ville, April 29, 1893 (N.Y.); idem 4713, swampy places in pine 
barrens, near Jacksonville, April 21, 1894 (Field ; Gray) ; Asa Gray, 
Apalachicola, Florida, April, 1875 (Gray); Roland M. Harper 1001, 
alt. 70 meters, wet pine barrens near Collins, Georgia, July 4, 1901 
(Berl.; Gray; N.Y.); idem 2198, cypress pond near Douglas, Coffee 
County, Georgia, May 11, 1904 (Berl.; Gray; N.Y.); Albert S. 
Hitchcock, Jacksonville, Florida (Field); B. F. Saurman, wet pine 
barrens, Apalachicola, Florida (Field); John K. Small, John B. 
DeWinkeler, & Charles A. M osier 11252, Apalachicola prairies, 
Florida, April 25, 1924 (N.Y.); John Donnell Smith, near Starke, 
Florida, April 9, 1880 (Field) ; C. S. Williamson, Waycross, Georgia, 
April 15 (N.Y.). 

110. Coreopsis Mildbraedii Muschler, Wiss. Ergebn. Deutsch. 
Zentr. Afr.-Exped. 1907-1908, 2: 381. 1911. 

Herba perennis vel suffruticosa, erecta, 1-1.5 m. alta; cauli- 
bus ramisque subhirtis vel deinde mox glabratis glaberrimisve, 
striato-sulcatis, teretibus, plus minusve ochraceis. Folia opposita 
petiolata petiolis concavis basi subvaginatis et linea prominente ad 
medium adpresse retrorsumque hirsuta conjunctis usque ad 2 cm. 
longis, petiolo adjecto 8-17 cm. longa, semel bis vel terne pinnati- 
secta, segmentis principalibus circumambitu ovatis vel rhomboideo- 
lanceolatis, faciebus dense glanduloso-punctulatis sed glabratis, 
marginibus ciliatis ac perspicuissime dentatis (dentibus elongatis 
saepius lineari-lanceolatis apicaliter acerrimis saepe 1-1.5 cm. 
longis) apice acerrime elongato-acuminatis. Capitula subnumerosa, 
tenuiter pedicellata pedicellis (1-3-bracteatis vix ultra 0.75 mm. 
crassis, infra glabris supra hispidis) saepius 4-12 cm. longis, radiata, 



444 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

pansa ad anthesin 4.5 cm. lata et 11 mm. alta. Involucri bracteae 
exteriores (1-3 proximae vere pedicella genitae exclusae) circ. 6-9, 
virides, elongato-lineares, dorso glabratae, marginibus infra plerum- 
que glabratae supra spinuloso-ciliatae, apice induratae, saepius 
1.3-1.8 cm. longae et circ. 1.5 mm. latae; interiores ovatae, margi- 
naliter diaphano-flavae alibi intense purpurascenti-atrae, tergo 
pubescentes, apice pulverulentae, tantum 5-8 mm. longae. Flores 
ligulati verisimiliter 8-10, flavi, ligula elliptico-oblanceolati, apice 
minute denticulati, circ. 2 cm. longi et 4-6 mm. lati. Paleae oblongo- 
lineares, pluristriatae, apice saepe subabrupte acutae, 6-8 mm. 
longae. Disci florum stigmata terminaliter elongato-caudata. 
Achaenia obovato-oblonga vel lineari-oblonga, obcompressa, apicem 
versus rarius contracta, atra, unaquaque facierum circ. 8-sulcata et 
sparsim erecto-setosa, marginibus anguste alata alis brunneis 
antrorsum setosis, corpore ipso 4.5-6 mm. longa et 1.5-2 mm. lata, 
apice dense breviterque erecto-setoso biaristata aristis erectis 
tenuibus sursum hispidis 1-2.5 mm. longis. 

Type specimen: Collected by Mildbraed (Exped. d. Herzogs Ad. 
Friedr. z. Mecklenburg Afr. 1907-1908), No. 2539, growing 1-1.5 
meters high at altitude of 3,000 meters, at edge of mountain forest 
and near the Ericaceae, Butagu Valley, Ruwenzori West, Belgian 
Congo, middle of February, 1908 (Bert.). 

Distribution: Known only from type locality in Belgian Congo. 

Specimens examined: JeanLebrun 4507, alt. 2,475 meters, Valley 
of the Nyamuwamba, Ruwenzori, Belgian Congo, November, 1931 
(Mus. Cong., 2 sheets); Mildbraed 2539 (type, Berl.). 

The achene wings are narrow and thickish, offering an approach 
to Bidens. The species is at once distinguished from all other species 
in both Bidens and Coreopsis by its (sharply cut) foliage. The 
exceptionally elongate exterior involucral bracts are an additional 
aid in identification. Lebrun's label describes the plant as having 
a strong odor of tansy (Tanacetum vulgare L.). 

111. Coreopsis bracteosa Sherff, Bot. Gaz. 76: 88. 1923. 

Herba glabra, verisimiliter perennis et 7-10 dm. alta, caulis 
internodiis folia superantibus, mediis inferioribusque saltern 18-22 
cm. longis. Folia sessilia, indivisa, linearia, 6-20 cm. longa et 4-10 
mm. lata, in apicem acutum sensim angustata, sparsim dentata 
dentibus saepe ad spinas vel etiam ad pilos spinulosos flexiles reduc- 
tis, margine revoluta et interdum minutissime spinuloso-ciliata. 
Capitula pedunculata pedunculis maximam partem glabris usque ad 



REVISION OF THE GENUS COREOPSIS 445 

5 cm. longis, radiata, pansa ad anthesin 3.5-4.5 cm. lata et 6-9 mm. 
alta. Involucri bracteae exteriores numerosae 19-21, anguste 
lineares, margine spinuloso-ciliatae, aliter glabrae, apice acerrime 
apiculatae, 1-1.7 cm. longae, interioribus lanceolatis dorso hispidis 
(pilis inferne squamellato-dilatatis) plerumque dimidio longiores. 
Flores ligulati circ. 12, ligula elliptici, lutei, apice denticulati, cire. 
1.7-2 cm. longi et 5-7 mm. lati. Paleae oblongo-lanceolatae, apiee 
subito angustatae, costa mediana brunnea perspicua. Disci florum 
stigmata caudato-apiculata. Achaenia atra, linearia vel oblonga, 
valde obcompressa, plerumque alata, margine et supra ad faciem 
sursum hispida, corpore circ. 4-5 mm. longa et (alis inclusis) 1-1.5 
mm. lata, biaristata aristis antrorsum hispidis 0.6-3 mm. longis. 
Planta Bidenti chaetodontae Sherff affinis sed foliis indivisis non 
pinnatipartitis, involucri bracteis exterioribus 19-21 non tantum 12, 
achaeniis plerumque alatis non exalatis, etc. differt. 

Type specimen: Collected by Hans Meyer, No. 532, at altitude of 
1,200-1,400 meters, bush and tree-grass-steppe, Jhangiro and 
Karagwe, German East Africa, summer of 1911 (Berl.). 

Distribution: British and German East Africa. 

Specimens examined: Meyer 532 (type, Berl.); A. Whyte, on 
5th march from Nandi to Kagamagas, Tropical East Africa, 1898 

(Kew). 

112. Coreopsis Prestinaria Schz. Bip. in Walpers, Repert. 6: 
163. 1846. Prestinaria Bidentoides Schz. Bip. loc. cit. Verbesina 
veris A. Richard, Tent. Fl. Abyssin. 1: 407. 1847. C. Prestinaria f. 
typica Vatke and f. latisecta Vatke, Linnaea 39: 498. 1875. 

Herba annua, subglabra, erecta, 2-6 dm. alta, ramis tenuibus 
saepe subnumerosis. Folia opposita, inferiora (ac interdum mediana) 
tenuiter petiolata petiolis saepius glabratis usque ad 2 cm. longis, 
superiora (ac plerumque mediana) breviter alato-petiolata vel sub- 
sessilia, omnia 3-6 cm. longa, circumambitu triangulata vel ovato- 
triangulata, bipinnatisecta, lobis membranaceis linearibus vel 
oblongo-lanceolatis glabris vel aegre ciliatis apice calloso-mucronatis. 
Capitula tenuiter pedicellata pedicellis usque ad 6 cm. longis, sub- 
numerosa, radiata, pansa ad anthesin 2-3 cm. lata et circ. 8 mm. alta. 
Involucri glabri bracteae exteriores 8-10, uniseriales, lineares vel 
lineari-spathulatae, trinerviae, calloso-apiculatae, circ. 6 mm. 
longae; interiores oblongo-lanceolatae, badiae, margine diaphanae, 
saepe paulo longiores. Flores ligulati circ. 8, flavi, ligula lineari- 
elliptici, apice integri, circ. 1-1.4 cm. longi et 3-4 mm. lati. Paleae 



446 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

anguste oblongae, nitidae, paucistriatae, usque ad circ. 12 mm. longae 
et circ. 1.5 mm. latae. Disci florum stigmata tenuia terminaliter 
angustata. Achaenia obcompressa, corpore lineari-fusiformia vel 
lineari-clavata nigra faciebus adpresse erecteque papillato-setosa 
6-9 mm. longa et 0.9-1.1 mm. lata, lateraliter alata alis angustis 
stramineis vel subbadiis superne 0.3-0.5 mm. latis marginibus 
exterioribus erecto-ciliatis, apice erecto-setosa et biaristata aristis 
subulatis stramineis antrorsum hispidulis 4-6 mm. longis. 

Type specimen: Collected by Wilhelm Schimper, No. 62, near 
Djeladjeranne (Dscheladscheranne), Abyssinia, 1840 (Par.; nomen 
vulg., Gellgelle Maskal). 

Distribution: Abyssinia. 

Specimens examined: J. M. Hildebrandt 417, during rainy 
season, September, 1872 (Berl.); A. Petit, Abyssinia (Berl., 3 sheets; 
Par., 3 sheets); Quartin Dillon 612, Abyssinia, November, 1840 
(Par., 2 sheets); Schimper 62 (type, Par.; cotypes, Berl., 4 sheets; 
Gray; N.Y.; Par., 2 sheets); idem 429, mountains and valleys at 
1,860 meters alt., Gowwo Soria, Tigre, Abyssinia, September 26, 
1862 (Berl., 2 sheets; type collection of f. latisecta Vatke); H. Steud- 
ner 349, Keren, etc., Tigre, Abyssinia (Berl.; nom. tigrense, Ade"). 

Often confused with C. macrantha but easily distinguished by its 
annual habit, lower stature, smaller flowering heads, smooth inner 
involucral bracts, more narrowly winged achenes, etc. The f. 
latisecta Vatke has leaf divisions slightly wider than in the type speci- 
mens but apparently is only a robust state of the species. 

Vatke (loc. cit.) confused C. macrantha with this species, as is 
shown by his inclusion of Schimper 71 and 1423 under C. Prestinaria. 
Oliver and Hiern (in Oliv. Fl. Trop. Afr. 3: 387. 1877) stated incor- 
rectly that the aristae exceeded the achene in length. Seldom do 
they even equal it, although the coincidence of certain aristae with 
others underneath in a mature fruiting head often gives an exag- 
gerated effect to the aristal length. 

The native names recorded by Schimper and others for this and 
closely similar species are: Gellgelle Maskal (Schimp.; spelled 
Gueulguel-Makhul, A. Rich. loc. cit.); Ad (Steudner; spelled Adde", 
A. Rich. loc. cit.); Embobahgad (A. Rich. loc. cit.). 

113. Coreopsis macrantha Schz. Bip. in Walpers, Repert. 6: 
163. 1846. Verbesina macrantha (Schz. Bip.) A. Rich. Tent. Fl. Abys- 
sin. 1: 408. 1847. C. macroptera Schz. Bip. ex Schweinf. & Aschers. 
in Schweinf. Beitr. Fl. Aethiop. 284. 1867 (nomen subnudum); 



REVISION OF THE GENUS COREOPSIS 447 

cf. Pirotta, Fl. Delia Colonia Eritrea 1: 184. 1904. C. Prestinaria f.> 
elatior Vatke, Linnaea 39: 498. 1875 (excl. Schimperi plantam n. 22). 
Prestinaria macrantha Schz. Bip. ex 0. & H. in Oliver, Fl. Trop. Afr. 
3: 391. 1877. 

Herba perennis, adscendenti-erecta e radice lignescenti fibrosa 
multicauli, 3-15 dm. alta, caulibus subtetragonis vel subteretibus, 
glabris, internodiis quam foliis saepius multo longioribus. Folia 
opposita, plemmque breviter petiolata petiolis glabris late alato 
marginatis tantum 2-8 mm. longis, petiolo adjecto 3-8 cm. longa, 
bipinnatisecta, lobis linearibus glabris membranaceis, corpusculo 
subcorneo terminatis, saepius 1.5-4 mm. latis. Capitula tenuiter 
pedicellata pedicellis glaberrimis usque ad 18 cm. longis, radiata, 
pansa ad anthesin 3-4 (-5) cm. lata et 8-13 mm. alta. Involucri 
bracteae subaequilongae, exteriores 9-13 saepe subbiseriales, lanceo- 
lato-lineares vel rarius anguste lineares, 3- (vel sub 5-) nerviae, 
omnino virides vel marginibus angustis albescentes, glabrae, apice 
subabrupte acutae, circ. 8-13 (rarius -18) mm. longae; interiores 
oblongo-lanceolatae, coloratae, marginibus diaphanae, ad tergi 
costam medianam hispido-fimbriatae. Flores ligulati circ. 8-10 
(12 fide Schultzii), flavi, ligula ovato-oblongi, apice minute denti- 
culati, 1.7-2.5 cm. longi et 5-10 mm. lati. Paleae lineari-oblongae, 
glabrae, 1-1.4 cm. longae. Disci florum stigmata caudato-appendi- 
culati. Achaenia plana, corpore nigra anguste lineari-oblonga ad- 
presse erecto-setosa 6-8 (-13) mm. longa et 0.9-1.2 mm. lata; 
margine moderate vel latissime alata alis hispidis et densissime 
erecto-ciliatis supra interdum 2-4 mm. latis; apice setis numerosis 
albescentibus erectis nitidis elongatis ac 2 aristis erectis subulatis 
stramineis antrorsum hispidis 3-7 mm. longis coronata. 

Type specimen: Collected by Wilhelm Schimper (sect. 2, No. 1236, 
Enschedcap, Abyssinia (Par.). 

Distribution: Eritrea and Abyssinia. 

Specimens examined: Anon., cult, in Hort. Berol., 1860 (Berl.; 
Par.); Antinori, Province of Scioa (Shoa), Abyssinia, 1878 (Flor.); 
Emilio Chiovenda, District of Semien, Province of Amhara, Abys- 
sinia, December 3, 1909 (Flor.); idem 1820, District of Dembia 
(Dembea), Province of Amhara, September 3, 1909 (Flor.); idem 
2205, Gondar, Province of Amhara, September 25, 1909 (Flor.); 
Quartin Dillon, Abyssinia (Par.) ;Ellenbeck (Exped. Baron vonErlanger) 
1660, alt. 3,500 meters, Schoa, Province of Schoa (Shoa), Abyssinia, 
November 16, 1900 (Berl.); Adriano Fiori 1826, alt. 2,000 meters, 
Hamasen region, Eritrea, April 1, 1909 (Flor.) ; G. Negri 56, Province 



448 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

of Scioa, Abyssinia, 1909 (Flor.) ; idem 235 bis, eodem loco et tempore 
(Flor.); idem 618, alt. 2,650 meters, eodem loco (Flor.); 0. Neumann 
75, alt. 1,800-2,300 meters, between Kontschu and Tomata (Gudji), 
Abyssinia, December 24, 1900 (BerL; forma involucre exteriore C. 
Prestinariae adpropinquans) ; A. Pappi 1194, alt. 2,800-3,000 
meters, toward the Torrent of Arigot, Mount Mamahot, Soyra 
Mountains, August 23, 1902 (Flor.); idem 1303, alt. 2,600-2,800 
meters, Torrent of Arigot, Soyra Mountains, Ocule' Cusae, Eritrea, 
August 23, 1902 (Flor.); idem 2642, Eritrea, July 14-15, 1902 
(Flor.); A. Petit, Abyssinia (Berl., 2 sheets); V. Ragazzi, Antoto, 
Province of Scioa, October 2, 1887 (Flor.); idem, Let-Marefia, 
Province of Scioa, September 4, 1886 (Flor.); idem, eodem loco, 
September 6, 1886 (Flor.); idem 250, alt. 2,000 meters, Amasen 
Quazien west of Dorfu, Eritrea, September, 1892 (Flor.); Rohlfs 
& Sleeker, alt. 2,450 meters, Katz, Abyssinia, December 30, 1880 
(Berl., 2 sheets) ; Rilppell, between Temben and Siemen, Abyssinia, 
June or July, 1832 (Par.); W. Schimper (itin. abyssin. sect. 5. n. 22, 
edit. Buchinger, 1854), Abyssinia (Berl.; Del.; Kew; Par., 3 sheets; 
type material of Coreopsis macroptera Schz. Bip.); idem 4$, in fields, 
alt. 2,790 meters, Dibra Eski (Siemen), Abyssinia, November 6, 
1850 (Par., 2 sheets; "blooming in August and September"); idem 
71, alt. 1,890 meters, mountains and valleys, Mai Sigamo, Abyssinia, 
September 26, 1862 (Berl., 3 sheets); idem 85, Abyssinia (Par.); 
idem (itin. abyssin. sect. 5. n. 95, edit. Buchinger, 1854), Abyssinia 
(Par.); idem (itin. abyssin. sect. 5. n. 1005, edit. Buchinger, 1854), 
Province of Agow, Abyssinia, 1853 (Par., 3 sheets); idem 1236 (type, 
Par.; cotypes, Berl., 2 sheets; N.Y.; Par.); idem 1423, alt. 3,150 
meters, Mount Gunna, Abyssinia, December 18, 1863 (Berl. ; Kew) ; 
Hugh Scott, alt. 2,700-2,900 meters, Mount Zuguala, Abyssinia, 
October 21-27, 1926 (Kew) ; H. Steudner 253, Province of Amhara, 
May 20, 1862 (Berl.); A. Terracciano & A. Pappi 2474, alt. about 
2,000 meters, Alibaret-Sella Mogasas, Bogos, Eritrea, February 5, 
1893 (Flor.). 

Vatke (loc. cit.) appears to have completely ignored the perennial 
habit and proportionately shorter achaenial aristae of this species. 
He merged it (as f . elatior) with C. Prestinaria, a course not followed 
by Oliver and Hiern, or by Otto Hoffmann, or by other subsequent 
students of the group. 

C. macroptera was known to Oliver and Hiern (in Oliv. Fl. Trop. 
Afr. 3: 391. 1877) only through achenes, and they referred it inter- 
rogatively to C. macrantha Schz. Bip. Schultz Bipontinus' private 



REVISION OF THE GENUS COREOPSIS 449 

herbarium (Par.) contains several sheets of excellent material 
labeled C. macroptera in his own writing. These, also several found 
elsewhere under his determination C. macroptera, have thicker and 
taller stems, mostly different leaves, mostly larger capitula with the 
larger, exterior bracts mostly more numerous and with their margins 
more conspicuously whitish, etc. (Folia subsessilia, 4-8 cm. longa, 
nunc bipinnatisecta lobis linearibus acriter apiculatis 1.5-5 mm. 
latis, nunc anguste oblongo-lanceolata lamina ipsa 1-1.5 cm. lata 
marginibus dentibus vel lobis linearibus perspicuissime elongatis 
[etiam 4.5 cm. longis] munita.) But a study of numerous other 
specimens more recently collected, especially by various Italian 
collectors, shows all intergradations, unless as to foliage. (In C. 
macroptera some of the principal leaves appeared to have a petiole 
and midrib grossly expanded into an abnormal blade, this irregularly 
toothed with greatly elongate, linear lobes. The appearance was 
unnatural and apparently teratological. However, we may con- 
sider this character as of no important value, since C. Prestinaria is 
known to produce at times a precisely comparable leaf -form. More- 
over, it may be noted that some authors who have retained C. macrop- 
tera have overlooked even this one seeming distinction. Thus 
R. Pirotta [Fl. Delia Colonia Eritrea 1: 184. 1904] lists C. macroptera 
and cites for it Terracciano & Pappi 2474, a plant which is typical 
C. macrantha.) 

114. Coreopsis pachyloma 0. & H. in Oliver, Fl. Trop. Afr. 
3: 391. 1877. C. involucrata Schz. Bip. in Walpers, Repert. 6: 163. 
1846 (non Nutt. Journ. Acad. Phila. 7: 74. 1834). Verbesina involu- 
crata (Schz. Bip.) A. Rich. Tent. Fl. Abyssin. 1: 409. 1847. C. 
callosa Schz. Bip. in Schweinf. & Aschers. Enum. 284. 1867. 

Achaenia biaristata C. pachyloma sensu stricto. 

Achaenia exaristata var. /3 inanis. 

Herba annua, erecta vel adscendens, ramosa, 3-7 dm. alta; caule 
ramisque (siccis) viridi-stramineis, perspicue striatis, nunc glabratis 
nunc hispidis, nitidis. Folia opposita tenuiter petiolata petiolis ba- 
saliter connatis interdum hispidis usque ad 2 cm. longis, petiolo ad- 
jecto usque ad 6 cm. longa, circumambitu late oblonga vel deltoideo- 
ovata, bipinnatipartita, lobis valde membranaceis, nervis margini- 
busque subsparsim setosis, saepe 1 vel 2 dentibus acriter serratis, 
abrupte mucronulatis, saepius 2-4 mm. latis. Capitula subcorym- 
bosa, pedunculata pedunculis tenuibus hispidis 5-14 cm. longis, 
radiata, pansa ad anthesin circ. 2 cm. lata et 6-8 mm. alta. Involucri 
bracteae exteriores circ. 8, lineari-spathulatae, valde foliaceae, facie- 



450 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

bus glabrae vel raro subhispidae, marginibus hispido-ciliatae, apice 
acuto vel subobtuso flavido-albidae ac cartilagineo-induratae, 7-12 
mm. longae et superne 1-2.5 mm. latae: interiores ovato-oblongae 
dorso hispidae saepe paulo vel etiam fere dimidio breviores. Flores 
ligulati 6-8, flavi, ligula elliptico-oblanceolati, 6-12-striati, apice 
integri vel denticulati, 8-10 mm. longi et 3-5 mm. lati. Paleae 
subalbidae convexe conduplicatae, acutae, 7-9 mm. longae. Disci 
florum stigmata terminaliter caudata. Achaenia corpore ipso 
dorsaliter lineari-oblanceolata, moderate obcompressa, nigra (nitida), 
glabra vel supra erecto-setosa, 4.5-5.5 mm. longa et dorsaliter fere 
1 mm. lata, lateraliter alis stramineis crassis erecte hispidis margi- 
naliter rotundatis ventraliter fere 0.5 mm. latis bialata, apice biaristata 
aristis lanceolato-subulatis, hyalinis, infra valde membranaceis supra 
rigidis acerrimisque, marginibus minute erecto-ciliolatis, 4-5.5 mm. 
longis. 

Type specimen: Collected by Wilhelm Schimper in Abyssinia 
(Kew). 

Distribution: Abyssinia. 

Specimens examined: Anon., under bushes at Gaffat near Debra 
Tabor, May 10, 1862 (Berl.); Eduard Ruppell, between Temben and 
Siemen, June or July, 1832 (type of C. involucrata Schz. Bip., Par.); 
Schimper (itin. abyssin.) sect. 5. nos. 15 (Del.; Par.) and 933 (Par.). 

An annual with the foliar habit of Bidens pilosa var. bimucronata 
f. odorata (Cav.) Sherff. The achenial characters separate it at once 
from other species of Coreopsis. 

Coreopsis pachyloma var. ft inanis Sherff, Bot. Gaz. 90: 
386. 1930. 

A specie achaeniis etiam primo exaristatis differt. 

Type specimen: Collected by Wilhelm Schimper, No. 933 pro 
parte, Abyssinia, commun. 1853 (Par.). 

Distribution: Abyssinia. 

Specimen examined: Schimper 933 pro parte (type, Par.). 

PLANTAE EXCLUSAE 

Coreopsis abyssinica Schz. Bip. ex Walp. Repert. 6: 163. 1846= 
Bidens chaetodonta Sherff, Bot. Gaz. 90: 387. 1930. 

Coreopsis abyssinica var. bipinnato-partita Chiov. in Pirotta, Fl. 
Colon. Eritrea 185. l9Q4:=Bidens setigera (Schz. Bip.) Sherff, op. cit. 
90: 390. 1930. 



REVISION OF THE GENUS COREOPSIS 451 

Coreopsis abyssinica var. glabrior 0. & H. in Oliv. Fl. Trop. Afr. 
3: 389. l877=Bidens chaetodonta var. glabrior (0. & H. in Oliv.) 
Sherff, op. cit. 90: 388. 1930. 

Coreopsis abyssinica f. latisecta Vatke, Linnaea 39: 499. 1875 
(nom. subnudum)=5zdews articulata Sherff, op. cit. 94: 591. 1932. 

Coreopsis Acmella (L.) E. H. L. Krause, Beih. Bot. Centralbl. 
32, pt. 2: 340. l9U=Spilanthes Acmella (L.) Murr. Syst. Veg. ed. 
13. 610. 1774. 

Coreopsis Acmella var. uliginosa E. H. L. Krause, loc. cit.= 
Spilanthes uliginosa Swartz, Prodr. Veg. Ind. Occ. 110. 1788. 

Coreopsis alata Cav. Icon, et Descr. 3: 30, pi. 260. 1795= Ver- 
besina occidentalis (L.) Walt. Fl. Carol. 213. 1788. 

Coreopsis alba L. Sp. PI. 908. l753.=Bidens pilosa var. radiata L 
Dondiaefolia (Less.) Sherff, Bot. Gaz. 97: 607. 1936. 

Coreopsis alternifolia L. op. cit. 909. YI53=Actinomeris alterni- 
folia (L.) DC. Prodr. 5: 575. 1836. 

Coreopsis ambacensis Hiern, Cat. Welw. Afr. PL 1, pt. 3: 586. 
1898=Bidens ambacensis (Hiern) Sherff, Bot. Gaz. 59: 309. 1915. 

Coreopsis ambigua Nutt. Journ. Acad. Phila. 7: 75. lSB4=Bidens 
mitis (Michx.) Sherff, op. cit. 81: 43. 1926. 

Coreopsis amplexicaulis Cav. Descr. 226. l8Q2=Simsia foetida 
(Cav.) Blake, Proc. Amer. Acad. n. ser. 41: 385. 1913. 

Coreopsis Anagallis E. H. L. Krause, Beih. Bot. Centralbl. 32, 
pt. 2: 344. Vd\=Erihydra Anagallis Gardn. Lond. Journ. Bot. 7: 
409. 1848. 

Coreopsis Anthemoides DC. Prodr. 5: 573. 1836=Bidens Anthe- 
moides (DC.) Sherff, Bot. Gaz. 56: 493. 1913. 

Coreopsis arenicola S. L. Moore, Journ. Linn. Soc. 37: 170. 1905= 
Bidens musoziana Sherff, op. cit. 59: 309. 1915. 

Coreopsis arguta Pursh, Fl. Amer. Sept. 2: 567. 1814=Bidews 
mitis (Michx.) Sherff, op. cit. 81: 43. 1926. 

Coreopsis aristata Willd. Sp. PI. 3: 2253. 18Q4:=Bidens aristosa 
(Michx.) Britt. Bull. Torr. Bot. Club 20: 281. 1893. 

Coreopsis aristosa Michx. Fl. Bor. Amer. 2: 140. 18Q3=Bidens 
aristosa (Michx.) Britt. loc. cit. 

Coreopsis aristosa var. mutica A. Gray, Syn. Fl. N. Amer. 1, pt. 
2: 295. 1884:=Bidens aristosa var. mutica A. Gray ex Gattinger, Fl. 
Tenn. 172. 1901. 



452 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Coreopsis arizonica (A. Gray) O. Hoffm. in Engl. & Prantl, Nat. 
Pflanzenf. 4, pt. 5: 243. 189Q=Coreocarpus arizonica (A. Gray) 
Blake, Proc. Amer. Acad. n. ser. 49: 344. 1913. 

Coreopsis Artemisiaefolia Jacq. Icon. PI. Rar. 3: pi. 595. 1786- 
1793; Coll. Suppl. 155. 1796= Cosmos sulphureus Cav. Icon, et 
Descr. 1:56. pi. 79. 1791. 

Coreopsis Artemisifolia Sess & Moc. Fl. Nov. Hispan. ed. 2. 
137. 1893= Cosmos sulphureus Cav. loc. cit. 

Coreopsis asperata Hutch. & Dalz. Fl. West Trop. Afr. 2: 141-143. 
1931; cf. Sherff, Bot. Gaz. 93: 219. l932=Bidens asperata (Hutch. 
& Dalz.) Sherff, loc. cit. 93: 220. 1932. 

Coreopsis Aspilioides Bak. Kew Bull. 153. 1898=Bidens Aspilioides 
(Bak.) Sherff, op. cit. 94: 590. 1932. 

Coreopsis aurea Dryand. in Ait. Hort. Kew. ed. 1, 3: 252. 1789= 
Bidens aurea (Dryand. in Ait.) Sherff, op. cit. 59: 313. 1915. 

Coreopsis Bacana Heyne ex DC. Prodr. 5: 632. \836=Glossogyne 
pinnatifida (Buch.-Ham. in Wall.) DC. in Wight, Contr. 19. 1834. 

Coreopsis baccata L. PI. Surin. 14. 1775; Amoen. Acad. 8: 262. 
H85=Wulffia stenoglossa DC. 5: 563. 1836. 

Coreopsis Baumii 0. Hoffm. in H. Baum, Warb. Kunene-Zambesi 
Exped. 419. lW3=Bidens Baumii (0. Hoffm.) Sherff, Bot. Gaz. 59: 
309. 1915. 

Coreopsis Bidens L. Sp. PL 908. 1753=Bidens cernua L. op. cit. 832. 

Coreopsis Bidentoides Nutt. ex Torr. & Gray, Fl. N. Amer. 2: 
339. 1843=Bidens Bidentoides (Nutt.) Britt. Bull. Torr. Bot. Club 20: 
281. 1893. 

Coreopsis biternata Lour. Fl. Cochinch. ed. 1. 508. 179Q=Bidens 
biternata (Lour.) Merr. & Sherff ex Sherff, Bot. Gaz. 88: 293. 1929. 

Coreopsis Buchingeri Schz. Bip. in Schweinf. & Aschers. Enum. 
284, nomen=Bidens rotata Sherff, op. cit. 90: 391. 1930. 

Coreopsis Buchneri Klatt, Leopoldina 25: 107. 1889=Bidens 
Buchneri (Klatt) Sherff, op. cit. 76: 158. 1923. 

Coreopsis Buchneri Klatt, Ann. Naturh. Hofmus. Wien 7: 103. 
l892=Bidens grandis Sherff, op. cit. 59: 309. 1915. 

Coreopsis chrysantha Vatke, Linnaea 39: 499. 1875=Bidens 
Vatkei Sherff, op. cit. 90: 388. 1930. 

Coreopsis chrysantha var. simplicifolia Vatke, op. cit. 500= 
Bidens Dielsii Sherff, op. cit. 90: 388. 1930. 



453 

Coreopsis cordifolia (Schz. Bip.) Drake del Cast. 111. Fl. Ins. Mar. 
Pacif. 208. 1890; Fl. Polyn. Franc. 109. 1892=Bidens cordifolia 
Schz. Bip. Flora 39: 361. 1856. 

Coreopsis cordifolia Sess & Moc. PL Nov. Hispan. ed. 2. 137. 
1893= Dahlia cardiophylla Blake and Sherff, nom. nov. 

The type is still extant (Herb. Bot. Card. Madrid). Dr. Sidney 
F. Blake, Senior Botanist of the United States Bureau of Plant 
Industry, has independently examined it at my suggestion and writes: 
"I agree that it is undoubtedly a hitherto unrecognized species of 
Dahlia." 

Coreopsis coriacea 0. Hoffm. in Engl. Pflanzenw. Ost-Afr. C: 
414, 1899=Bidens coriacea (0. Hoffm.) Sherff, Bot. Gaz. 81: 52. 1926. 

Coreopsis corymbifolia Buch.-Ham. ex DC. Prodr. 5: 602. 1836= 
Bidens pilosa L. Sp. PI. 832. 1753. 

Coreopsis Cosmoides A. Gray, Proc. Amer. Acad. 5: 126. 1861= 
Bidens Cosmoides (A. Gray) Sherff, Bot. Gaz. 70: 98. 1922. 

Coreopsis crassifolia Sesse* & Moc. Fl. Mex. ed. 2. 194. 1894= 
Dahlia variabilis Desf. Cat. Hort. Par. ed. 3. 182. 1829. The SessS & 
Mocino type came from mountains in Temascaltepec, State of 
Mexico. Its leaves are simple (although the lower ones are described 
as ternate). The general appearance is thus quite different from 
that in Dahlia variabilis Desf. With that species, however, it is 
definitely seen to belong from a study of Pringle 3165. The plant of 
this number at St. Louis (Mo.) has closely matching foliage, but 
the plant at Chicago (Field) has the leaves mostly 5-partite, as 
very commonly found in D. variabilis. 

Coreopsis Crataegifolia 0. Hoffm. Bot. Jahrb. 30: 431. 1901= 
Bidens Crataegifolia (0. Hoffm.) Sherff, op. cit. 76: 158. 1923. 

Coreopsis cuspidata Bertol. Misc. Bot. 7: 44. 1848 (ex descript. et 
patri&)=Bidens mitis (Michx.) Sherff, op. cit. 81: 43. 1926. 

Coreopsis dichotoma Drake del Cast. 111. Fl. Ins. Mar. Pacif. 209. 
1890 (nee alior.)=Bidens Degeneri Sherff, op. cit. 85: 3. 1928. 

Coreopsis diffusa Jones, Extracts Contr. West. Bot. 18: 73. 1933= 
Chrysanthellum mexicanum Greenm. Proc. Amer. Acad. 39: 114. 
1903 (forma humilis sed non aliter atypica). 

Coreopsis discoidea Torr. & Gray, Fl. N. Amer. 2: 339. 1843= 
Bidens discoidea (Torr. & Gr.) Britt. Bull. Torr. Bot. Club 20: 281. 
1893. 



454 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Coreopsis dissecta (Benth.) Blake ex Ind. Kew. Suppl. 5: 66. 1921 
Coreocarpus dissectus (Benth.) Blake, Proc. Amer. Acad. n. ser. 49: 
344. 1913. 

Coreopsis Elliotii S. L. Moore, Journ. Linn. Soc. 35: 346. 1902= 
Bidens Elliotii (S. L. Moore) Sherff, Bot. Gaz. 59: 309. 1915. 

Coreopsis exaristata 0. Hoffm. in Engler, Pflanzenw. Ost-Afr. C: 
414. lS99=Bidens microcarpa Sherff, op. cit. 76: 84. 1923. 

Coreopsis exaristata var. gracilior 0. Hoffm. loc. cit.=Bidens 
gracilior (0. Hoffm.) Sherff, loc. cit. 1923. 

Coreopsis Ferulaefolia Jacq. Hort. Schoenbr. 3: 65. 1798=Bidens 
Ferulaefolia (Jacq.) DC. Prodr. 5: 603. 1836. 

Coreopsis filifolia Hook. Bot. Mag. pi. 3505. 1836=Thelesperma 
filifolium (Hook.) A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 301. 1884. 

Coreopsis Fischeri 0. Hoffm. in Engler, Pflanzenw. Ost-Afr. C : 
414. 18W=Bidens Fischeri (0. Hoffm.) Sherff, Bot. Gaz. 76: 158. 1923. 

Coreopsis flammula Banks ex Steud. Nomencl. ed. 1. 108. 1821= 
Bidens laevis (L.) Britton, Sterns & Poggenb. Prel. Cat. 29. 1888 
(fide syn. B. Chrysanthemoidis in Steud. loc. cit.). 

Coreopsis Foeniculacea Sesse" & Moc. ex DC. Prodr. 5: 603. 1836= 
Bidens Ferulaefolia var. Foeniculaefolia (DC.) Sherff, Bot. Gaz. 81: 
39. 1926. 

Coreopsis foetida Cav. Icon, et Descr. 1: 55, pi 77. l79l=Simsia 
foetida (Cav.) Blake, Proc. Amer. Acad. n. ser. 41: 385. 1913. 

Coreopsis formosa Bonato Pisaura autom. Coreopsis formosa 22, 
pi. 2. 1793=Cosraos bipinnatus Cav. Icon, et Descr. 1: 10, pi. 14- 1791. 

Coreopsis frondosa 0. Hoffm. in Engler, Pflanzenw. Ost-Afr. C: 
414. l899=Bidens magnifolia Sherff, Bot. Gaz. 90: 390. 1930. 

Coreopsis fruticosa Forst. Fl. Ins. Austr. Prodr. 91. 1786, nomen= 
Bidens australis Spreng. Syst. Veg. 3: 453. 1826. 

Coreopsis Georgina Cass. Diet. Sci. Nat. 18: 441. 1820= Dahlia 
variabilis (Willd.) Desf. Cat. Hort. Par. ed. 3. 182. 1829. 

Coreopsis Georgina var. nuda Cass. op. cit. 442= Dahlia variabilis 
(Willd.) Desf. loc. cit. 

Coreopsis Georgina var. pruinosa Cass. op. cit. 443= Dahlia vari- 
abilis (Willd.) Desf. loc. cit. 

Coreopsis glaucescens O. & H. in Oliv. Fl. Trop. Afr. 3: 389. 1877= 
Bidens articulata Sherff, Bot. Gaz. 94: 591. 1932. 

Coreopsis Grantii Oliv. Trans. Linn. Soc. 29: 98. pi 65. 1874= 
Bidens Grantii (Oliv.) Sherff, op. cit. 59: 309. 1915. 



REVISION OF THE GENUS COREOPSIS 455 

Coreopsis hawaiensis (A. Gray) Drake del Cast. 111. Fl. Ins. Mar. 
Pacif. 209. 189Q=Bidens hawaiensis A. Gray, Proc. Amer. Acad. 5: 
128. 1861. 

Coreopsis heterophylla Cav. Icon, et Descr. 3: 34, pi. 268. 1795; 
Descr. 226. l8Q2=Iostephane heterophylla (Cav.) Benth. in Benth. 
& Hook. Gen. PL 2: 368. 1873. 

Coreopsis heterophylla Bertol. Misc. Bot. 7: 43. 1848 (ex descript. 
et patria)=M2ens mitis (Michx.) Sherff, Bot. Gaz. 81: 43. 1926. 

Coreopsis Hillebrandiana Drake del Cast. 111. FL Ins. Mar. Pacif. 
209. l89Q=Bidens Hillebrandiana (Drake del Cast.) Deg. ex Sherff, 
op. cit. 85: 6. 1928. 

Coreopsis hirta Raf. Ann. Nat. 15. 1820= Rudbeckia triloba L. 
Sp. PL 907. 1753 (fide Torr. & Gray, Fl. N. Amer. 2: 349. 1843). 

Coreopsis Hoktii 0. Hoffm. in Engler, Pflanzenw. Ost-Afr. C: 
415. 18W=Bidens Holstii (0. Hoffm.) Sherff, Bot. Gaz. 76: 79. 1923. 

Coreopsis incisa Ker, Bot. Reg. 1: 7 (pi.}. l8l5=Bidens incisa 
(Ker) G. Don in Sweet, Hort. Brit. ed. 3. 360. 1839. 

Coreopsis incurva Moench, Meth. Suppl. 245. lSQ2=Bidens 
Ferulaefolia (Jacq.) DC. Prodr. 5: 603. 1836 (fide auctorum). 

Coreopsis insecta S. L. Moore, Journ. Bot. 46: 42. 19Q8=Bidens 
insecta (S. L. Moore) Sherff, Bot. Gaz. 59: 309. 1915. 

Coreopsis involucrata Nutt. Journ. Acad. Phila. 7: 74. 1834= 
Bidens polylepis Blake, Proc. Biol. Soc. Wash. 35: 78. 1922. 

Coreopsis Jardinii (Schz. Bip.) Drake del Cast. 111. Fl. Ins. Mar. 
Pacif. 209. 1890; FL Polyn. Franc. 109. l892=Bidens Jardinii Schz. 
Bip. Flora 39: 360. 1856. 

Coreopsis Jasminifolia Bertol. Misc. Bot. 7: 44. l848=Bidens 
mitis (Michx.) Sherff, Bot. Gaz. 81: 43. 1926. 

Coreopsis kilimandscharica 0. Hoffm. Bot. Jahrb. 20: 234. 1894= 
Bidens kilimandscharica (O. Hoffm.) Sherff, op. cit. 59: 309. 1915. 

Coreopsis Kirkii 0. & H. in Oliv. FL Trop. Afr. 3: 390. 1877= 
Bidens Kirkii (0. & H. in Oliv.) Sherff, op. cit. 59: 309. 1915. 

Coreopsis leucantha L. Sp. PL ed. 2. 1282. H63=Bidens pilosa var. 
radiata Schz. Bip. in Bark.-Webb & Berth. Hist. Canar. 3, 2, part 2: 
242. 1836-1850. 

Coreopsis leucanthema L. Cent. PL 1: 29. 1755; Amoen. Acad. 4: 
291. l759=Bidens pilosa var. radiata Schz. Bip. loc. cit. 

Coreopsis leucorhiza Lour. FL Cochinch. ed. 2. 622. 1793 (ex 
descript. et patr\B.}=Bidens pilosa var. minor (Bl.) Sherff, Bot. Gaz. 
80: 387. 1925. 



456 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Coreopsis leucorrhiza Lour. op. cit. ed. 1. 508. 1790 (ex descript. et 
pa.tria)=Bidens pilosa var. minor (Bl.) Sherff, loc. cit. 

Coreopsis Liebmannii Schz. Bip. ex Hemsl. Biol. Centr. Amer. 
Bot. 2: 196. l88l=Verbesina intermissa Blake, Contr. U. S. Nat. 
Herb. 23: 1578. 1926. 

Coreopsis limensis Jacq. Coll. 2: 299. 1788=Encelia canescens 
Lam. Encycl. Meth. 2: 356. 1786 (fide Blake, Proc. Amer. Acad. n. 
ser. 41: 369. 1913). 

Coreopsis linearifolia DC. Prodr. 5: 570. 1836=Bidens angustis- 
sima HBK. Nov. Gen. et Sp. 4: 183 (233). 1820. 

Coreopsis linearifolia 0. & H. in Oliv. Fl. Trop. Afr. 3: 390. 1877= 
Bidens Schweinfurthii Sherff, Bot. Gaz. 59: 309. 1915. 

Coreopsis lineata Klatt, Ann. Naturh. Hofmus. Wien 7: 103. 1892 
=Bidens grandis Sherff, loc. cit. 

Coreopsis lucida Cav. Descr. 224. \8Q2=Bidens aurea (Dryand. in 
Ait.) Sherff, op. cit. 59: 313. 1915. 

Coreopsis Macraei A. Gray, Proc. Amer. Acad. 5: 126. 1861= 
Bidens Campylotheca Schz. Bip. Flora 39: 359. 1856. 

Coreopsis macrocarpa A. Gray, loc. cit.=Bidens macrocarpa (A. 
Gray) Sherff, Bot. Gaz. 70: 97. 1920. 

Coreopsis Menziesii A. Gray, op. cit. 127=Bidens Menziesii (A. 
Gray) Sherff, op. cit. 70: 98. 1920. 

Coreopsis micrantha (Gaud.) A. Gray, loc. cit. Bidens micrantha 
Gaud. Voy. Freycinet Bot. PI. 85 (sine descript.). 1830; ibid. 464. 

Coreopsis mitis Michx. Fl. Bor. Amer. 2: 140. 18Q3=Bidens mitis 
(Michx.) Sherff, Bot. Gaz. 81: 43. 1926. 

Coreopsis molokaiensis (Hillebr.) Drake del Cast. 111. Fl. Ins. 
Mar. Pacif. 210. l89Q=Bidens molokaiensis (Hillebr.) Sherff, op. cit. 
70: 97. 1920. 

Coreopsis multifida DC. et var. mutica DC. Prodr. 5: 573. 1836= 
Bidens pilosa var. bimucronata f. odorata (Cav.) Sherff, op. cit. 81: 
41. 1926. 

Coreopsis nitida H. R. M. ex Elench. PI. Hort. Bot. J.-J. Destremx 
1805: 10. 1806, nomen (ex synon.)=Bidens aurea (Dryand. in Ait.) 
Sherff, op. cit. 59: 313. 1915. 

Coreopsis odorata Poir. in Lam. & Poir. Encycl. Suppl. 2: 350. 
1811; etiam Lam. & Poir. Atlas pi. 704, f- 1 (sine anno) -Bidens 
pilosa var. bimucronata f. odorata (Cav.) Sherff, op. cit. 81: 41. 1926. 

Coreopsis odoratissima Cav. ex Pers. Syn. PL 2: 477. 18Q7=Bidens 
pilosa var. bimucronata f. odorata (Cav.) Sherff, loc. cit. 



REVISION OF THE GENUS COREOPSIS 457 

Coreopsis Oerstediana Benth. ex Oerst. Kjoeb. Vid. Medd. 93. 
1852=Bidens Oerstediana (Benth. ex Oerst.) Sherff, op. cit. 80: 385. 
1925. 

Coreopsis ovata Cav. Icon, et Descr. 3: 41, pi. 280. 1795= Acti- 
nomeris ovata (Cav.) Nutt. ex DC. Prodr. 5: 576. 1836 (cf. Nutt. 
Gen. 2: 181. 1818). 

Coreopsis parviflora Jacq. Hort. Schoenbr. 3: 65, pi. 374- 1798= 
Cosmos parviflorus (Jacq.) HBK. Nov. Gen. et Sp. 4: 189 (241), 
1820. 

Coreopsis polycephala (Schz. Bip.) Benth. & Hook, ex Drake del 
Cast. 111. Fl. Ins. Mar. Pacif. 217. 1890; etiam Fl. Polyn. Fr. 108. 
1892=Bidens polycephala Schz. Bip. Flora 39: 360. 1856. 

Coreopsis procera Dryand. in Ait. Hort. Kew. ed. 1. 3: 253. 1789= 
Actinomeris alternifolia (L.) DC. Prodr. 5: 575. 1836. 

Coreopsis pukhella 0. Hoffm. Bot. Jahrb. 38: 204. l9Q6=Bidens 
microphylla Sherff, Bot. Gaz. 90: 390. 1930. 

Coreopsis quadricornis Krock. Fl. Siles. 2, pt. 2: 481. 1793= 
Bidens cernua L. Sp. PI. 832. 1753. 

Coreopsis Remyi Drake del Cast. 111. Fl. Ins. Mar. Pacif. 210. 
l89Q=Bidens micrantha Gaud. Voy. Freycinet Bot. PI. 85 (sine 
descript.). 1830; ibid. 464. 

Coreopsis reptans L. Syst. Nat. ed. 10. 2: 1228. 1759; Amoen. 
Acad. 5: 381 (nomen) et 407. l76Q=Bidens reptans (L.) G. Don in 
Sweet, Hort. Brit. edit. 3. 360. 1839. 

Coreopsis Ridens Gunn. Fl. Norveg. 2: 87. 1772 (sphalm)- 
Coreopsis Bidens Li.=Bidens cernua L. Sp. PI. 832. 1753. 

Coreopsis Rueppellii Schz. Bip. ex Walp. Repert. 6: 163. 1846= 
Bidens Rueppellii {Schz. Bip. ex Walp.) Sherff, Bot. Gaz. 90: 389. 
1930. 

Coreopsis ruwenzoriensis S. L. Moore, Journ. Linn. Soc. 35: 345. 
l9Q2=Bidens coriacea(0. Hoffm.) Sherff, op. cit. 81: 52. 1926. 

Coreopsis Sambucifolia Cav. Descr. 225. 18Q2=Bidens Sambucifolia 
Cav. Icon, et Descript. 3: 15, pi. 229. 1795. 

Coreopsis sandvicensis (Less.) Benth. & Hook, ex Drake del Cast. 
111. Fl. Ins. Mar. Pacif. 210. 1890= J Bz'dews sandvicensis Less. Linnaea 
6: 508. 1831. 

Coreopsis scandens Sess & Moc. PI. Nov. Hispan. ed. 2. 137. 
1893; etiam Fl. Mex. ed. 2. 194. 1894=Bidens squarrosa HBK. 
Nov. Gen. et Sp. 4: 187 (238). 1820. 



458 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Coreopsis Schaffneri A. Gray, Proc. Amer. Acad. 19: 15. 1883= 
Bidens Schaffneri (A. Gray) Sherff, Bot. Gaz. 56: 493. 1913. 

Coreopsis Schkchteri (Sherff) Burtt Davy in Kew Bull. 568. 1935= 
Bidens Schkchteri Sherff, Bot. Gaz. 76: 146. 1923. 

Coreopsis Seretii DeWild. Ann. Mus. Congo, ser. 5. 2: 212. 1907= 
Bidens Seretii (DeWild.) Sherff, op. cit. 76: 162. 1923. 

Coreopsis serrata DC. Prodr. 5: 595. 1836=Bidens laevis (L.) 
Britton, Sterns, & Poggenb. Prel. Cat. 29. 1888. 

Coreopsis serrulata Poir. Encycl. Suppl. 2: 352. 1811=Bidens 
serrulata (Poir.) Desf. Cat. Hort. Par. ed. 3. 186. 1829. 

Coreopsis serrulata (Schz. Bip.) Benth. & Hook, ex Drake del 
Cast. 111. Fl. Ins. Mar. Pacif. 210. 1890; etiam Fl. Polyn. Franc. 109. 
1892=Bidens bipontina Sherff, Bot. Gaz. 85: 10. 1928. 

Coreopsis setigera Schz. Bip. ex Walp. Repert. 6: 163. 1846= 
Bidens setigera (Schz. Bip.) Sherff, op. cit. 90: 390. 1930. 

Coreopsis simplicifolia (Vatke) Engler, Abh. Preuss. Akad. Wiss. 
1891, 2: 435. l892=Bidens Dielsii Sherff, op. cit. 90: 388. 1930. 

Coreopsis speciosa Hiern, Cat. Welw. Afr. PI. 1, pt. 3: 585. 1898= 
Bidens grandis Sherff, op. cit. 59: 309. 1915. 

Coreopsis Steppia Steetz in Peters, Nat. Reise Mossamb. Bot. 
496. 1862-1864=Bidens Steppia (Steetz) Sherff, op. cit. 76: 82. 1923. 

Coreopsis Stuhlmannii 0. Hoffm. in Engler, Pflanzenw. Ost-Afr. 
C: 415. 1899=Mens Stuhlmannii (0. Hoffm.) Sherff, op. cit. 76: 
158. 1923. 

Coreopsis tanna Forst. ex Steud. Nomencl. ed. 2. 1: 420. 1840= 
Glossogyne tenuifolia Cass. Diet. Sci. Nat. 51: 475. 1827. 

Coreopsis tannensis Spreng. Fl. Hal. Mant. 53. 18Q7=Glossogyne 
tenuifolia Cass. loc. cit. 

Coreopsis Taylori S. L. Moore, Journ. Bot. 44: 22. 19Q6=Bidens 
Taylori (S. L. Moore) Sherff, Bot. Gaz. 59: 309. 1915. 

Coreopsis ternata Chiov. Ann. Bot. Roma 9: 74. 1911=Bidens 
ternata (Chiov.) Sherff, op. cit. 90: 391. 1930. 

Coreopsis tetragona Cerv. ex La Llave & Lex. Nov. Veg. Descr. 
1: 31. 1824=Bidens aurea (Dryand. in Ait.) Sherff, op. cit. 59: 313. 
1915. 

Coreopsis trichosperma Michx. Fl. Bor. Amer. 2: 139. 1803= 
Bidens coronata (L.) Britt. Bull. Torr. Bot, Club 20: 281. 1893; cf. 
Sherff, op. cit. 56: 495. 1913. 



REVISION OF THE GENUS COREOPSIS 459 

Coreopsis trichosperma var. tenuiloba A. Gray, Syn. Fl. N. Amer. 
1, pt. 2: 295. lSS4:=Bidens coronata var. tenuiloba (A. Gray) Sherff, 
op. cit. 86: 446. 1928. 

Coreopsis trifida Poir. Encycl. Suppl. 2: 353. I811=Thelesperma 
filifolium A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 301. 1884. 

Coreopsis trifoliata Bertol. Fl. Guat. 36. 184Q=Bidens squarrosa 
HBK. Nov. Gen. & Sp. 4: 187 (238). 1820. 

Coreopsis trilobata Vahl ex Klatt, Leopoldina 23: 90. 1887= 
Montanoa pauciflora Klatt, loc. cit. 

Coreopsis tripartita M. B. Moss, Kew Bull. 184, 196. 1929=Bidens 
Mossii Sherff, Bot. Gaz. 92: 202. 1931. 

Coreopsis ugandensis S. L. Moore, Journ. Linn. Soc. 35: 347. 1902 
=Bidens ugandensis (S. L. Moore) Sherff, op. cit. 59: 309. 1915. 

Coreopsis variifolia Salisb. Prodr. 206. !796=Bidens reptans (L.) 
G. Don in Sweet, Hort. Brit. ed. 3. 360. 1839. 

SPECIES INCERTAE SEDIS 

Coreopsis acuta Pursh, Fl. Amer. Sept. 2: 569. 1814 (Conopsis 
acuta Pursh ex Less. Syn. Gen. Compos. 228. 1832 sphalm). 
Georgia. "Is perhaps Actinomeris squarrosa Nutt." (Torr. & Gray, 
Fl. N. Amer. 2: 349. 1843.) 

Coreopsis aspera Pursh Fl. Amer. Sept. 2: 570. 1814. Maryland. 
"Not identified and probably not of the genus" (A. Gray, Syn. Fl. 
N. Amer. 1, pt. 2: 290. 1884). 

Coreopsis bituminosa Noronha, Verh. Batav. Gen. 5. ed. 1. Art. 
IV. 11. 1790. Malaya. 

Coreopsis brasiliensis Colla, Herb. Pedem. 3: 479. 1834. 

Coreopsis emarcida Noronha, Verh. Batav. Gen. 5. ed. 1. Art. 
IV. 11. 1790. Malaya. 

Coreopsis flexicaulis Raf. Med. Repos. N.Y. Hexade 2. 5: 361. 
1808. Southern New Jersey. "Not identified and probably not of 
the genus" (A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 290. 1884). 

Coreopsis (per errorem typ. Cereopsis) gracilis Blanco, Fl. Filip. 
ed. 2. Suppl. 591. 1845. (Cf. Fernandez-Villar in Blanco, op. cit. 
ed. 3. pi. 287. 1878-1880; etiam Merrill, Sp. Blanc. 380. 1918.) 

Coreopsis japonica Hort. Dammann ex Wien 111. Gartenzeit. 20: 
438, /. 41. 1895. Japan. 

Coreopsis lanceolata var. longipes Kew Bull. App. 2: 68. 1931. 
Nomen nudum. 



460 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 

Coreopsis minima Hort. ex Steud. Nom. ed. 2. 1: 420. 1840. 
North America. 

Coreopsis Neaei DC. Prodr. 5: 573. 1836. South America. 

Coreopsis perfoliata Walt. Fl. Carol. 214. 1788. Carolina. Cf. 
Bidens laevis (L.) Britton, Sterns, & Poggenb. Prel. Cat. 29. 1888. 
(The small Walter Herbarium in London (Brit.) lacks a specimen of 
C. perfoliata.) 

Coreopsis philadelphica L. ex Jacks. Ind. Linn. Herb. 62. 1912. 
Nomen nudum. 

Coreopsis radiata Mill. Gard. Diet. ed. 8. Coreopsis No. 5. 1768. 
South Carolina. Perhaps Bidens laevis (L.) Britton, Sterns, & 
Poggenb. Prel. Cat. 29. 1888. 

Coreopsis repens Sesse* & Moc. Fl. Mex. ed. 2. 194. 1894. Mexico. 

Coreopsis scabra Raf. in Robin & Raf. Fl. Ludov. 72. 1817 
(Conopsis scabra Raf. ex Less. Syn. Gen. Compos. 228. 1832 sphalm). 
Louisiana. 



INDEX OF COLLECTORS CITED 1 

Page numbers in italic type 



Abrams, L. R., nos. 3342, 3628, 11788, 
11915, 313 

no. 9465, 29 
Achten, L., no. 253B, 387 
Adams, J. W M no. 480, 423 
Allard, F., nos. 65, 330, 387 
Allen, J. A., Iowa, 398 
Andre, E., no. 4387, 334 
Anonymous, ex T. Nuttall, 409 

no. 575, 418 

no. 576, Arkansas, Sept., 1835, 401 

Georgia, 438 

mts. of N. Carolina, Sept., 1843, 416 

Arkansas Nat. Forest, 1909, 355 

Bar Harbor, Aug., 1906, 406 

Gaffat near Debra Tabor, 1862, 450 

Macon, Georgia, 438 

Santa Maria R., Feb. 7, 1880, 308 

Smithfield, 1871, 423 

Tampa Bay, 344 

cult., June, 1818, 401 

cult, ex seminibus ex herb. DC. 
anno 1839, 429 

cult, in Hort. Berol. anno 1860, 44? 
Anthony, A. W., no. 216, 306 

no. 394, 305 

Antinori, Prov. Scioa, 1878, 447 
Armstrong, Margaret, no. 741, 306 
Arsene, Bro. G., Cerro Azul, Oct., 1909, 
320 

nos. 3185, 5295, 6026, 6060, 318 
Ashe, W. W., Chapel Hill, July, 1898, 
401 

Babcock, H. H., Chicago, June 17, 

1874, 344 

Bachelew, F. S., no. 223, 305 
Bacigalupi, R., no. 1209, 309 
Baikie, W. B., Niger River, 372 
Bailey, L. H. & E. Z., no. 13047, 433 
Bain, S. M., no. 95 pro parte, 349 

no. 95 pro parte, 406 
Baites, Mexico, 1864, 298 
Baker, C. F., Blount Springs, May 5, 

1898, 406 

St. Croix Falls, Aug. 15, 1900, 398 
no. 4090, 313 
Ball, J., near Chicla, Apr. 21-23, 1882, 

339 

Barker, Carrie, no. 31, 418 
Barnes, C. R., Converse, June 30, 1877, 

399 



Barnes, C. R., & W. J. G. Land, no. 
155, 360 

no. 158,317 

nos. 165, 313, 301 
Barnhart, J. H., no. 2715, 431 
Barter, C., Borgu, 372 

no. 933, 395 

Bartlett, H. H., no. 1638, 09 
Batchelder, C. Foster, Falmouth, Aug. 

14, 1906, 423 

Bates, J. M., no. 2958, 418 
Battiscombe, E., no. 83, 382 

no. 945, 379 
Baum, H., no. 878, 387 
Bavicchi, no. 14, 387 
Beardslee, H. C., & C. A. Kofoid, 
Great Smoky Mts., Aug., 1891, 411 

Swain County, Aug., 1891, 349 

Swain, Aug., 1891, .406 
Bebb, M. S., District of Columbia, 
1863, 401 

Marion Co., 1860, 399 

Winnebago Co., 1859, 399 

Fountaindale, 411 
Bebb, R., nos. 50, 1048, 399 

no. 1174, 418 

Beckwith, Florence, no. 54, 411 
Becquet, A., no. 62, 391 
Benke, H. C., no. 327, 118 

no. 1841, 411 

no. 3183, 342 

no. 3279, 353 

no. 3967, 399 

nos. 4306, 4657, 418 
Bequaert, Joseph, Leopoldville, Apr. 9, 
1915, 388 

no. 113, 387 

no. 7205, 388 

Berlandier, J. L., nos. 566, 882, 1073, 
426 

no. 1802, 359 

nos. 1956, 2312, 2503, 2580, 426 
Berlin Bot. Garden (Hort. Berol.), ex, 
353 

1806-1812, 350 

cult., July 25, 1815, 402 

Sept., 1832, 343 

cult., Sept., 1834, 402 

cult., 1837, 402 

July, 1837, 421 

cult., Sept. 11, 1876, 413 
Bernoulli, Gust., no. 196, 298 



1 Unnumbered specimens are cited, generally, in the order of exactness as to 
geographic location, in alphabetical order as to locality, or in chronological order. 

461 



462 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 



Beyrich, H. C., banks of rivers, Georgia, 
349 

thickets, Georgia, 406 
Bigelow, J. M., Ft. Smith, 1853-1854, 342 

Ft. Smith, June 23, 1853, 344 

Mohave Creek, 1853-1854, 309 
Billington, C. B., no. 131, -439 
Biltmore Herbarium, Hammond, Sept. 
17, 1900, 427 

near Pembroke, June, 1901, 438 

no. 173a, 411 

nos. 175a and b, 406 

nos. 176, 1766, 557 

no. 307c, 351 

no. 437a, 401 

no. 2064A, 404 

no. 20666, 439 

no. 2067a, 342 

no. 20706, 443 

no. 2139, 441 

no. 53746, J>16 

no. 56936, 344 

no. 147586, 438 

no. 14759a, 442 
Bischoff , G. W., ex herb., 357 
Blair, Walter, Prince Edward Co., 

June, 1881, 401 
Blake, S. F., no. 9542, 402 
Blakley, O. W., no. 1451, 399 

no. 3420, 411 
Blankinship, J. W., Southern Pines, 402 

Greene Co., 411 
Blanton, F. S., no. 6872, 431 
Bogusch, E. R., no. 994, 421 
Bolander, H. N., Julian City, SIS 

no. 4520, 31 6 

Bonpland, A., equatorial America, 339 
Boott, William, herb, of, Winter Pond, 

July 30, 1876, 423 
Boriani, no. 42, 395 

Bourgeau, Eugene, Saskatchewan, Sept. 
17, 1857, 429 

Winnipeg Valley, 1859, 399 
Boykin, S. B., Georgia, 342 
Boynton, F. E., Highlands, 1889, 349 
Brandegee, K., San Diego, Apr. 25, 1900, 
313 

Redstone Park, July 25, 1905, 418 
Brandegee, T. S., The Dalles, 1882, 429 

Isl. Santa Cruz, Apr., 1888, 308 

Alcalde, Apr., 1891, 311 

San Martin Isl., Apr., 1897, 306 

San Diego, June, 1900, 306 

San Diegc, May, 1906, 306 

no. 3368, 313 
Brannon, M. A., near Crown Point, 

1889, 342 

Bray, W. L., no. 48, 428 
Brendel, Frederick, Illinois, 342 

Illinois, 399 

Peoria, 399 

Peoria, 411 



Brewer, W. H., no. 241, 309 
Bridges, Thomas, no. 260, 316 
Brinton, J. B., Egg Harbor, Sept. 3, 
1888, 423 

Egg Harbor, Aug. 18, 1889, 424 
Britton, N. L., Norfolk, June 10, 1892, 

418 

Britton, Mrs. N. L., no. 132, 431 
Bross, Mason, Hyde Park, June 3, 1878, 

342 
Brown (S.), Britton (N. L.), & Bisset 

(P.), no. 2011, 354 
Brownfield, J., Summerville, Oct., 1892, 

439 

Bruce, Mrs. C. C., no. 1987, 316 
Buchanan, John, no. 380, 377 
Buchner, no. 32, 387 
Buchtien, Otto, no. 97, 337 

no. 595, 339 
Buckley, S. B., southeastern U. S., 402 

Alabama, 349 

pine woods, mts. of Alabama, 05 

Alabama, June, 1841, 357 

Alabama, July, 1841, 06 

mts. of Carolina and Georgia, 416 

Florida, 431 

Illinois, 399 

Virginia, 357 
Buettner, R., no. 142, 397 

no. 408, 388 
Burgess, A. B., no. 433, 411 

no. 513, 412 
Bush, B. F., no. 84, 344 

no. 158, 355 

no. 159, 418 

no. 160, 399 

no. 172A, 3.49 

no. 696, 422 

no. 3017, 418 

no. 3249, 399 

no. 3416, 397 

nos. 6041, 6137, 6531, 7872, 3*9 
Butler, G. D., no. 14, 418 

no. 16, 353 

Calkins, W. W., nos. 135, 145, 412 

Cambridge Bot. Gard. (see Hort. Bot. 
Harvard Univ.) 

Camp, S. H. & D. R., Bay View, 

July 14, 1896, 342 
Jackson Co., Aug. 18, 1897, 412 
Jackson Co., Aug. 18, 1898, 412 

Canby, W. M., Felton, Sept., 1867, 424 
near Wilmington, Oct., 1867, 441 
vicin. Wilmington, Oct., 1867, 39 
swamps, Delaware, Aug., 1877, 424 
Florence, July 3, 1878, 409 
Indian River, Feb., 1889, 431 
nos. 53, 54, 357 
no. 55, S44 
no. 157, 418 

Carey, John, North Carolina, 351 



INDEX OF COLLECTORS 



463 



Carey, John, & Asa Gray, North 
Branch of New River, etc., 351 

Virginia, July, 1841, 351 
Carpenter, W. M., Louisiana, 349 
Carson, Alex., no. 75, 393 

no. 106, 376 
Chandler, H. P., no. 5123, 306 

no. 7010, 1*18 
Chang, Tang Chung, & Nong Sing Po, 

no. 3790, 1*19 
Chapman, A. W., wet barrens, 1+36 

middle Florida, 344 

ex herb., southern Florida, 1*39 

Georgia, 1*15 

mts. of Georgia, 349 

mts. of Georgia, 357 

mts. of Georgia, 3*16 

Apalachicola, W 

Spring-brook, 44# 

Aspalaga, 1897, 1*1*1 

ex herb., no. 1746, 1*36 

ex herb., 20666, 1*1*1 

ex herb., 2066c, 1*1*1 

herb, of, 20706, 1*1*3 

Chase, Agnes, Rogers Park, Aug. 27, 
1896, 1*12 

nos. 2382, 2474, 1*02 
Chase, V. H., n. of Wady Petra, Aug. 

12, 1896, 1*12 

Cheo, H. C., & W. F. Wilson, no. 11 
pro parte, 31*2 

no. 11 pro parte, 1*18 

no. 29, 1*18 
Chevalier, Aug., nos. 2083, 9633, 9721, 

395 

Chiovenda, Emilio, District of Semien, 
Dec. 3, 1909, 1*1*7 

no. 970, 371 

no. 1090, 372 

no. 1148, 371 

no. 1820, 1*1*7 

nos. 1869, 1953, 1954, 371 

no. 2205, 1*1*7 
Chipp, T. F., no. 66, 362 
Churchill, J. R., shore of L. Michigan, 
1*12 

Lookout Mt., June 2, 1911, 1*06 
Claessens, J., nos. 67, 73, 88, 400, 387 
Clark, H. L., Falmouth Rd., Aug. 16, 

1899, 31*2 
Clemens, Mr. & Mrs. Joseph, no. 978, 

1*18 

Clements, F. E., no. 2778, 1*18 
Clements, F. E. & E. S. f nos. 224, 225, 
313 

no. 226, 306 
Cleveland, Daniel, San Diego, 306 

San Luis Rey, 1877, 313 
Clokey, I. W., no. 2428, 399 
Clokey, I. W. & B. Templeton, no. 4520, 
309 



Clothier, G. L., & H. N. Whitford, 

no. 1037, 399 
Cocks, R. S., Cameron, July, 1903, 1*26 

no. 3631, 1*27 
Coe, Mary A., Atlanta, May 6, 1903, 

357 
Collins, J. F., Spectacle Pond, Sept. 

11, 1900, 1*21* 
no. 8, 1*26 
Collins, J. F., M. L. Fernald & H. H. 

York, no. 294, 1*21* 
Combs, Robert, & C. R. Ball, no. 517, 

399 
Congdon, J. W., Lincoln, Aug. 22, 1871, 

424 

Mariposa Water Ditch, March 28, 

1901, 316 
Mariposa Water Ditch, Apr. 4, 1903, 

315 
Conzatti, Cassiano, no. 1431, 298 

nos. 1955, 2074, 299 
Conzatti, Cassiano, & Gonzalez, nos. 

545, 546, 299 
Coombs, Mrs. A. L., Warner's Springs, 

1 Q1 1 010 
lifll, Old 

Coulter, J. M., near LaPorte, July 22, 
1878, 399 

Kankakee River, 1882, 399 
Coulter, Thomas, no. 316, 308 

no. 339, 313 

Coville, F. V., no. 127Ark., 355 
Coville, F. V.,&F. Funston, no. 742, 309 
Cowles, R. B., Coronado Isls., March 

29, 1921, 306 
Craig, Newsom, & Hiland, no. 80, 309 

no. 129, 311 
Cratty, R. I., Iowa, 399 

Armstrong, July 18, 1897, 399 

no. 16, 399 

Croft, Mary B., San Diego, 1885, 1*26 
Curtis, M. A., N. Carolina, 1*37 
Curtis, Mrs. R. C., cult, in 1930 from 

Angola seeds, 385 
Curtiss, A. H., Florida, November, 37 

Virginia, 1*02 

Bedford Co., July 1, 1868, 07 

Bedford Co., June 1, 1871, 357 

Bedford Co., June 10, 1871, 401 

Bedford Co., June 30, 1871, 07 

Bedford Co., May 31, 1872, 357 

Bedford Co., July 3, 1872, 07 

Chattahooche, 1875, 439 

near Jacksonville, Sept., 437 

Pensacola, 1885, 31*2 

Pensacola, summer of 1885, 31*6 

no. 17, 349 

no. 1470, 437 

no. 1477 pro parte, 437 

no. 1477 pro parte, 439 

no. 1477 pro parte, 441 

no. 1480, 433 

no. 1481, 342 



464 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 



no. 1484, MS 

no. 1485, 557 

no. 1487, 1>07 

no. 4487, 441 

no. 4488, 445 

no. 4489, 457 

no. 4713, 445 

no. 5266, 457 

no. 5348 pro parte, 457 

no. 5348 pro parte, Ml 

no. 5375, 431 

nos. 5568, 6278, 457 

no. 6362, 359 

no. 6390, 544 

no. 6471, 406 

no. 6734, 452 

no. 6825, 404 

no. 10223, 441 
Curtiss, C. C., Martha's Vineyard, 

Aug. 13, 1892, 424 

Cuthbert, Alfred, Augusta, Sept, 25, 
1898, 441 

Georgia, Oct. 10, 1898, 38 

Davis, John, near LaGrange, June 9, 
1915, 412 

no. 1855, 406 

nos. 2097, 7818, 407 
Deam, C. C., & H. A. Gleason, Bluffton, 

412 
Deam, Mrs. C. C., no. 1709, 431 

no. 1864, 421 

Dedrick, Allan, dry, sandy soils, 406 
De Giorgi, S., vicin. Elisabethville, 1923, 
597 

no. 142, 231 

no. 170, 592 
De Jelski, Const., no. 705, 555 

no. 765, 525 
De Selm, A. W., no. 140, 342 

nos. 275, 276, 394, 599 

nos. 499, 621, 412 
Dewart, F. W., St. Cloud, Sept., 1892, 

599 
Dick, W. M., Lorain Co., July 31, 1895, 

412 

Dickey, S. S., no. 96, 412 
Dillon, Quartin, Abyssinia, 447 

no. 612, 446 
Dixon, R. A., no. 17, 418 

no. 528, 342 
Dixon, R. A., & Cornelia Gage, no. 713, 

342 
Dodge, C. K., near Port Huron, Aug. 

4, 1895, 412 
Douglas, David, N. America, 429 

California, 312 

California, 515 

California, 1833, 512 

Ft. Vancouver, 429 
Douglas, A. E., Arequipa, 1892, 340 



Drake, D., & J. H. Dickson, Bradfords 

Isl., June, 1889, 429 
Dreisbach, R. R., no. 1042, 41 8 
Drummond, Thomas, Texas, 41 8 

no. 68, 426 

no. 69, 421 

no. 199, 559 

no. 200, 421 

Dudley, W. R., no. 340, 509 
Duesner, C. W., Miller, 1908, 412 

Pine, 1908, 342 
Duges, Alfredo, no. 472, 500 
Dummer, R. A., no. 3304, 361 
Dunkle, M. B., no. 1857, 508 
Dunlap, no. 204, 574 
Dunn, G. W., Coronado Isls., Apr. 17, 
1891, 306 

Eames, E. H., no. 8825, 551 
no. 8839, 421 

Earle, F. S., Auburn, Sept. 4, 1899, 412 
nos. 2060, 2061, 406 
no. 2147, 454 

Earle, F. S. & E. S., no. 73, v555 

Earle, F. S., & C. F. Baker, Auburn, 

Apr. 4, 1897, 557 
Auburn, Aug. 17, 1897, 406 
Auburn, May 28, 1898, 407 
no. 243, 542 

Eastwood, Alice, Kaweah, Apr. 27, 

1895, 509 

Cuyama, May 2, 1896, 512 
Point Sal, May 12, 1896, 308 
no. 2565, 506 
no. 2574, 515 
no. 3187, 512 
nos. 3194, 4377, 515 

Eaton, A. A., no. 469, 455 

Eggert, Heinrich (Henry), St. Louis, 

Aug. 25, 1874, 412 
St. Louis, June 29, 1875, 599 
Jefferson Co., July 8, 1879, 549 
St. Louis, June 18, 1886, 599 
Jefferson Co., May 20, 1887, 542 
St. Louis, Aug. 23, 1887, 412 
Jefferson Co., June 10, 1891, 544 
Williamsonville, June 11, 1893, 555 
Hematite, May 25, 1896, 544 
Jefferson Co., May 25, 1896, 542 
Sherwood, June 8, 1897, 406 
Stone Mt. and vicin., July 23, 1897, 

454 

Yellow R., July 27, 1897, 409 
no. 122, 412 

Eggert, H., & Sydow, St. Louis, July 1, 
1874, 599 

Eggleston, W. W., no. 19577, 308 
no. 19748, 515 

Ehlers, J. H., no. 518, 542 

Ehrenberg, K. A., no. 354 pro parte, 295 
no. 354 pro parte, 299 



INDEX OF COLLECTORS 



465 



Ellenbeck, no. 529, 368 
no. 1436, S72 
no. 1660, 447 
Elmer, A. D. E., no. 601, 429 

no. 3634, 308 
Emig, W. EL, no. 146, 399 
Engelmann, George, Mountain Creek 

bottom, 1833, 349 

vicin. East St. Louis, May, 1845, 342 
St. Louis, Sept., 1850, from Fendler's 

seeds, 418 

St. Louis, Aug., 1863, 349 
Norristown, Aug. 27, 1876, 407 
herb, of, Aiken, Apr., 1882, 344 
Engelmann, Henry, Ft. Laramie, July, 

1858, 418 

Engler, Adolph, no. 1266, 377 
Evermann, B. W., L. Pend d'Oreille, 
Aug. 7, 1893, 418 

Fauntleroy, Juliet, no. 654, 407 
Fendler, August, no. 397 pro parte, 418 
no. 397 pro parte, 426 
no. 441, 426 

Ferguson, A. M., Tampa, 1898, 432 
Fernald, M. L., E. B. Bartram, & B. 

Long, nos. 24687, 24688, 424 
Fernald, M. L., C. H. Bissell, C. B. 

Graves, B. Long, & D. H. Linder, 

no. 22859, 424 
Fernald, M. L., & B. Long, nos. 10664, 

10669, 17591, 17594, 424 
no. 19265, 351 

nos. 24689, 24690, 24691, 24692, 424 
Fernald, M. L., B. Long, & D. H. 

Linder, nos. 22860, 22861, 22862, 

424 

Ferris, Roxana S., no. 7466, 313 
no. 7572, 308 
no. 7710, 313 

Feudge, J. B., no. 1382, 312 
Fiebrig, Karl, no. 3476, 339 
Fiori, Adriano, no. 1826, 447 
Fish, Fanny E., All Saints' Bay, May, 

1882, 306 

Fisher, G. L., no. 5007, 422 
Fitzpatrick, T. J. & M. F. L., ex herb., 

Allamakee Co., June 27, 1895, 399 
Flamand, F., Kiamohampa, Sept. 20, 

1911 387 
Fleming, 'G. L., & W. W. Eggleston, 

no. 19676, 306 

Fletcher, E. F., Westford, 418 
Ford, H. C., Santa Barbara Co., 1889, 

310 

Francis, Mary E., no. 155, 437 
Fredholm, Adolph, nos. 5473, 5778, 

6094, 431 
Frost, W. D., Willmar, July, 1892, 399 

Galen, James, no. 4289, 412 
Galeotti, H. G., no. 2086, 298 



Garber, A. P., keys of Florida, 1876, 
431 

Tampa Bay, 1876, 431 

L. Monroe, March, 1876, 431 

Mellon ville, March, 1876, 431 

Tampa, May, 1876, 431 

Rosewood, June, 1876, 431 

Levy Co., Nov., 1877, 437 
Garvens, Wolfgang, vicin. Palm Beach, 

1908, 431 
Gates, F. C., no. 508, 342 

no. 598, 399 

no. 808, 412 

no. 942, 344 

no. 2478, 342 

no. 2817, 344 

no. 3148, 399 

no. 10104, 412 

Gates, F. C. & M. T., no. 10547, 342 
Geyer, C. A., Beardstown, 344 
Ghiesbreght, A. B., nos. 133, 539, 298 
Gibbs, Mrs. E. T., South Pond, 424 
Gillet, J., nos. 806, 885, 387 
Gironcourt, Mission of, no. 256, 372 
Gleason, H. A., east of Rantoul, 
Oct. 5, 1907, 412 

Shirland, June 29, 1908, 399 

Milroy, July 6, 1908, 399 

no. 326, 399 

nos. 812, 1813, 1977, 412 

nos. 2749, 2750, 2753, 412 
Goetze, W., no. 731, 382 

no. 790, 369 

no. 1041, 377 

no. 1069, 378 

Grabham, G. W., Bahadur Gorges, 371 
Graebner, P., cult. Hort. Berol., Aug. 5, 

1910, 350 
Grant, G. B., nos. 681, 682, 313 

no. 5424, 310 
Grant & Walter Wheeler, nos. 680, 

2228, 308 

Grassly, C. W., Hyde Park, June 10, 
1877, 344 

Miller, Aug. 31, 1880, 412 
Gray, Asa, Hickory Nut Gap, 1843, 416 

Apalachicola, Apr., 1875, 443 
Gray, A., & John Carey, mts. of 
Virginia and N. Carolina, July, 
1841, 407 
Gray, A., & W. S. Sullivant, mts. of 

Carolina, 1843, 416 
Greenman, J. M., no. 437, 424 

no. 544, 412 

nos. 2017, 2592, 342 

no. 3771, 350 

no. 3991, 412 

Gregg, Josiah, no. 901, 426 
Griffiths, David, Highmore, July, 1897, 

429 
Grimes, E. J., no. 3768, 402 



466 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 



Gronovius, D., ex herb., 402 
Groth, B. H. A., no. 66, 422 
Guenther, Ernesto, & O. Buchtien, no. 
55, 319 

Hale, Josiah, Louisiana, 3^5 

Louisiana, 412 

Louisiana, 4-18 

Louisiana, 426 

Louisiana, 1842, 342 

Alexandria, S57 
Hall, Elihu, Oregon, 429 

central Illinois, 350 

Athens, 1861, 342 

Athens, 1861, 399 

Athens, 1861, 412 

cult, from Texas seeds, 1864, 359 

Athens, Aug., 1864-1867, 412 

Athens, Aug., 1868, 412 

Hempstead, June, 1872, 421 

cult, from Dakota seeds, 1877, 429 

no. 342, 3 5 It 

no. 343, ^21 

no. 344 pro parte, 418 

no. 344 pro parte, 426 

no. 345, 418 

Hall, E. E., Texas, 1877, 353 
Hall, H. M., no. 3074, 310 

no. 3973, 313 
Hall, James, Virginia, 1828-1834, 402 

Montgomery Co., 1828-1834, 407 
Haller, W. H., no. 982, 353 
Hamerlinck, P., nos. 1502, 1512, 387 
Hanson, H. C., no. A13, 426 
Harbison, T. G., Waynesville, 1897, 407 

Highlands, May, 1902, 357 

Highlands, 1905, 407 

Highlands, Aug. 1, 1905, 403 

Highlands, Aug., 1906, 403 
Harford, W. G. W., & G. W. Dunn, 

Cascades, May 28, 1869, 429 
Harper, R. A., no. 74, 357 
Harper, R. M., vicin. Sawdust, June 18, 
1927, 409 

no. 5, 402 

no. 22, 342 

no. 46, 407 

no. 131, 407 

no. 545, 405 

no. 653, 441 

no. 1001, 443 

no. 1666 pro parte, 439 

no. 1666 pro parte, 441 

no. 1954, 412 

no. 2198, 443 

Hartweg, Theodor, no. 116, 321 
Harvard Univ. Bot. Gard. (reading 
Hort. Bot. Harv. Univ. or Hort. 
Cantabr. on some labels), 1842, 404 

cult., 1842, 407 

cult., 1844, 421 

ex Cambridge, Aug., 1845, 406 



cult., 1848, from Lindheimer's Texas 

seeds, 359 
cult., 1849, 409 

cult, from Carolina seeds, 1849, 416 
cult, from Texas seeds, 1849, 426 
cult, from Fremont's seeds, 1851, 

1852, 306 

cult, from Wright's seeds, 1852, 422 
cult., Sept., 1856, 359 
cult., 1871, 312 
cult., 1873, 407 
cult., 1873, 1882, 316 
cult., 1882, 313 

Harvey, F. L., Little Rock, 418 
no. 22, 355 

Hasse, H. E., Mojave Station, 312 
Desoto, July, 1887, 350 
Los Angeles, Apr. 22, 1888, 313 
Hayes, Sutton, near Amatitlan, July 20, 

1860, 298 

Heddle, J. R., no. 698, 399 
Heiland, Lychen, Aug. 16, 1882, 418 
Heller, A. A., vicin. Salisbury, Aug. 21, 

1890, 412 
near mouth of Tucquan Creek, Sept. 1, 

1890, 412 
Coeur d'Alene River, July 12, 1892, 

429 
about L. Pend d'Oreille, Aug. 1-10, 

1892, 429 
no. 99, 407 
no. 933, 402 
no. 1013, 407 
no. 1548, 428 
no. 1730, 422 
no. 7662, 310 
no. 7674, 313 
no. 7723, 312 
no. 11752, 418 

Heller, A. A., & E. G. Halbach, near 
McCall's Ferry, Sept. 9, 1893, 413 
Heller, A. A. & E. G., no. 4129, 413 
Heller, Edmund, Mt. Garguez, Aug. 26, 

1911, 363 

Hens, Fr., no. 45, 387 
Herrera, F. L., nos. Ill, 391, 1186, 1481, 

S0 

Hexamer, A. C., & F. W. Maier, 
Summerville, May 27, 1855, 404 
Abbeville Co., June, 1855, 406 
ex herb., Soluda R., June 14, 1855, 

357 

rich woods, June 16, 1855, 342 
Heyde, E. T., & E. Lux, no. 3792, 298 
Hildebrandt, J. M., no. 417, 446 
Hill, E. J., Morgan Park, June 17, 1876, 

O t 

nos. 1108, 2746, 345 
Hinkley, Mr. & Mrs. F. E., no. 51, 3^0 
Hitchcock, A. S., Jacksonville, 443 

Iowa City, 1888, -413 

New River, 1896, 431 



INDEX OF COLLECTORS 



467 



Palm Beach, 1896, 1*31 
Levy Co., 1898, 431 
Madison Co., June-July, 1898, 31*3 
no. 139, 433 
no. 279, 1*18 
no. 666, M8 
no. 910, 439 
no. 13943, 1>18 
no. 21564, 829 
Hitchcock, C. L., no. 12224, 313 

no. 12340, 310 
Hoffman, W., Corn Creek, June 21, 

1870, 899 

vicin. Rolla, July 15, 1871, 350 
Holm, Theodor, Brookland, June, 1894, 

402 
Holway, E. W. D., Decorah, July 16, 

1888, 399 

nos. 5, 52, 154, 613, 298 
Holzinger (Holtzinger), J. M., Lamoille, 

June, 1895, 899 
Santa Fe, Aug. 13, 1911, WO 
Hombl<, no. 329, 397 
Hooker, J. D., Arkansas, 1+18 

Kentucky, 407 

Hort. Cantabr., see Harvard Univ. Bot. 
Gard. (for Hort. Berol., etc. see 
Berlin, etc.) 
Houghton, Douglass, 200 miles above 

St. Anthony's Falls, 399 
above St. Anthony's Falls, July, 1832, 

399 
Houghton, H. W., no. 3567, 853 

no. 3864, 350 

House, H. D., no. 150, 402 
no. 4023, 350 
no. 9660, ^It 

Howell, A. H., no. 808, 407 
Howell, J. T., no. 8220, 308 
Howell, T. J., Oregon, 1881, 429 
Klickitat, June, 1881, 429 
banks of Columbia, July, 1881, 429 
no. 224, 1*29 
Hrdlicka, vicin. San Damian, Jan.- 

Feb., 1913, 340 
Huett, J. W., Ottawa, 1*13 
Hulier, Hort., no. 8, 300 
Humboldt, Alexander von, & Aime" 

Bonpland, near Loja, 329 
Paramo de Saraguru, 329 
Hunnewell, F. W., 2nd, White Sulphur 

Springs, July 4-6, 1914, 407 
Hunt, Edwin, Sudbury, Aug. 12, 1874, 

4*4 

Hutchinson, Mary E., Randolph, 343 
Hyams, M. E., Statesville, July, 1880, 
357 

Jackson, F. J., Kikuyu region, 1889, 361* 
Jaeger, E. C., no. 140, 313 
Jameson, William, Ecuador, 335 

hills near Loja, commun. 1865, 334 



Jermy, Gustav, no. 657, 423 
Johnson, L. N., Chicago, June 19, 1891, 

343 
Johnston, I. M., near Mojave, May 5, 

1920, 312 

near Mojave, May 5, 1920, 318 
Victorville, May 15, 1920, 313 
5 miles s.e. of Victorville, May 17, 

1920, 311* 
1 mile s. of Crutts P.O., May 14, 1922, 

312 

no. 1251, 314 
Jones, M. E., Pasadena, Feb. 22, 1882, 

314 

Fallbrook, March 25, 1882, 314 
San Diego, Apr. 3, 1882, 314 
Todos Santos Bay, Apr. 11, 1882, 306 
Darwin, Apr. 28, 1897, 310 
Shepherd's Canyon, Apr. 30, 1897, 

310 

Pleasant Canyon, May 6, 1897, 310 
Whitewater, May 11, 1903, 314 
Cajon Pass, May 16, 1903, 310 
Victor, May 17, 1903, 314 
Mojave, May 20, 1903, 312 
Marsh Lake, Sept. 17, 1903, 435 
San Diego, Apr. 14, 1906, 314 
Julian, Apr. 17, 1906, 314 
Sand Point, Sept. 18, 1908, 1*29 
Cave Spring, March 12, 1924, 310 
Santa Cruz Isl., Apr. 13, 1924, 308 
ranch 29 miles s.w. of Tia Juana, 

Apr. 13, 1925, 306 
near Santa Ana Canyon, March 10, 

1926, 314 

Malibu Hills, Apr. 26, 1926, 303 
Cuyama Canyon, Apr. 28, 1926, 312 
Coronado Isls., June 10, 1926, 306 
Adelanto, May 12, 1927, 312 
Santa Cruz Isl., March 25, 1929, 308 
no. 3134, 306 

nos. 3361 pro parte, 3373, 314 
nos. 28012, 28013, 423 
no. 29474, 418 
nos. 29511, 29512, 359 
Jones, P. M., Santa Rosa Isl., Apr., 

1901, 308 
Jones, W. W., no. 160, 426 

no. 227, 1*18 
Joor, J. F., Galveston Bay, June, 421 

Kalbreyer, W., no. 125, 373 
Kassner, T., no. 2776, 398 

no. 2871, 383 
Katzenstein, Otto, herb., s.e. U.S., Sept. 

23, 1897, 437 
Kearney, T. H., Jr., Clear Creek, Sept., 

1893, 415 
White Cliff Springs, July 14, 1894, 

407 

no. 553, 413 
Kellerman, W. A., no. 6296, 298 



468 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 



Kellogg, A., & W. G. W. Harford, no. 

436, 312 
no. 439, 816 
Kellogg, J. H., Allenton, Aug. 12-13, 

1884, 850 

St. Louis, July 15, 1903, 350 
cult., June 30, 1905, 421 
cult., Mo. Bot. Gard., May 23, 1911, 

554 
Kellogg, William, Rancho Encinal, 

April 15, 1903, 31 1* 
Owanita, 1907, 451 
Kennedy, G. G., Orleans, July 31, 1891, 

4#4 

Ormond, March 29, 1897, 545 
Sharon, Sept. 2, 1904, 424 
Kennedy, W. L., Fort Tejon, spring, 

1876, 310 
Kenoyer, L. A., & F. W. Pennell, no. 

3224, 424 

Kersting, nos. 14, 422, 395 
Kew., Hort., cult, from R. Diespecker's 

seeds, 584 
Killip, E. P., & A. C. Smith, no. 21536, 

31*0 

no. 21823, 325 
no. 22171, 555 
no. 23304, 540 
King, Anna, no. 116, 405 

nos. 534, 566, MS 
Kling, nos. 60, 166, 597 
Knopf, E. C., nos. 56, 258, 508 
Krebs, William, Cleveland, 415 
Kreger, F. 0., no. 315, 1*29 

Lang, H. A., Egg Harbor, Sept. 10, 

1905, 425 

Langlois, A. B., western Louisiana, 
Apr., 1880, 545 

Spring Hill, Oct. 1, 1880, 436 
Lansing, O. E., Jr., no. 243, 545 

no. 307, 599 

no. 647, 1*13 

nos. 848, 935, 545 

no. 1646, MS 

nos. 2715, 2782, 5*5 

no. 2797, WO 

no. 2832, 415 

no. 3113, 400 

nos. 3822, 3911, MS 
Lapham, I. A., Milwaukee, 545 

Milwaukee, 400 
Laurent, Em., between Lusambo and 

Lomani, 587 
Laurent, Em. & Marc., Kondue, Nov. 

25, 1903, 587 

Laurent, Marc., no. 512, 587 
Leavenworth, M. C., Louisiana, 555 

Louisiana, 1*00 

Tampa Bay, 1*31 
Lebrun, Jean, no. 4507, 1*1*1* 
L4card, no. 290, 595 



Le Conte, North America, 55.4 
Ledermann, C., no. 1855, 372 

no. 5243, 595 

Leeds, B. F., Chattanooga, Sept. 29, 
1892, 1*06 

Chattanooga, Sept. 29, 1892, 1*07 
Lehmann, ex, 1*00 

Leiberg, J. B., Kootenai Co., June, 1890, 
1*18 

L. Pend d'Oreille, July, 1892, 1*19 

no. 1542, 1*29 

Lely, H. V., nos. 383, 449, 568 
Lemmon, J. G., top of Mt. Diablo, 310 

San Bernardino, March, 1880, 31 J* 

no. 137?, California, 312 

no. 138, 510 

no. 4159, 1*29 

no. 4520, 312 

Leonard, E. C., no. 4103, 505 
Lescrauwaet, E., no. 188, 587 
Letterman, G. W., Allenton, 550 

near Franklin Co. Road Bridge, July, 
1883, 5-45 

Allenton, Aug., 1883, 545 

Merely, July, 1884, 5-45 

Gratiot, May 30, 1897, 351* 

near St. Louis, Aug., 1899, 554 
Levine, C. 0., no. 778, 5-45 
Lewton, F. L., Forest City, 1894, 1*33 

Cedar Hammock, Aug. 3, 1894, 455 
Lighthipe, L. H., Monticello, Apr. 5, 
1891, 545 

no. 557, 457 

Linder, D. H., no. 1706, 588 
Lindheimer, F., w. of Brazos River, 
Apr., 559 

w. of Houston, early May, 1840, 545 

Houston, Apr., 1842, 554 

Galveston Id., Nov., 1842, 1*21 

Texas, 1849, 1*21 

no. 54, 1*21 

no. 101, 1*22 

no. 102 pro parte, 1*19 

no. 102 pro parte, 1*26 

no. 363, 559 

no. 548, 419 ; 

no. 897, 419 
Lips., Hort., ex Amer. septentr., 550 

ex Illinois, 599 
Lloyd, C. G., near Rockwood, Aug. 10, 

1880, 409 
Lloyd, F. E., & S. M. Tracy, no. 527a, 

54ff 

Lloyd, R. N., Ravenswood, June 3, 
1887, 545 

Ravenswood, July 11, 1887, 400 

Ravenswood, Aug. 31, 1887, 415 
Loring, Lake Superior, 545 
Luckhardt, no. 626, 575 
Lyall, David, Cascade Mts. to Ft. 
Colville, 1860, 429 

Colville to Rocky Mts., 1861, 429 



INDEX OF COLLECTORS 



469 



Macbride, J. F M no. 2932, 337 

nos. 3152, 3504, 321* 
Macbride, J. F., & Featherstone, no. 
210 337 

nos. 436, 690, 1237, 340 
MacCarthy, Gerald, District of Co- 
lumbia, 402 

eastern N. Carolina, 1883, 439 
MacDougal, D. T., no. 293, 426 

no. 617, 429 

MacKay, Ernest, no. 25, 430 
Mackensen, Bernard, no. 76, 422 
Mackenzie, K. K., no. 100, 351* 

no. 328, 350 
Macoun, J. M., Waneta, July 28, 1902, 

430 

Macoun, John, Sandwich, Aug. 4, 1892, 
US 

Winnifred, Aug. 12, 1895, 419 

no. 38, 419 

Magee, E. E., Hayward Co., Sept., 
1897, 407 

Waynesville, Sept. 9, 1897, 407 

below Satulah Mt., Aug. 2, 1902, 407 

Highlands, Sept. 2, 1902, 407 
Mann, Gustav, no. 1922, 373 
Manning, W. H., Lake City, Aug. 13, 
1883, 400 

Mackinac Isl., July 11, 1915, 343 
Manz, no. 29, 422 

Mathews, Alexander, Prov. Chacha- 
poyas, 1835, 1836, 328 

no. 571, 340 

no. 1376, 331 

no. 1418, 323 
McClatchie, A. J., San Diego, Apr., 

1893, 306 
McDonald, F. E., Peoria, June, 1891, 00 

Peoria, June, 1899, 343 

Peoria, Aug., 1900, 413 

Peoria, 1900-1901, 00 

Peoria, Sept., 1901, 413 

Peoria, June, 1904, 343 

Peoria, June, 1904, 400 

Peoria, Aug., 1904, 413 

Peoria, May, 1915, 345 

Peoria, June, 1915, 400 

Peoria, June, 1916, 343 

Peoria, July, 1919, 419 
McLean, Andes, Peru, 336 
Mead, S. B., Mason Co., June 18, 
1845, 343 

Illinois, 1850, 400 
Mearns, E. A., no. 144, 400 

nos. 1291, 1722, 364 
Mendez, 300 

Mexia, Ynes, no. 445, 358 
Meyer, Hans, no. 532, 444 
Michaux, Andr6, North America, 350 

North America, 404 

North America, 413 

North America, 439 



Migeod, F. W. H., no. 211, 374 
Mildbraed, F., no. 2539, 444 

no. 3392, 373 

Miller, E. S., Long Pond, Aug. 30, 1871, 
425 

Wading River, Aug. 18, 1877, 425 
Milligan, Mrs. J. M., Jefferson Co., 345 
Millspaugh, C. F., no. 3865, 413 

no. 4630, 308 
Mintz, no. 40, 373 
Moffatt, W. S., no. 105, 343 

no. 492, 413 
Mohr, Charles, Mentone, 403 

Mobile, 406 

ex herb., eastern shore Mobile Bay, 
Apr., 1868, 422 

Gulfport, July 4, 1874, 346 

Bladon, July, 1875, 403 

Lawrence Co., May, 1880, 357 

ex herb., Winston Co., May 1, 1881, 
357 

Cullman, June, 1883, 346 

vicin. Ashland, June, 1883, 346 

Mulberry R. Valley, June 6, 1883, 346 

cult, from Texas seeds, Mobile, 
May 10, 1887, 360 

Columbus, June 4, 1888, 346 

Mentone, Sept. 18, 1892, 403 

Gallion, May 25, 1893, 347 

Poplarville, Oct. 18, 1894, 435 

Yellow Pine, Aug. 8, 1895, 441 

DeSoto Falls, Sept. 12, 1898, 403 
Moore, A. H., no. 1832, 419 

nos. 2552, 2553, 413 
Morgan, R. T., vicin. Fountaindale, 

Aug., 1877, 400 
Morong, Thomas, ex herb., Winchester, 

Aug. 5, 1881, 425 
Morris, E. L., no. 105, 402 

no. 1050, 407 
Mosely, E. L., Oxford, Aug. 21, 1895, 

413 

Muenzner, no. 159, 382 
Munroe, H. F., Hyde Park, May, 1879, 
3L.3 

Hyde Park, June, 1879, 343 
Munz, P. A., no. 1589, 406 

nos. 2148, 2538, 3814, 314 

no. 4466, 310 

no. 7955, 306 

nos. 9136, 9186, 310 

no. 9357, 314 

no. 10042, 310 

no. 11378, 308 

no. 12415, 314 
Munz, P. A., & E. Crow, nos. 11553, 

11691, 11750, 308 
Munz, P. A., & R. D. Harwood, nos. 

3385, 3475, 314 

Munz, P. A., R. D. Harwood, & I. M. 
Johnston, no. 4077, 310 



470 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 



Munz, P. A., & D. D. Keck, nos. 7842, 

7879, 310 
no. 7930, 311,. 
Munz, P. A., L. Street, & G. Williams, 

nos. 2362, 2388, Silt 
Munz, P. A., & J. Voss, no. 11877, 308 
Murdoch, John, Jr., Orleans, Aug. 22, 

1900, 425 
Murdoch, Joseph, Rochester, Aug. 24, 

1902, 425 

Nash, G. V., nos. 832, 1257, 1876, 431 

no. 2255, 433 

no. 2585, 415 

Nealley, G. C., Brazos Santiago, 1889, 
428 

Chenates, 1889, 419 
Negri, Giovanni, no. 56, 447 

nos. 2356is, 618, 448 

no. 9156is, 367 

Nelis, Bokala, May, 1913, 387 
Nelson, E. W., no. 1718, 299 

nos. 2185, 2249, 318 

nos. 6104, 6813, 426 
Nelson, J. C., no. 1802, 419 

no. 1980, 430 
Ness, H., no. 2078, 360 
Neumann, Oscar, no. 75, 448 

no. 134, 379 
Newberry, Sitgreaves Pass, March 26, 

314 

Nicolas, Puebla, Oct., 1905, 299 
Norton, J. B., e. of Hartsville, Nov. 5, 
1921, 441 

no. 1455, 419 
Nutt, W. H., between L. Tanganyika 

and L. Rukwa, 1896, 382 
Nuttall, L. W., no. 571, 308 
Nuttall, Thomas, without data, 314 

Red River, 419 

Cakes, Plymouth, 425 

Olive, E. W., no. 72, 426 

Orcutt, C. R., Point Loma, March 12, 

1884, 314 

Point Loma, March 24, 1884, 306 
northern Baja California, Apr. 17, 

1886, 81k 
Orcutt, C. R., & W. S. Bowne, San 

Diego, 1884, 306 
Oyster, J. H., no. 4014, 426 

Palmer, Edward, no. 41, 308 
no. 126, 314 
no. 172, 360 
no. 217, 314 
no. 286, 431 
nos. 489, 530,317 
no. 568, SOS 
nos. 635, 655, 426 
no. 677, 314 



Palmer, E. J., Newton Co., July 15, 
1906, 419 

no. 17, 343 

no. 2272, 419 

nos. 2317, 2317A, 350 

nos. 2386, 2386A, 343 

no. 2513, 350 

no. 6962A, 352 

no. 7740, 422 

no. 8120, 355 

no. 8468, 415 

no. 8468, 415 

no. 9144, 300 

no. 12115, 423 

no. 18078, 00 

no. 27056, 345 

no. 29347, 350 

no. 29421, 415 
Pammel, L. H., no. 621, 413 
Pappi, A., nos. 1194, 1303, 2642, 448 

no. 7733, 395 
Paris., Hort., 1816, 350 

1820, 404 

1823, 418 
Paris, Hort. Bot. de 1'Ecole de M<d. de, 

Aug., 1820, 413 
Parish, S. B., Marengo, Apr., 1882, 310 

nos. 2211, 5191, 314 

no. 9771, 312 

no. 10415, 314 

Parish, S. B. & W. F., San Bernardino, 
Apr., 1881, 314 

foothills of San Bernardino Mts., 
May, 1888, 314 

no. 226, 310 
Parker, C. F., Bristol, Aug. 11, 1865, 

425 
Parlin, J. C., North Berwick, July 4, 

1892, 419 

Parry, C. C., Pacific coast of N. 
Amer., 1880, 310 

California, 312 

San Bernardino, 314 

San Diego, 308 

San Diego, 1850, 305 

San Diego, 1883, 306 

no. 115, 310 

no. 116, 314 

Parry, C. C., & J. G. Lemmon, no. 184, 
310 

no. 185, 314 

Patterson, H. N., Henderson Co., Aug., 
1871, 413 

near Oquawka, July, 400 

near Oquawka, June 20, 1872, 400 
Pearce, R., Cordillera Huanta, Feb., 

1867, 338 

Pease, A. S., s. of Champaign, Oct. 9, 
1920, 400 

no. 12394, 400 

no. 17993, 343 



INDEX OF COLLECTORS 



471 



Peattie, D. C., no. 8, 400 

nos. 164, 2029, 413 
Peirson, F. W., no. 3900, 430 
Peng, To Kang, Ts'ang Wai Tak, & 

Ts'ang Un Kin, no. 936, 419 
Pennell, F. W., no. 4029, 434 

no. 13244, 340 

no. 14342, 537 

nps. 14396, 14710, 340 
Petit, A., Abyssinia, 446 

Abyssinia, 448 
Pfund, T., Dar-Fur, 1876, 395 

no. 336, 395 
Piemeisel, R. L., & L. W. Kephart, no. 

166, 366 

Pierce, W. M., 20 mi. e. of Victorville, 
Apr. 12, 1922, 314 

desert 20 mi. e. of Victorville, Apr. 

12, 1922, 31 2 
Pitcher, Zina, Arkansas, 354 

Arkansas, 419 
Plowden, Abyssinia, 371 
Plummer, Miss, mts. above Santa 

Barbara, 1878, 308 
Pogge, P., nos. 257, 1282, 387 

no. 1292, 389 

nos. 1295, 1309, 387 
Pollard, C. L., no. 1222, 414 
Pollard, C. L., & W. R. Maxon, no. 496, 

347 

Poole, S. F., Sharon, July, 1905, 343 
Porter, T. C., middle Georgia, 1846, 407 

near Augusta, Apr., 1847, 343 

Susquehanna R., Aug. 22, 1861, 413 

40 mi. s. of Lynchburg, July 17, 1880, 
402 

9 mi. above Kanawha Falls, July 21, 

1880, 350 

Powell, B. T., Hastings, July, 1886, 419 
Preuss, Kamerun, 1891, 373 

no. 689, 373 

Pringle, C. G., banks of Columbia R., 
Sept. 10, 1881, 430 

bluffs of seashore, San Diego, Apr. 
25, 1882, 306 

hills bordering Mojave Desert, May 
11, 1882, 310 

Mojave Desert, May 14, 1882, 312 

nos. 2197, 2303, 317 

no. 2367, 361 

nos. 3570, 3841, SOS 

no. 4871, 304 

no. 4896, 299 

no. 5508, 318 

no. 7866, 320 

no. 8218, 298 

no. 8328, 426 

nos. 8332, 8431, 320 

no. 8781, 301 

no. 9885, 361 

no. 9886, 320 



no. 9895, 300 
no. 10050, 320 
no. 11506, 361 
no. 11507, 317 
no. 11546, 303 
no. 11900, 301 
no. 13041, 298 
no. 13547, 300 
Purpus, C. A., no. 1339, 300 
no. 1550, 298 
nos. 2581, 2581a, 302 
no. 3099, 300 
no. 4098, 304 
no. 5004, 316 
nos. 5063, 5531, 310 

Quarre", P., no. 419 pro parte, 386 
no. 419 pro parte, 390 
no. 1616, 391 
no. 1662, 394 
nos. 2173, 2826, 2921, 389 

Ragazzi, V., Let-Marefia, Sept. 4, 1886, 
448 

Let-Marefia, Sept. 6, 1886, 448 

Antoto, Oct. 2, 1887, 448 

no. 250, 448 
Raimondi, A., no. 436, 330 

no. 1774, 323 

no. 3022, 335 

no. 6463, 329 

nos. 6695, 7732, 7914, 333 

no. 12060, 329 

Randolph, Fannie R., no. 136, 431 
Randolph, L. F. & F. R., no. 150, 402 

no. 1066, 407 
Ravenel, H. W., Santee Canal, 345 

damp pine land, Santee Canal, Sept., 
441 

damp rich soils, Santee Canal, Sept., 
442 

Santee Canal, July, 1846, 409 
Redfield, J. H., no. 5630, 439 
Reed, F. M., no. 6147, 306 
Reverchon, Julian, White Creek, July 4, 
422 

Dallas, May, 1874, 354 

Dallas, May, 1876, 354 

western Texas, 1882, 423 

Pine Isl., May 5, 1903, 348 

nos. 513, 1478, 354 

nos. 2041, 2077, 348 
Reynolds, Carrie A., Stony Is!., Aug. 24, 

1909, 413 
Reynolds, H. S., Judsonia, May 31, 

1877, 355 
Riddell, J. L., Kentucky, 407 

Ohio, 413 

Ridgway, Robert, no. 60, 413 
Riehl, Nicholas, no. 41, 400 

no. 65, 350 



472 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 



Robinson, B. L., Normal, July, 1887, 
1>00 

Hendrix, Aug. 31, 1904, 413 

no. 56, 413 

no. 79, 407 

Robyns, Walter, no. 138, 389 
Rohlfs & Stacker, Katz, Dec. 30, 1880, 

448 

Godofelassi, Jan. 7, 1881, 370 
Rolfs, P. H., no. 309, 343 

no. 392, 357 

Rose, J. N., no. 2344, 301 
Rose, J. N., A. Pachano, & G. Rose, 

no. 23146, 334. 
Rose, J. N., & J. H. Painter, no. 7480, 

303 
Rose, J. N., J. H. Painter, & J. S. Rose, 

no. 8837, 298 
no. 9717, 300 

Rosendahl, C. O., no. 656, WO 
Rothrock, J. T., cult., Philadelphia, 

June 25, 1877, 419 
no. 157, 426 
no. 317, 419 

Rowland, western Lagos, 1893, 355 
Rugel, Ferdinand, Broad R., June-July, 

1841, 350 

Broad R., July, 1841, 416 
valley of Blue Ridge Mts., Aug., 1841, 

41.4 

Florida, 1842-1849, 439 

between Quincy and Aspalaga, May, 
1843, 350 

Florida, 1844, 407 

no. 102, 432 

no. 133, 433 

no. 142, 432 

no. 480, 437 

no. 570 pro parte, 441 
Runyon, Robert, no. 335, 428 
Rtippell, Eduard, between Temben and 

Siemen, 1832, 448 
Rusby, H. H., no. 1685, 337 
Russell, Margaret P., Asheville, 407 

Hood R., Aug. 15, 430 

herb, of, Rochester, Aug. 17, 1918, 

425 

Ruth, Albert, Chilhowee Gap, July, 
1893, 409 

Mount Nebo, July, 1893, 408 

Knoxville, June, 1893, 407 

Knoxville. May, 1895, 357 

Knoxville, June, 1895, 408 

Knoxville, May, 1896, 409 

Lookout Mt., July 20, 1897, 408 

no. 67, 357 

no. 69, 414 

no. 482, 354 

nos. 612, 624, 403 

no. 652, 357 

nos. 655, 665, 350 
Rydberg, P. A., no. 9445, 35.Z 



Salvin, Osbert, Volcan del Fuego, 

Aug., 1873, 298 
Sandberg, J. H., Hennepin Co., Aug., 

1889, 400 

Hennepin Co., July, 1890, 400 
Sandberg, J. H., D. T. MacDougal, 

& A. A. Heller, no. 617, 430 
Sapin, A., N'Dolo, May, 1906, 387 

N'Kolokosso, June 28, 1906, 387 

Sokomini L., Sept., 1906, 387 

Katola Msek, Apr., 1908, 388 

Munungu, Apr., 1910, 388 
Sargent, C. S., no. 27, 340 
Sartwell, H. P., Maumee R., 4*4 
Saurman, B. F., Apalachicola, 343 

Apalachicola, 443 

Savage, T. E., J. E. Cameron, & F. E. 
Lenocker, Spokane, July, 1898, 419 

The Dalles, Aug., 1898, 419 
Savage & Stull, no. 60, 400 
Scammon, Franklin, Chicago, 345 

Chicago, June, 1859, 343 

Chicago, June 3, 1859, 343 
Schimper, Wilhelm, no. 48, 448 

no. 62, 446 

nos. 71, 85, 448 

no. 429, 446 

no. 933 pro parte, 450 

no. 1173, 371 

no. 1236, 448 

no. 1386, 370 

no. 1423, 448 

sect. 5, no. 15, 450 

sect. 5, nos. 22, 95, 448 

sect. 5, no. 933, 450 

sect. 5, no. 1005, 448 
Schindler, A. K., no. 335, 419 

no. 345a, 343 

Schlechter, Rudolph, no. 12474, 388 
Schneck, Jacob, LaGrange, May, 1894, 

422 

Schott, Arthur, near Georgetown, Aug., 
1857, 402 

near Georgetown, June 22, 1860, 402 
Schroeder, F., no. 74, 395 
Schuette, J. H., Door, July 30, 1887, 343 

St. Anthony Park, July 2, 14, 1888, 
400 

St. Anthony Park, July 8, 1888, 400 

St. Anthony Park, July 16, 1888, 400 

Plover, July 26, 1888, 400 

cult., Green Bay, Sept. 14, 1888, 419 

cult., Green Bay, Sept. 14, 1888, 422 

n. of Sturgeon Bay Canal, July 14, 

1890, 343 

cult., Green Bay, Aug. 23, 1897, 419 
Green Bay, Aug. 13, 1899, 400 
Green Lake, Aug. 13, 1899, 400 
cult., Green Bay, July 31, 1901, 419 
cult., Green Bay, Aug. 2, 1901, 419 
no. 76, 354 
Schulz, Ellen D., no. 367, 423 



INDEX OF COLLECTORS 



473 



Schweinfurth, G. (Fl. Gallabat 439), 
Matamma, Oct., 1865, 395 

(Fl. Gallabat 585), Matamma, Oct., 

1865, 396 

Scott, Hugh, Mt. Zuguala, 1926, 448 
Seemann, Berthold, no. 675, 33%. 
Seymour, F. C., no. 1384, 1*25 
Shaw, Marjories, no. 1941, 310 
Shaw, Marjorie, Edna Spaulding, & 
Mrs. C. Li. Walton, Antelope 
Valley, 310 

Sheldon, E. P., Sleepy Eye, July, 1891, 
ItOO 

Ft. Snelling, June, 1895, 400 

Pike Island, July, 1895, 1*19 

no. 11081, 1*30 
Shepard, E. M., Greene Co., July, 1880, 

350 
Sherff, E. E., no. 250 pro parte, 419 

no. 250 pro parte, 430 

no. 253, 402 

no. 278, 354 

no. 284, 400 

no. 498, 422 

nos. 499, 507, 419 

no. 989, 350 

no. 1038, 414 

nos. 1642, 1781, 400 

nos. 5000, 5000a, 419 

no. 5003, 404 

no. 5004, 402 

no. 5005, 350 

no. 5006, 343 

no. 5007, 408 

nos. 5013, 5014, 345 

no. 5015, 422 

no. 5016, 345 

no. 5017, 360 

no. 5018, 422 

Sherwood, W. L., Waynesville, July, 
1897, 408 

Tuckasiegee Valley, Sept. 20, 1897, 

408 

Shimek, Bohumil, Monte Sano, July, 
1891, 408 

Oklahoma City, July, 1892, 419 
Short, C. W., hilly lands, Kentucky, 408 

Lexington, 1835, 357 

no. 42, 350 
Shriver, Howard, Wytheville, 1875, 409 

Wytheville, July, 1877, 408 
Simon, Gust., no. 3, 373 
Simpson, J. H., Manatee, 1889, 437 
Skeels, H. C., no. 304, 343 

no. 375, 400 

Skehan, Josephine, Ocean Springs, 
Sept. 16, 1895, 436 

no. 137, 406 

Small, J. K., n. of Marysville, Aug. 15, 
1888, 414 

vicin. mouth of Tucquan Creek, 
Sept. 1, 1890, 414 



vicin. mouth of Tucquan Creek, 

Oct. 11, 1890, 414 

Round Top Mt., July 2, 1892, 408 
Farmer Mt., July 12, 1892, 408 
Nick's Creek, Aug. 5, 1892, 408 
Stone Mt., July 3, 1893, 406 
Little Stone Mt., July 7, 1893, 354 
Little Stone Mt., July 7, 1893, 409 
banks of Yellow R., July 11, 20, 1893, 

355 
between Alcovy R. and No Business 

Creek, July 14, 1893, 354 
Tallulah Falls, Aug. 4, 1893, 414 
Estatoah Falls, Aug. 12, 1893, 41 4 
Thomas Bald, Aug. 19, 1893, 350 
Little Long Creek, Aug. 22, 1894, 402 
Altamaha R. Swamp, June 18-21, 

1895, 404 

Stone Mt., Aug. 1-6, 1895, 434 
base of Currahu Mt., July, 1896, 350 

Small, J. K., & J. J. Carter, Bull Key, 

Nov. 6, 1903, 432 
no. 694, 439 
no. 954, 440 
no. 1028, 437 
no. 1219, 432 
no. 2731, 440 
no. 3025, 433 
no. 3401, 440 

Small, J. K., J. B. DeWinkeler, & 
C. A. Mosier, no. 11252, 443 

Small, J. K., & A. A. Heller, near R. 

& D. Railroad, June 3, 1891, 357 

n. of Hickory, June 25, 26, 1891, 409 

Green's Hill, July 10, 1891, 408 

between Cranberry and Linville, July 

17, 1891, 408 

Rocky Knob, July 24, 1891, 408 
Grandfather Mt., July 25, 1891, 408 
no. 100, 414 
no. 328, 408 
no. 372, 402 

Small, J. K., & G. V. Nash, Everglades, 
1901, 437 

Smith, C. L., Sierra de San Felipe, 
Oct. 11, 1894, 299 

Smith, E. C., no. 510, 400 

Smith, H. H., no. 1089, 350 
no. 1150, 355 
no. 2456, 350 
no. 2526, 409 
no. 2582, 350 
no. 5034, 308 

Smith, J. D., near Starke, Apr. 9, 1880, 

443 

Warm Springs, July 29, 1880, 408 
Caesar's Head, July, 1881, 408 
Pellicier's Creek, March 4, 1882, 433 
Richland Gap, Aug. 14, 1882, 351 
Wildcat Ridge, Aug. 21, 1882, 350 
Wildcat Ridge, Aug. 21, 1882, 351 
Sumter County, March 21, 1883, 432 



474 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI 



Cumberland Falls, Aug. 23, 1883, 351 

Columbia, Sept., 1883, 44* 

banks of Soquee R., Sept. 13-15, 
1883, 351 

Orangeburg District, Aug. 9, 1884, 
406 

Orangeburg District, Aug. 9, 1884, 
^09 

banks of Oconee R., Aug. 16, 1884, 
351 

near Montgomery, Aug. 26, 1885, 415 

west of Cahawba R., Sept. 2, 1885, 351 

Biloxi Bayou, Sept. 16, 1885, 436 

no. 602, 347 

no. 2028, 441 

Smith, L. C., nos. 25, 808, 299 
Snyder, Mrs. M. S., seashore, La Jolla, 
Feb. 25, 1895, 306 

La Jolla, March 15, 1895, 31 5 
Spalding, Clear Water, 430 
Sparano, no. 25, 388 
Spencer, Mary F., no. 86, 306 

no. 172, 315 

no. 407, 310 

nos. 563, 653, 653a, 315 

no. 1114, 345 

nos. 1954, 2040, 310 

no. 2123, 315 
Standley, Jeanette P., no. 195, 432 

no. 423, 437 

Standley, P. C., no. 4410, 344 
Steele, E. S., Washington, Aug., 1889, 

402 

Steetz, ex, ex Lexington, 357 
Steudner, H., no. 253, 448 

no. 349, 446 
Stevens, G. W., no. 784, 419 

no. 1212, 419 

no. 2387, 351 

no. 2713, 400 
Steward, A. N., no. 1510, 419 

nos. 1518, 1583, 1922, 344 
Stewart, J. T., northern Mississippi, 
1863, 557 

near Charleston, 1865, 440 

near Charleston, June 1, 1865, 404 

around Peoria, 1867, 400 
Stillman, valley of upper Sacramento R., 

316 

St. John, Harold, no. 2939, 425 
Stolz, Ad., no. 764, 382 

no. 1306, 377 

Straub, F. C., nos. 97, 159, 432 
Strauss, H., San Diego and near-by isls., 
Sept. 24, 1908, 306 

San Diego and near-by isls., Feb. 3, 

1912, 306 
Street, L., San Elejo Lagoon, May 12, 

1917, 306 

Stubel, A., no. 35 pro parte, 330 
Suksdorf, W. N., Columbia R., Aug. 26, 
1882, 430 



Taylor, Thomasyille, Sept. 30, 1903, 441 
Taylor, Katherine A., ex herb., Sum- 
merville, 440 

Columbia, May, 1890, 357 

Summerville, June 10, 1891, 404 
Terracciano, A., & A. Pappi, no. 2474, 

448 
Tharp, B. C., no. 5627, 427 

no. 5635, 360 

Thaxter, Roland, Cullowhee, June- 
July, 1887, 351 

Cullowhee, June-July, 1887, 408 
Thone, Frank, no. 232, 414 
Thoreau, D. B., no. 2.11, 384 
Thurber, George, Cranston, Aug., 1844, 

425 
Thurrow, W. F., Hockley, 1890, 344 

Hockley, 1890, 419 

Hockley, 1890, 422 

Hockley, 1890, 427 
Tourney, J. W., no. 594, 427 
Townsend, C. H. T., A192, 339 

no. A211, 326 

Townsend & C. M. Barber, no. 150, 427 
Tracy, S. M., no. 3456, 427 

no. 4778, 436 

no. 6920, 432 

no. 6921, 432 

no. 7142, 437 

no. 7256, 433 

no. 7355, 433 

no. 7704, 432 

no. 7902, 420 

no. 8542, 3*6 

no. 8550, 422 

no. 8565, 436 

no. 8670, 4H 

no. 8925 pro parte, 360 

no. 8925 pro parte, 422 

no. 9049, 408 

no. 9454, 344 

Tracy, S. M., & Lloyd, no. 508, 344 
Trask, Blanch, no. 76, 305 
Trelease, William, Hesperia, Apr. 10, 

1892, 310 

Troll, C., no. 3157, 340 
Tuckermann, Edward, Jr., ex herb., 
New England, 425 

Uhde, no. 627, 300 

Uline, E. B., Fish L., July, 1891, 400 

Mishawaka, July 1, 1891, 400 
Umbach, L. M., Miller, June 5, 1897, 
344 

Clarke, Aug. 28, 1897, 414 

Clarke, June 4, 1898, 344 

DuPage, July 6, 1898, 400 

Clarke, Aug. 20, 1898, 4H 

no. 2752, 419 

U. S. Expl. Exped. under Capt. Wilkes, 
above Obrajillo, 337 



INDEX OF COLLECTORS 



475 



Vanderyst, Hyacinthe, Ndembo, Aug. 
20, 1906, 388 

Ndembo, Jan. 7, 1907, 388 

Inkissi Falls, May, 1907, 388 

Leopoldville, May 23, 1907, 388 

Kumpako, 1908, 388 

Leopoldville, May 27, 1910, 388 

N'Boma, Apr. 28, 1911, 388 

nos. 5224, 700, 1099, 1187, 1341, 
1582, 4304, 388 

nos. 4885, 10097, 388 

nos. 12366, 21227, 389 
Van Hermann, no. 2686, WO 
Vasey, G. R., northern Alabama, 1878, 
408 

N. Carolina, 1878, 357 

N. Carolina, 1878, 408 

Lookout Mt., 1878, 414 

no. 290, 311 

no. 551, 430 

Verdick, E., Lukafu, 388 
Vermoesen, F., no. 2403, 389 
Vesterland, Otto, Marion Co., Novem- 
ber, 1889, 441 
Von Mechow, Alexander, nos. 33, 459, 

387 
Von Tuerckheim, H., no. 11-2043, 298 

Wallace, Los Angeles, 1854, 315 
Ward, L. F., Potomac flats, Aug. 12, 

1877, 414 

Ford Co., Oct. 5, 1897, 420 
Warner, S. R., Willis, June 1, 422 
Warszewicz, no. 37, 340 
Watson, Sereno, no. 217, 430 
Webb, R. J., Pippin L., Sept. 6, 1909, 

414 

Webber, H. J., Sanibel Isl., Jan. 29, 
1896, 432 

no. 414, 432 

Weberbauer, A., nos. 203, 2394 pro 
parte, 340 

no. 3812, 838 

no. 4048, 322 

nos. 5764, 5782, 340 

no. 6995, 332 

Wellman, F. C., Bailundo District, 388 
Werdermann, Erich, no. 1114, 336 
Werner, W. C , no. 92, 408 



Wharf, E. H., N. Carolina, 357 
Wheeler, H. E., no. 62, 354 

no. 82, 344 

White, Mark, no. 218, 420 
Whitford, H. N., nos. 64, 203, 344 
Whyte, Alexander, northern Nyassa- 
land, 382 

Mt. Zomba, Dec., 1896, 377 

no. 35, 377 

Wiegand, K. M., & W. E. Manning, 
no. 3410, 438 

no. 3414, 408 

no. 3415, 408 

nos. 3421, 3422, 434 
Williams, E. F., Cataumet, Sept. 15, 
1901, 425 

Boston, Sept. 3, 1903, 414 

Wickford, June 17, 1908, 345 

Concord, June 20, 1908, 344 
Williamson, C. S., Waycross, Apr. 15, 

443 
Willich, C. L., & F. W. Weiss, ex herb., 

427 

Woodson, R. E., Jr., no. 655, 351 
Woodson, R. E., Jr., & E. S. Anderson, 

no. 1545, 345 
Woytkowski, Felix, no. 23, 330 

no. 24, 327 
Wright, Charles, Texas, 354 

Texas, 422 

Texas, 423 

northern Georgia, 408 

no. 217, 423 

nos. 339, 341, 420 

nos. 340, 1236, 427 

no. 1237, 423 

Wright, W. G., San Bernardino, 1880, 
311 

Ventura Co., March, 1894, 308 

no. 172, 311 
Wright, W. H., no. 50, 344 

no. 221, 414 

Young, J. D., no. 153, 368 
Young, M. S., near Paloduro Canyon, 
Sept. 7, 1917, 427 

Zech., Graf., no. 168, 396 

Zeller, Cyril, Ellis Co., 1908-1911, 420 



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